Yale Peruvian expeditions, 1911-1915, Publications no. 34. "Igi'oe thffe Moots I for. the founding of a. Cclleg? in iHis Colonf •ifaLE^MViEiasi'inf- 1921 CONTRIBUTION PROM THE PERUVIAN EXPEDITIONS.! BR AR^ OF YALE UNIVERSITY AND THE NATIONAL GEOGRAPHIC SOCIETY PEKTJVIAN MOSSES R. S. WILLIAMS NEW YORK 1916 ffibrintatf-, wiihqit change of paging, from Bulletin of the Tokrbt Botanical Club 43: 323-334. pi. 17-80. June 29, 1916 [From ihe Bulletin of the Torrey Botanical Club, 43: 323-334,//. /7-20. 30 Je 1916.] ' Peruvian mosses R. S. Williams (with plates 17-20) The following list is made up of two collections. The first, obtained by Harry Ward Foote, on the Yale Peruvian Expedition of 191 1, was kindly forwarded by Dr. A. W. Evans of Yale. It consisted of forty-eight packets and contained thirty-seven species among which appear to be two novelties, also two species known before only from Patagonia. The specimens are from localities varying from about 900 to 3,300 meters in altitude. They were without number, and I have accordingly rearranged and numbered them from 1 to 48. The second collection consisted of forty-three packets, ob tained by Messrs. Cook and Gilbert, with the exception of three species by Hiram Bingham, while on the Yale University-National Geographic Society Peruvian Expedition in 1915. In this second collection four species are apparently new to science. I may say that it was through the cooperation of the United States Department of Agriculture that Messrs. Cook and Gilbert were detailed to accompany this expedition and the specimens were forwarded to me by Mr. William R. Maxon of the United States National Museum. The altitude at which this second collection was obtained varies from 1,800 to 4,100 meters, and there are thirty-three species not in the first collection, making seventy species in this list. The type specimens of the new species are deposited in the herbarium of the New York Botanical Garden. DlCEANUM MlTTENII C. Mull. Above Arma Valley, July, 1915, H. Bingham 2063. I have not been able to find any publication of this species except that by Brotherus in Engler & Prantl's Nat. Pflanzenfam. (1: 328), under subgenus 4. It is a plant of medium size with the ovate leaf-base quickly narrowed tq^SKfong, subulate (and serrulate point; the alar cells are numerous|m<^^afipwe|jif the 323 324 Williams: Peruvian mosses broader part of the leaf narrow and elongate with pitted walls. The collection is scanty and sterile. Campylopus leucognodes (C. Mull.) Paris. Lucumayo Valley, 3,600 m., June, 1915, Cook & Gilbert 1264a. Fissidens asplenioides (Sw.) Hedw. San Miguel, on rock, 1,820 m., September, 1911, Foote 1. Fissidens crispus Mont. Ollantaytambo, about 3,000 m., May, 1915, Cook & Gilbert 754. Fissidens rigidulus Hook. f. & Wils. Ollantaytambo, on rock, about 3,000 m., May, 1915, Cook & Gilbert 670. Trichostomum brachydontium Bruch. Cuzco, on dry rock, about 3,500 m., July, 191 1, Foote 2; these Cuzco specimens have leaves shorter and broader than in typical specimens and possibly should not be referred here, but they seem to be only a stunted form of the species; Lucma, on dead wood, 2130 m., August, 191 1, Foote 15a. Trichostomum semivaginatum Schimp. Ollantaytambo, about 3,000 m., May, 1915, Cook & Gilbert 501a. Leptodontium integrifolium sp. nov. Dioecious, the male plants about like the fertile, the flowers with outer leaves similar to those of the stem, the inner broadly oblong, rounded obtuse, scarcely one third the length of the outer, costate, of mostly golden-brown, elongate, smooth cells enclosing five or six antheridia and numerous, longer, filiform paraphyses: fertile plants simple or slightly branched, the stems 3-4 cm. long with scattered, loose tufts of radicles and leaves, when dry rather loosely imbricate and incurved-flexuous, when moist spreading-recurved ; stem-leaves 3—3.5 mm. long, ovate- lanceolate, the entire margins strongly recurved from a little below the apex to near the base and papillose; costa stout, papillose on the back, excurrent into a serrulate point sometimes 0.5 mm. long; cells of the upper part of leaf rather obscure, scarcely elon gate, 8-9 n in diameter, densely papillose on both sides, in the lower fourth of leaf, elongate, narrowly rectangular, smooth, except near the margin and costa; perichaetial leaves about 6 mm. long, convolute, with a subulate, abruptly spreading, serrulate point; fruit unknown. (Plate 17, figs. 1-7.) Williams: Peruvian mosses 325 Ollantaytambo, ' about 3,000 m., May, 1915, Cook & Gilbert 704a. Growing with Thuidium peruvianum. Globulina peruviana sp. nov. Inflorescence unknown: plants in rather compact mats with simple stems 1-4 mm. high, bearing radicles at the base and imbricate-appressed leaves rather uniformly placed along the stem or somewhat clustered at the apex and forming a globose bud about 0.5 mm. in diameter; stem in cross-section about 200 fi in diameter with cells of outer walls not differentiated and with a distinct central strand; leaves rotundate or slightly oblong, about 0.5 mm. long, more or less cucullate, the flat margins crenulate half way down or more; costa vanishing abruptly a little below the apex, rather weak in the lower part, widest near the apex, slightly rough on the back and papillose on the inner face, in cross-section showing two rows of medium-sized, thick- walled cells on the dorsal side and a single row of much larger cells on the ventral side; cells mostly gradually smaller from the base to the apex of the leaf, more or less shortly rectangular below, 12-16 n by 16-30 n, toward the apex square to rhomboidal, with sides 6-8 n long, finely papillose on both sides about the upper half of the costa, the marginal cells smooth; fruit unknown. (Plate 17, figs. 8-15.) Araranca, south side of Roya Pass, about 4,100 m., April, 1915, Cook & Gilbert 177a. Growing apparently on dry soil with Pogonatum polycarpum, at the highest elevation given for any of the collections. Barbula costata Mitt. Ollantaytambo, about 3,000 m., May, 1915, Cook & Gilbert 610b. A small amount of what is apparently this species occurred, with immature fruit, growing with Bryum argenteum. What appears to be this also was found sterile, mixed with Leskea gracillima, at the same locality by these collectors, 663a. Barbula pruinosa (Mitt.) Jacq. Cuzco, on dry rock, 3,500 m., July, 1911, Foote 3. Barbula replicata Tayl. Cuzco, on dry rock, 3,500 m., July, 191 1, Foote 4; Ollantay tambo, about 3,000 m., on soil, May, 1915, Cook & Gilbert 610a. 326 Williams: Peruvian mosses Mitten was correct, I believe, in thinking B. apiculata Hampe not distinct from this species. Barbula subulatula C. Mull. Urubamba, on damp earth, 2,900 m., July, 191 1, Foote 5. Tortula affinis Hampe. Sicuani, Vilcanota Valley, on stone, about 3,600 m., April, 1915, Cook & Gilbert 157; Ollantaytambo, on rock, about 3,000 m., May, 1915, Cook & Gilbert 612; Urubamba, on rock, 2,900 m., July, 191 1, Foote 6. Tortula bipedicellata Besch. and T. confusa Card., I believe, do not differ from T. affinis, which in turn is very close to T. pichin- chensis (Tayl.) Mitt., differing chiefly in having a low basal mem brane of the peristome, the cell structure of which also varies. In T. pichinchensis the cells are often as high as broad, and some what rhomboidal in slightly oblique rows; in T. affinis the cells are low, mostly two or three times broader than high. Tortula andicola Mont. Ollantaytambo, about 3,000 m., May, 1915, Cook Sf Gilbert 771. Tortula lacerifolia sp. nov. Dioecious : plants with stems (in this small collection scattered through cushions of Fabronia andina) mostly simple, 6-8 mm. high, with radicles at the base and leaves more or less aggregate toward the apex; stem-leaves twisting incurved when dry, erect- spreading when moist, oblong, mostly 2-2.5 mm- long, the margins usually deeply incised in the upper half into readily separating, narrow, irregular segments, often 0.25 mm. long and twisted part way round or inflexed or reflexed, the margins of the lower leaf being entire and recurved; costa golden-brown, stout, near the base 80 ix wide, smooth on the back, more or less papillose on the upper side, excurrent into a thorn-like spine, in cross-section showing two large guide-cells, a large band of stereid cells on the dorsal side, with two medium-sized cells on the ventral side; cells throughout upper leaf rather obscure, mostly hexagonal, mamillose and covered on both sides with small often C-shaped papillae, those of lower leaf pale, with few or no papillae and mostly short- rectangular, up to about 18 fi wide by 40-50/4 long; outer peri- chaetial leaves like those of the stem, the inner small, pale, more or less deeply cleft into numerous segments, the archegonia numerous, with few paraphyses; fruit unknown. (Plate 18, figs. 1-6.) Williams: Peruvian mosses 327 Ollantaytambo, about 3,000 m., July, 191 1, Foote 7. Grimmia micro-ovata C. Mull. Temple of Viracocha near Tinta, on rock, about 3,500 m., April, 1915, Cook & Gilbert 206; Ollantaytambo, on rock, about 3,000 m., May, 1915, Cook & Gilbert 6og. Grimmia ovata Web. & Mohr. Cuzco, on dry rock, about 3,500 m., 191 1, Foote 8. Grimmia rivulariopsis sp. nov. Dioecious, the male plant rather more slender than the fertile with often 2-3 flowers scattered along the stem; the inner peri- gonial leaves small, ovate, ecostate, or faintly costate, enclosing rather numerous, fusiform antheridia about 0.5 mm. long, without paraphyses: fruiting-plants abundantly branching, 2-3 cm. high, without radicles and bearing slightly secund leaves, erect-spreading when moist, mostly somewhat incurved-appressed when dry; stem-leaves oblong-ovate, about 2 mm. long, rather abruptly narrowed to an acute, serrulate apex, the margins entire, of a double thickness of cells and recurved from a little below the apex to near the base; costa stout, percurrent, smooth on both sides and flat or convex on the ventral side in the upper part; cells of stem-leaves distinct, smooth, with slightly thickened, scarcely or not sinuous walls, the median mostly 6 /j, wide by 6-8 n long, the basal more or less rectangular, 8 /j. wide by 16-22 n long; perichaetial leaves larger than those of the stem, about 3.5 mm. long, the costa of the inner leaves rather faint below, widest toward the apex, not quite percurrent, the cells of the lower half of leaf narrowly rectangular or linear with slightly thickened, straight walls; capsule ovate, immersed, about 1.5 mm. high, without stomata, a band of five or six rows of small, trans versely elongate cells about the rim, the median exothecal cells scarcely or not elongate, rather irregular with scarcely thickened or sinuous walls, up to 25 n in diameter; seta erect, scarcely one half the capsule in height; peristome-teeth lanceolate, about 120 /J, wide at the base and 400 n high, red, papillose, entire or often split at the apex or along the median line and slightly lacunose ; annulus none; lid convex, obliquely apiculate; calyptra little more than covering the apiculus, the base cleft into several lobes. (Plate 19.) Ollantaytambo, about 3,000 m., on rock in stream bed, May, 1915, Cook & Gilbert 753; and on rock, same place and collectors, 56i. Most closely related to G. amblyophylla C. Mull, of the South American species, but the apex of the leaf is very different, the 328 Williams: Peruvian mosses cells of the stem-leaves much smaller, with thinner, scarcely sinuous walls and the inner perichaetial leaves have much longer, narrower cells throughout the lower leaf. Rhacomitrium crispipilum (Tayl.) Jacq. Lucumayo Valley, 3,000 m., June, 1915, Cook &f Gilbert 1264. Funaria calvescens Schwaegr. Ollantaytambo, about 3,000 m., on wet soil, May, Cook & Gilbert 669; Manchu Picchu, about 2,100 m., May, Cook & Gil bert 996. MlELICHHOFERIA ANDINA Sull. Araranca, south side of La Roya Pass, about 4,100 m., on earth wall, April, 1915, Cook & Gilbert 182a. MlELICHHOFERIA bogotensis Hampe. Ollantaytambo, about 3,000 m., May, 1915, Cook & Gilbert 705. LEptobryum Wilsoni (Mitt.) Broth. Arequipa, on wet rock, 2,300 m., June, 191 1, Foote 9. Anomobryum filiforme (Dicks.) Husn. Santa Rosa, on dry hillside, 3,960 m., July, 1911, Foote 10. Bryum andicola Hook. Above San Miguel, on rocks, 1,830 m., September, 1911, Foote 11. Bryum argenteum L. Cuzco, on dry rock, 3,500 m., 1911, Foote 12; Ollantaytambo, on soil, about 3,000 m., May, 1915, Cook & Gilbert 610. Bryum biforme sp. nov. Pseudoautoicous, the male plants minute, 1-2 mm. high, attached by radicles to the tomentum of the fertile stems, each plant bearing one or two flowers containing two or three antheridia, with few, somewhat longer paraphyses; the outer antheridial leaves broadly ovate-lanceolate, scarcely serrulate, with costa vanishing well below the apex: fertile plants with more or less branching, tomentose stems 3-4 cm. high, the short branches often bearing at the apex abundant, filiform flagella, 3-4 mm. long and covered with minute scale-like leaves; stem-leaves widely spreading when moist, somewhat spreading-flexuous and twisted when dry, serrulate above, rather broadly ovate, acute, slightly decurrent, 3-3.5 mm. long, the margins flat above, recurved below with a border of elongate, narrow, pale cells most distinct half Williams: Peruvian mosses 329 way down the leaf where it becomes four or five cells wide ; costa not quite percurrent, from a broad, reddish brown base, tapering gradually to the slender apex; cells in upper leaf rhomboidal or hexagonal, in lower leaf larger and more or less rectangular, all with somewhat thickened walls distinctly pitted, at least in the lower leaf; the median cells about 16/4 wide and 40-50 ix long; outer perichaetial leaves about the length of the stem-leaves, from a broad base, oblong, acute, serrulate and with costa van ishing a little below the apex; seta about 2 cm. long; capsule pendent with sporangium rather narrowly ovate and tapering into a neck about one half the length of the sporangium, the median exothecal cells irregularly elongate with evenly thickened, mostly slightly curving walls, the stomata abundant in the neck, about 40 n long; annulus large; peristome- teeth lanceolate, about 100 p wide at the base and 600 > high, with hyaline borders and the outer plates finely punctate, mostly one and one half to two times wider than high, the inner lamellae without cross- walls; inner segments about the height of the teeth, keeled and solid along the median line, very slender-pointed with mostly two long, slender cilia between them, sometimes distinctly appendiculate ; spores rough, up to 20 fi in diameter; lid highly convex-apiculate. (Plate 20.) Torontoy, Urubamba Valley, about 3,600 m., May, 1915, Hiram Bingham 730. This is, I believe, the only Bryum known having inflorescence as above described ; the flagella also are unusual. Bryum densifolium Brid. Urubamba, on damp earth, 2,900 m., July, 191 1, Foote 13; Santa Anna, 912 m., August, 191 1, Foote 14. Mnium ligulatum C. Mull. Lucma, on dead wood, 2,130 m., Aug. 1911, Foote 15, growing mixed with Cyclodictyon aeruginosum (Mitt.) Broth, and Tricho stomum brachydontium Bruch.; Urubamba, on wet earth, 2,900 m., July, 191 1, Foote 16. Anacolia subsessilis (Tayl.) Broth. Urubamba, on loose earth and rocks, 2,900 m., July, 1911, Foote 17; Ollantaytambo, about 3,000 m., Cook & Gilbert, May, 1915, 504, 749, 771a. Breutelia bryocarpa Herzog. Lucumayo Valley, 1,800-3,600 m., June, 1915, Cook & Gilbert, 1320a. 330 Williams: Peruvian mosses A few stems with one capsule were found mixed in with Breutelia tomentosa. This is evidently the same species as that collected by Weddell in Peru and named, apparently in manu script only, Bartramia nutans by Montagne. It is also the same as my Bolivian collection, No. 2825, referred to Breutelia nutans (Mont.). Breutelia tomentosa (Sw.) Schimp. San Miguel, Urubamba Valley, about 1,800 m., May, 1915, Cook &f Gilbert 988; Lucumayo Valley, 1,800-3,600 m., June, 1915, Cook & Gilbert 1320. Pogonatum campylocarpum C. Mull. Lucumayo Valley, 1,800-3,600 m., June, 1915, Cook & Gil bert 1322. Pogonatum polycarpum (Schimp.) Broth. Araranca, south side of La Roya Pass, about 4,100 m., April, 191 5, Cook & Gilbert 177 and 182. Polytrichum antillarum Rich. Torontoy, Urubamba Valley, about 2,400 m., June, 1915, Cook & Gilbert 1194; Lucumayo Valley, 1,800-3,600 m., on wet earth, June, 1915, Cook &¦ Gilbert 1318. Hedwigia albicans (Web.) Lindb. Temple of Viracocha, near Tinta, about 3,500 m., April, 1915, Cook & Gilbert 204; Ollantaytambo, about 3,000 m., May, 1915, Cook & Gilbert 501. Hedwigidium imberbe (Sm.) B.S.G. Urubamba, on rocks, 2,900 m., July, 191 1, Foote 18; Ollantay tambo, 2,750 m., July, 1911, Foote 19; Temple of Viracocha, near Tinta, about 3,500 m., April, 1915, Cook & Gilbert 204a. Braunia canescens Schimp. Near Calca, on rock, April, 1915, Cook & Gilbert 245; Ollan taytambo, about 3,000 m., April and May, 1915, Cook & Gilbert 473, 502, and 503. This species has been considered scarcely distinct from B. cirrifolia (Wils.) Jaeg., but it seems to me to be a rather smaller species with more closely imbricate leaves. The leaves also, below the apex of the stem, are broader in the upper part, often slightly obovate, and more abruptly narrowed to the pale apex of blade and hair point. B. cirrifolia has leaves rather Williams: Peruvian mosses 331 lanceolate and gradually narrowed to a yellowish apex and flexuous hair point. Braunia argentinica C. Miill. and B. cirri- olia var. falcatula Herzog belong, I should say, under B. canescens. Prionodon bolivianus C. Miill. Torontoy, Urubamba Valley, about 3,600 m., May, 1915, Hiram Bingham 729. Squamidium rotundifolium (Mitt.) Broth. Above San Miguel, 1,830 m., September, 191 1, Foote 20. Pilotrichella viridis (C. Miill.) Jacq. San Miguel, Urubamba Valley, 1,800 m., May, 1915, Cook & Gilbert 919. Papillaria imponderosa (Tayl.) Broth. Above San Miguel, on moist earth, 1,830 m., September, 191 1, Foote 21. Meteoriopsis remotifolia (Hornsch.) Broth. San Miguel, 1,500-1,830 m., July, 191 1, Foote 22. Neckera eucarpa Schimp. Urubamba, on bark, 2,900 m., July, 191 1, Foote 23; Ollan taytambo, about 3,000 m., May, 1915, Cook & Gilbert 763. Neckera Jamesoni Tayl. Ollantaytambo, about 3,000 m., on Alnus, May, 1915, Cook & Gilbert 763a. Neckera Lindigii Hampe. Lucma, on dead wood, 2,130 m., August, 1911, Foote 24; Urubamba, on bark, with N. eucarpa, 2,900 m., July, 1911, Foote 23a. Porotrichum valdiviae C. Miill. Ollantaytambo, about 3,000 m., on trunks, May, 1915, Cook & Gilbert 762a; apparently the farthest northern station for this species. Entodon platygyrioides C. Mull. Above San Miguel, on rock, 1,830 m., September, 191 1, Foote 25- Erythrodontium squarrosum (C. Miill.) Par. Santa Ana, on rock, 912 m., August, 191 1, Foote 26. 332 Williams: Peruvian mosses Fabronia andina Mitt. Ollantaytambo, on rock, about 3,000 m., May, 1915, Cook & Gilbert 662; Urubamba, 2,900 m., July, 1911, Foote 27; Ollantay tambo, 2,750 m., July, 1911, Foote 28. Fabronia polycarpa Hook. Santa Ana, 912 m., August, 191 1, Foote 29. Cyclodictyon aeruginosum (Mitt.) Broth. Lucma, on dead wood, 2,130 m., August, 1911, Foote 30. Rhacopilum tomentosum (Hedw.) Mitt. Santa Ana, on dead wood, 912 m., August, 1911, Foote 31, 32; San Miguel, 1,525 m., September, 1911, Foote 33. Leskea gracillima Tayl. Urubamba, 2,900 m., July, 191 1, Foote 34, on bark, 35 on rock; Ollantaytambo, on rock, about 3,000 m., May, 1915, Cook & Gil bert 609a, 663. Thuidium peruvianum Mitt. Ollantaytambo, about 3,000 m., May, 1915, Cook & Gilbert 704, 748. Hygroamblystegium irriguum (Wils.) Loeske. Urubamba, on rock, 2,900 m., July, 1911, Foote 36. Appar ently not before reported for South America. The next nearest known location is, I believe, Guatemala. Drepanocladus longifolius (Wils.) comb. nov. Amblystegium longifolium Wils.; Mitten; Jour. Linn. Soc. Bot. 12: 571. 1869. Temple of Viracocha, near Tinta, about 3,500 m., in a spring, April, 1915, Cook & Gilbert 216. These sterile specimens seem to belong under this species although only known from Patagonia up to the present time. Hygrohypnum peruviense sp. nov. Inflorescence unknown : growing in loose mats with irregularly branching stems without radicles, 2-3 cm. long and about 200 11 in diameter, in cross-section showing a distinct central strand and outer walls of about three rows of smaller, thick-walled cells; leaves loosely erect-spreading, more or less secund, rather broadly ovate-acute, somewhat cymbiform, 1.5-2 mm. long, and nearly or quite entire ; costa stout, 80 ix wide or more at the base, simple Williams: Peruvian mosses 333 and extending about three fourths up the leaf or shorter and dividing into two to five branches; leaf-cells all elongate, prosen- chymatose and mostly slightly sinuous, with rather thin walls, the median 5-6 fi wide and 40-60 fi long, the alar usually forming a distinct cluster of broad, short cells either greenish or finally reddish-brown; fruit not found. (Plate 18, figs. 7-12.) A variety evidently of the above, growing in flowing water, has elongate, hispid, wiry stems, 10-15 cm- long, without leaves below and in habit like H. Bestii, except much more slender. Cuzco, on wet rock, 3,500 m., July, 191 1, Foote 37 (type); Ollantaytambo, about 3,000 m., on rock under spring, May, 1915, Cook fif Gilbert 666 (the variety) . Ctenidium malacodEs Mitt. Ollantaytambo, about 3,000 m., on trunks, May, 1915, Cook & Gilbert 762b. Mittenothamnium andicola (Hook.) Card. Ollantaytambo, about 3,000 m., on trunks, May, 1915, Cook & Gilbert 762. Mittenothamnium oxystegum (Spruce) Card. Santa Ana, on rock, 912 m., August, 1911, Foote 38. Hypnum Schreberi Willd. Ollantaytambo, about 3,000 m., May, 1915, Cook & Gilbert 748a. Ectropothecium aeruginosum (C. Miill.) Mitt. Santa Ana, 912 m., on dead wood, August, 191 1, Foote 39. Palamocladium leskeoides (Hook.) E. G. Britton. San Miguel, on rock, 1,830 m., September, 1911, Foote 40. Brachythecium asperulum (Hampe) Jaeg. Ollantaytambo, about 3,000 m., May, 1915, Cook & Gilbert 7530- Brachythecium flaccum C. Miill. Huadquinia, on wet earth, 1,530 m., August, 191 1, Foote 41; Ollantaytambo, about 3,000 m., on trunks, May, 1915, Cook & Gilbert 762c. Brachythecium sericeovirens (C. Mtill.) Par. Ollantaytambo, on rock, 2,750 m., July, 1911, Foote 42. Known previously only from Patagonia. 334 Williams: Peruvian mosses Brachythecium stErEopoma (Spruce) Jaeg. ' San Miguel, on live wood, 1,830 m., July, 1911, Foote 48. Rhynchostegium aquaticum (Hampe) Jaeg. Urubamba, in water, 2,900 m., July, 191 1, Foote 43; Arequipa, in water, June, 191 1, Foote 44; Cuzco, on wet rocks, July, 191 1, Foote 45. Rhynchostegium conophyllum (Tayl.) Jaeg. Urubamba, on rock, 2,900 m., July, 1911, Foote 46; San Miguel, on live wood, 1,830 m., July, 191 1, Foote 47. New York Botanical Garden Explanation of plates 17-20 Plate 17 Leplodontium integrifolium. i. Plant about natural size. 2. Perichaetium, X 7- 3- Stem-leaf, X 15. 4. One side of base of leaf, X 115. 5. Median cells of leaf, X 240. 6. Apex of leaf, X SO. 7. Cross-section of leaf, X 150. Globulina peruviana. 8. Plant about natural size. 9. Plant, X 10. 10. Stem- leaf, X 45. n. Apex of costa and leaf, X 230. 12. Median cells of leaf, X 230. 13. Basal cells on one side of costa, X 230. 14. Cross-section of stem, X 230. 15. Cross-section of leaf, X 230. Plate 18 Tortula lacerifolia. 1. Plant about natural size. 2. Perichaetial leaf, X 25. 3. Upper stem-leaf, X 25. 4. A segment from the leaf-margin, X 130. 5. Base of leaf on one side of costa, X 180. 6. Cross-section of leaf, X 180. Hygrohypnum peruviense. 7. Plant about natural size. 8. Cross-section of stem, X 130. 9. Basal leaf-cells on one .side of costa, X 270. 10, n. Stem-leaves, X 25. 12. Median leaf-cells, X 270. Plate 19 Grimmia rivulariopsis. i. Fertile and male plant about natural size. 2. Cap sule, seta and perichaetial leaf, X 20. 3. Calyptra, X 20. 4. Lid, X 20. 5. Part of peristome and rim of capsule, X no. 6. Antheridia with leaf, X 20. 7. Upper part of stem leaf, X 180. 8. Stem-leaf, X 18. 9. Median cells of leaf, X 180. 10. Cells in lower part of perichaetial leaf, X 180. n. Perichaetial leaf, X 18. 12. Median exothecal cells, X 180. 13. Cross-section of stem-leaf, X 180. Plate 20 Bryum biforme. 1. Fertile and male plant about natural size. 2. Male plants, X 20. 3. Capsule, moistened, X 8. 4. Upper stem-leaf, X 16. 5. Outer perichaetial leaf, X 16. 6. Upper part of stem-leaf, X 135. 7- Border of leaf and adjoining cells about half way down, X 135. 8. Inner perichaetial leaf, X 16. 9. Stoma, X 135. 10. Median exothecal cells, X 135. n. Part of peristome, annulus and rim of capsule, X 100. Bull. Torrey Clue Volume 43 plate 17 1-7. LEPTODONTIUM INTEGRIFOLIUM R. S. Williams 8-15. GLOBULINA PERUVIANA R. S. Williams Bull. Torrey Club Volume 43, plate 18 1-6. TORTULA LACERIFOLIA R. S. Williams 7-12. HYGROHYPNUM PERUVIENSE R. S. Williams Bull. Torrey Club Volume 43, plate 19 GRIMMIA RIVULARIOPSIS R. S. Williams Bull. Torrey Club Volume 43, plate 20 'MOfM «i o^y io]p O^MV S§W»s BRYUM BIFORME R. S. William 9002 04055 8653