590 Eb3t CENTRAL CIRCULATION BOOKSTACKS The person charging this material is re¬ sponsible for its renewal or its return to the library from which it was borrowed on or before the Latest Date stamped below. You may be charged a minimum fee of $75.00 for each lost book. Theft/ mutilation, and underlining of books are reasons for disciplinary action and may result in dismissal from the University. TO RENEW CALL TELEPHONE CENTER, 333-8400 UNIVERSITY OF ILLINOIS LIBRARY AT URBANA-CHAMPAIGN AT JUL101998 When renewing by phone, write new due date below previous due date. L162 Digitized by the Internet Archive in 2017 with funding from University of Illinois Urbana-Champaign Alternates https://archive.org/details/textbookofnaturaOOebel TEXT BOOK OF atural History. by ADRIAN J. EBELL, Bli.B., M.D. PART x. ENDING TO THE STRUCTURAL FEATURES OF CLASSES IN TH ANIMAL KINGDOM. ebell & CO . Publishers of School and Popular Scientific Works, Room 18, Cooper Union Building, NEW YORK. Entered according to Act of Congress in the year 1870, by Adrian J Isbell, in the Office of the Librarian of Congress, at Washington. Stereotyped and printed by ^olhemus valves . Heart of two or three *4 lobes or valves. . . - Cold-blooded, or Haematocrya. Heart of four lobrs or valves. '] Wann-blooded, [> or J Haematotlierma. These four functions of Generation , Digestion , Respi¬ ration and Circulation , are called vegetative, because they are essential alike for the life of both plants and animals. Those of sensation and motion are peculiar to animals alone, and hence are called Animal Functions. SENSATION, 35 CHAPTEK XII. SENSATION. The special functions of sensation are those of— 1. Nervous Action, 2. Touch, 3. Taste, 4. Smell, 5. Hearing, 6. Sight. Nervous Action is the means by which the organs, or parts of the animal body, reciprocate with each other in the relations of sensation, or feeling and motion, whether voluntary or involuntary. In the lowest members of the animal kingdom this nerve, or vital power, is equally dis¬ tributed through all ..parts of the body, and is transmitted from one portion to another by the direct contact of the cells and organs, there existing no distinct sets of nervous organs. As we ascend in rank, however, distinct systems of nervous organs are found with increasing perfection, forming the chief basis of zoological classification , and the ladder on which the rank of the animal kingdom is arranged. The nerve systems of animals are arranged according to five plans, which, we shall hereafter find, give rise to the five great types or branches. 36 SENSATION. Table V. 1. Nervous system, without any distinct nerve organs. 2. Nervous system, consisting of a nerve thread around the stomach, with branching nerve filaments. 3. Nervous system of nerve bunches, called Ganglia, arranged chiefly about the stomach, and their connecting and branching nerves. 4. Nerve system of a row of nerve bunches, or Gang¬ lia, extending from one extremity of the body to the other, with their connecting and branching system of nerves, called the Ganglionic or Sympathetic System. 5. Nerve system, consisting of a governing nerve mass, of two parts (Cerebrum and Cerebellum), located in the head, from which extends the entire length of the trunk a nerve chain , called the Spinal Cord, with its branching sets of nerves / with which is associated the Ganglionic or Sympathetic System. The nervous systems may act independently, as in feel¬ ing, or in a special way peculiar to themselves, called Elec¬ trical or Magnetic, in connection with the Muscular Sys¬ tems, in motion, whether voluntary or involuntary, and in the several functions of life peculiar to the animal, and as agents in the higher faculties of volition, instinct, thought, and roason. FIVE SENSES. 37 CHAPTER XIII. ^ FI YE SENSES. The organs of special sense are those of Touch, Taste, Smell, Hearing, Sight. These are found by no means equally distributed through¬ out the animal kingdom; indeed, most of the lowest types of animals are almost, if not quite, destitute of them ; but as we ascend in rank, and find the nervous systems more developed, so also these organs of special sense keep pace with them in their development and perfect action. Touch Is the sense by which animals, through their surfaces, gain a knowledge of the bodies surrounding them, or the con¬ ditions in which they may exist. It is the physiological parent of the other senses, being not only the first educator of them in man, but the first one evinced among the lower animals. Thus, while of all the senses, it is the simplest and the least confined to any one part of the animal, and the most general in its action, V* it is at the same time the most direct, positive and certain of them all. 38 FIVE SENSES. It might be considered under the heads of Tactile Dis¬ crimination, Temperature and Muscular Sense. The first two, Tactile Discrimination and Temperature, are exercised through the surface of the body, and the third, Muscular Sense, through the action of the muscles. Taste Is the sense by which animals are enabled to perceive the differences of flavor in their food, and make a proper selection thereof. It is invariably located in the mouth, in the organ called the tongue, though the tongue is by no means always possessed of the function of taste. Among most of the lower animals the tongue is merely a prehensile organ for catching and holding the food, and is wholly destitute of the powers of taste. Among great numbers of the lowest members of the animal kingdom there is an entire want of any organ that may be called a tongue, and, in many cases indeed, of any mouth-organs whatever. Smell Is the sense by which animals are capable of taking knowledge of odors, and is chiefly a guide, as well as taste, in the selection of food. Most of the lower animals seem either to be destitute of it, or to have it merged with the functions of touch. Among the higher animals the nose and the nostrils are the organs of smell, and greatly vary in their form and development. Hearing Is the sense by which animals are enabled to receive and distinguish sounds. The organ of hearing consists funda- FIVE SENSES. 39 mentally of an elastic membrane to receive the vibrations of the air, and a set of nerves to impart a knowledge of those vibrations to the chief nerve centres. The lowest members of the animal kingdom are desti¬ tute of any organs of hearing whatever; and the highest members have a wonderful and delicate mechanism in the ear, comprised in three chambers, i.e., the anterior , middle , and posterior chambers. Between these two extremes exist organs of all degrees of complexity and perfection. Sight , Or vision, is the sense by which animals are enabled to re¬ ceive impressions from light. The organs of vision con¬ sist fundamentally of one or more transparent refractive membranes for lenses, to collect together the rays of light and form the image, together with an expanded nerve to receive it. In the lowest animals—when it exists at all, the entire eye consists of a mere speck, capable only of receiv¬ ing light, but not form or color. From this elementary form upward, we find all stages and degrees of develop¬ ment, to the wonderful visual organs of the birds and mammals. We do not propose to take up, at length, in this connec¬ tion the development of the organs of sense ; that would be the special work of a treatise on comparative anatomy and physiology. We would merely, in our methods of study, outline as it were, the several functions and organs to be taken into consideration in a review of animal me¬ chanism. Proceeding from the Organs of Sensation we would next sketch, in brief, those of Motion , 40 MOTION. CHAPTER XIV. MOTION. Motion , or movement in the animal mechanism, is of two kinds, Voluntary and Involuntary; and quite dif¬ ferent in its character from the movement of plants. It is executed through two sets of organs, first, Muscles , and second, Endo , or Exo- Skeleton, for leverage, protection and support; but in many of the lower animals, a muscu¬ lar system alone exists, and even that, indeed, often of a very weak and loose texture. The skeleton is either external, as the shell of the crab or lobster, or the hard skin of the insect, and is called Exo-Skeleton; or is internal, having the muscles attached to its exterior, as the bony skeleton of the vertebrae, and is called Endo-Skeleton. An outline of the model or Archetype Endo-Skeleton will be given in connection with the characteristic features of vertebrates. As has already been said, the structure of the animal is the basis of the classification adopted by this volume. CLASSIFICATION. 41 CHAPTEE XV. CLASSIFICATION. Thor© are two systems according to which plants and animals are classified: one called the Natural, and the other the Artificial system. The artificial system arranges plants and animals ac¬ cording to their appearance and external conditions, with¬ out reference to their structural relations, hut it is one of great uncertainty and ever subject to changes and varia¬ tions. It was adopted by the older zoologists, and hence we find their systems ever conflicting with each other. The end sought after by the artificial system, is merely to name objects, for which oftentimes the commonest points of external appearance are alone observed; and animals and plants that are, according to their structure, wide apart, are grouped together, and those that may be closely related are placed far asunder; thus answering the purposes of a dictionary or table of reference. The natural system, on the other hand, is founded wholly on points of structural resemblance and difference. Those animals that are alike in the most prominent fea¬ tures of structure, though they may differ in a number of lesser ones, it groups together in the great departments of Branches and Types, differing from each other in the most important features of their anatomy. Those of each of these types that resemble one another in some other lesser points of structure, but differ from a number of other 42 CLASSIFICATION. groups of the same type on those very points, it still further divides into classes, and these, by other successive grada¬ tions of resemblance and difference, into Orders, Families, Genera, and Species. This gradation of Types, Classes, Orders, Genera, and Species, Forms the nomenclature of the natural system of classi¬ fication. The last two (Genus and Species) combined together forms the specific name of an animal. The Generic pre¬ ceding the Specific name, as Smith John, instead of John Smith, e.g ., canis familiar is, the common or familiar dog, instead of familiaris cards. This method is one that greatly economises the number of words used in naming the animal, as comparatively a few names, variously com¬ bined and interchanged with each other, furnish distinctive terms for the hundreds of thousands of animals of which, if each were possessed of an entirely distinct name, the vocabulary would be far beyond the reach of any human intellect, and an acquaintance with the members of even an order or family an utterly hopeless task ; while now, a few names, that soon become familiar, are used over and over m different combinations, under the several larger groups, and so combine the utmost definiteness with the greatest simplicity. With these introductory remarks we will now enter upon the second head or department. METHODS OF STUDY. 43 CHAPTER XVI. METHODS OF STUDY, l.e ., CLASSIFICATION AND NATURAL HISTORY. Classification of the animal kingdom, or systematic zoology , relates to the arrangement of animals in their several groups of types, classes, orders, etc., according to their resemblance and difference of structure. Natural History is the science of descriptions relating to the appearances, methods of life, and habits of animals. Our methods of study shall be, first, to get a clear idea of the types and their points of distinction; next, of the classes they include; then, of the orders, and so on, noticing merely a few genera and species in connection with each of these divisions, to illustrate the structural peculiarities we may have under consideration. The learner would make much easier and quicker pro¬ gress in the attainment of an acquaintance, even with the details of classification of any special part of the animal kingdom, by first thoroughly mastering the distinctive fea¬ tures of the larger division of types, classes and orders, and this method, certainly, is of the first importance, if the two chief objects in the study of Natural History, spoken of in the first section of this volume, i.e., the de¬ velopment of our observing powers and learning the laws of nature, are the ends in view. We must, then, concisely review— 1. The special and distinctive features of all the types; 44 METHODS OF STUDY. 2. The classes of each type ; and 3. The order of each class. The compass of this volume will not, except in a few special cases, permit us to proceed beyond the sub-orders and families, and, indeed, not in all cases even beyond the orders. The animal kingdom, according to its structure, divides itself into five great types, or branches, i. e., beginning with the lowest, as we shall, in all cases: Protozoa, Radiates, Mollusks, Articulates, Vertebrates. € We have chosen to arrange our tables from the lowest upward, because we are thus enabled to study the struc¬ ture of the higher groups as taught synthetically, by the lower ones. These types are all constructed according to the same plan; though in the highest alone do we find the entire system brought out as given in our review of Structural Zoology. Each of these types, however, expresses a distinctive arrangement of the organs of the body, and is built on an idea or method exclusively peculiar to itself. That we may have clearly before us the method in which our studies of each of these divisions will be conducted, t we would again call to the mind of the student the table METHODS OE STUDY. 45 of functions and organs, as our studies of the divisions shall be conducted in relation to each of these, i. e.: Generation, Digestion, Eespiration, Circulation, Nerve Action, Organs of Sense, Organs of Movement, And also as regards their general appearance, habits, and places of residence. > 46 PROTOZOA. CHAPTER XVII. PROTOZOA.-GENERATION. The Type of Protozoa comprises the lowest and sim¬ plest organisms in the animal kingdom. In the lower orders, no definite organs exist, but the animals present, under the microscope, formless and shapeless masses of jelly, in which the various organs of the body are made for the occasion at the tim3 they are needed for use. Reproductio?i is non-sexual. No distinct sexes have been discovered among the Protozoa. They multiply ex¬ clusively by dividings (fissiparous), and by budding (gem- miparous). They comprise all the methods of fissiparous multiplication, i. e ., by the parent splitting transversely, longitudinally, or irregularly, or splitting open and dis¬ charging the young. Some of the Protozoa pass through several genera¬ tions in a few hours, and their spores or buds survive almost any exposure of heat or cold, and may be even sub¬ jected to chemical action, or be pulverised, without losing their vitality. Digestion among the Protozoa is of the lowest grade, and enacted by organs of the very simplest structure, con¬ sisting, among the greater number of them, of a mere cavity, or depression in the soft shapeless body, endowed with the vital power to dissolve the food taken into it. The entire alimentary canal consists of a mere sack, with PROTOZOA.-DIGESTION. 47 the same opening for entrance and outlet, and even this simple organ has to be made at the time of use among many of the lower orders; as the amoeba, sponge animals, etc., where, on the contact of the food, a bag or cavity is sunk into the body, or little filaments are projected as arms that encircle it, and so a stomach is formed of the part of the body where the food is floating. Several of these stomachs may be formed on different parts of the body at the same time, which, after having performed the special work for which they were extempo¬ rized, may again be merged and lost in the soft plastic tissue of the body, called Sarcode. This feature led Ehren- berg, the great microscopist of Berlin, to call this type Polygastria, or many-stomached. Some others of this type have even a simpler process of digestion ; subsisting entirely on liquid food, they have need of neither stomach nor digestive fluid, and so absorb their nutriment directly through the skin wall of the body. Others, again, are supplied with a more elaborate system, consisting of a permanent stomach, with tubes of inlet and exit. The existence and courses of these organs in such minute jelly-like animals, have been traced under the microscope, after they had been fed on colored liquids. Many of them—as the Stentor, Yorticella, Potifera— have numerous vibrating hairs or cilia around their mouth openings, that keep up a rapid vibration, and so sustain the circulation of a current of water through their stom¬ achs, and are thus supplied with the particles of decaying matter floating in the water that furnish their food. 48 PROTOZOA.-RESPIRATION-CIRCULATION. Distinct Respiratory and Circulatory Organs are usually wanting. The oxygen is absorbed from air, dissolved in the water, directly through the entire skin wall of the body. It is presumed that, in some of the higher orders, the vibratile cilia around the mouth answer, in a greater degree than the rest of the body, the functions of respira¬ tion. The circulating fluids are conducted in the lowest orders directly from cell to cell, through their partition walk ; though, in the higher, there are a few branching, circulat¬ ing tubes, through which, the fluids are conducted by the contraction and expansion of the body. PROTOZOA.-NERVOUS SYSTEM-ORGANS OF SENSE. 49 CHAPTER XVIII. PROTOZOA.-THE NERVOUS SYSTEM AND ORGANS OF SENSE. Among the Protozoa no distinct nervous organs have been found, although they are evidently possessed of a distinct nervous action and the functions of some of the organs of sense. The vital power is equally distributed through all parts of the body, to such an extent that the animals will con¬ tinue ther existence, unharmed, even if cut into a number of pieces ; each piece readily fashioning itself into and taking up the life of a distinct animal. All the organs of sense among them are wanting, yet they evidently are capable of the sense of touch ; but the entire lining of the body is equally capable of it, except among the higher orders, which have many hairs or cilia around the mouth, that seem to be possessed of a greater degree of tactile or sensitive power than the rest of the body. Some of the higher members of the type seem to be sensitive to light, but their powers of vision extend no further. Ehrenberg describes some red spots on the lin¬ ing of their bodies as rudimentary eyes, but this is not yet established as a fact. They evince a choice in the selection of their food, but this cannot be attributed to anything like a sense of taste, but rather to a vital principle, the same as that which guides the plant to a selection of its food. That they are possessed, in any degree, of the functions of smell and hearing we have no evidence. 50 PROTOZOA.-MOTION Motion , among a great many of this type, is active and constant, although only a few of them are possessed of even a rudimentary locomotive apparatus. They are all of them devoid of anything like a skeleton, either external or in¬ ternal ; although many of them secrete hard shells, or cases of lime or flint, on the exterior of their bodies, which aggregate and form masses, often of great extent. Yet this hard covering is merely a protective envelope, and those occupying it are usually fastened to one spot. Except among some of the higher orders, which have those vibratile cilia we spoke of, even distinct muscles are wanting, and the little animal moves by a contraction and expansion of its entire jelly-like mass, or the incessant vibration of many little hairs on all parts of the body. PROTOZOA.-NATURAL HISTORY. 51 CHAPTEK XIX. PROTOZOA-NATURAL HISTORY. The type of Protozoa includes countless numbers of microscopic animals, most of which are quite invisible to the naked eye. Their numbers, however, exceed all com¬ putation. Of the hard shells, of those that secrete limy or flinty coverings, immense strata of rock have been formed, tens and hundreds of feet in thickness, and thousands of square miles in extent; which, by subsequent wrinkling, have formed lofty mountains. The ocean, in its broad expanse and depth, and all its branches and streams ; the lakes and rivers of the land; the ponds and little runnels and ruts, and even the mud-puddle or little pocket contained in the hollow of the rock or a decaying log, or, indeed, any place where we may find a drop of water, if there is the least decaying organic matter in it, is found to contain countless hosts of these Protozoa. They are the first animals that were created on the sur¬ face of the earth, for we find their remains in the lower¬ most strata that contains animal fossils. Although of insignificant size, they are of immense importance in the life or economy of the earth ; not only do the shells of the hard covered ones form great layers of rocks, but the myriads of formless, organless, jelly-like atoms that swarm in all waters that contain the least decaying organic matter, act as scavengers, ridding the waters of the little 52 PROTOZOA.-NATURAL HISTORY. putrid masses, that else would render them unfit for sustain¬ ing higher kinds of life. The Protozoa all live, only in the water, although their bodies will sustain an indefinite amount of injury without loss of life. The ponds in which they swarm may be dried up, and the dust made up of their bodies blown abroad over the land, and subjected to a hundred various changes, yet, on the first application of sufficient moisture, they will again come to life and be as active as ever. The type of Protozoa includes such animals as the vari¬ ous kind of Sponges, Chalk Animals, Flint Animals, Amoeba, Vorticella, etc., which we shall consider hereafter. TYPE OF RADIATES.-GENERATION. 53 CHAPTEE XX. TYPE OF RADIATES. The type of Eadiates is characterized from all others, by having the organs arranged around a common centre, from which the branches or segments of the body radiate, usually in multiples of five. They are all of them inhabi¬ tants of the water, and nearly all of the ocean or salt and brackish waters. The vital organs of the animals are all soft, but many orders of them secrete and deposit struc¬ tures of lime within their bodies, forming masses called corals, or build either a scaffolding or frame-work, amid which the animal lives, as the star fishes ; or else constructs a shell or case around their bodies, composed either of lime, and studded with pencil-like spines, as in the Sea Urchin, or as in the Sea Cucumber, or Synapta, a cartilagenous shell, sometimes even garnished with hooks. We may now proceed to a review of the structure of the type of Eadiates, commencing with that of the organs of generation. Generation. The Eadiates increase in three ways : 1st By dividing into segments or parts. 2d. By budding. 3d. By emission from their bodies at certain times of the year of great numbers of spores, or ciliated round masses, resembling eggs. 54 TYPE OF RADIATES.-DIGESTION-RESPIRATION. The last two are the most prevalent methods of multi¬ plication, and are of extreme importance m the life and economy of the animals in the ocean. The little spores, or detached buds, float freely in the water, and so reach the places most conducive to their life and growth, and then, as they develop, either continue moving about in that region or else fix themselves permanently to the rock, and increase to an indefinite extent by buds from all parts of their surface. Multiplication by division is comparatively rare among them. Those that increase by spores or eggs are always bi¬ sexual, though both sexes may be either united in the same individual, or exist on separate ones. Digestion. The digestive apparatus of the Radiates is extremely simple in all except the highest class. It usually consists of a mere sack, with but one opening, into which the food is drawn by numerous arms or tentacles placed around it. These tentacles are endowed with great contractile power, so that they can be drawn back into the body or projected out to great length, at the option of the animal. They are also often armed with a powerful sting, and an acrid liquid that is capable of producing, even on the human surface, a smarting, and in some cases a severe inflammation ; but, among the animals on which they feed, it acts as a paralyzing or deadening agent. The bodies of some appear as mere sacks or stomachs, with the few soft radiating organs of the body arranged around. Respiration among the Radiates is altogether and invari¬ ably aquatic. None of them are capable of breathing in type: op radiates.— a iiteuLATiotf. 55 the atmosphere. The oxygen is taken from the dissolved air in the water either by the entire skin wall of the body or by the numerous tentacles that surround the mouth. Circulation is carried on by a scarcely more complex sys¬ tem than among the Protozoa. There are tubes that branch out from around the stomach and convey the chyle or digest¬ ed food to all parts of the body, but there is no distinct sys¬ tem of arteries and veins, nor is the fluid propelled through its tubes by means of a muscular heart. 56 RADIATES.-NERVOUS SYSTEM-ORGANS Of SENSE. CHAPTER XXI. RADIATES-NERVOUS SYSTEM AND ORGANS OF SENSE. A delicate nerve filament surrounding the mouth or stomach, opening and sending off branches into the several parts of the body, especially where the tentacles or arms are fastened, is the only rudimentary set of nervous organs found among the Radiates. And the Organs of Sense are in a similar state of devel¬ opment. The sense of touch apparently is equally distri¬ buted over the body. Among the members of the highest class, at the ends of certain nerve-lines, are organs which are capable of taking knowledge of light, but not of form or color. The remaining functions and organs of sens'; seem to be entirely wanting. The Organs of Locomotion are distributed under a wide variety of forms. Many of them, as the coral animals, are anchored to one spot during the whole of their ex¬ istence, and only such of their numbers as are thrown off from the parent animals in the form of spores, are capable of swimming around and changing their habitation, and that, only during the earliest stages of their lives. Others, as the Star Fishes and Sea Urchins, are only capable of crawling along, slowly and deliberately, at the bottom of the water, over the rocks and rubbish, amid which they live ; while yet others again, as the Jelly Fishes, dance along through the waters or float lightly upon its surface. RADIATES.-NATURAL HISTORY. 57 The special set of organs requisite for each of these will be taken into consideration in connection with the several classes of Radiates The solid formation within the coral animals can scarcely be called a skeleton, as it is more properly a deposit in the body as it grows old. In the highest kind of Radiates alone do we find what may be classed as a skeleton, in which the growing walls of the Sea Urchin and Star Fish, furnish the first instance of a growing skeleton. Among most of the Jelly Fishes, however, neither a growing skeleton or even a living de¬ posit is found ; but the body consists of a mere mass of jelly, in which not even muscular fibres are visible, and which perform their motions by a simultaneous contrac¬ tion and expansion of their entire body or disk. The higher classes, however, which have distinct arms, are possessed of delicate masses of muscular fibres. The natural history of the Radiates, as we have already observed, pertains entirely to the water. Countless myriads of them have inhabited the ocean, ever since the first appearance of animal life on the earth ; they swarm in the seas of all latitudes, and fill the two important parts in the economy of the ocean life, of purifying the waters and of furnishing food for hosts of higher animals. 58 TYPE OF MOLLUSKS.-GENERATION. CHAPTER XXII. TYPE OF MOLLUSKS.-GENERATION AND NUTRITION. Members of the type of Mollusks are distinguished from the two preceding types, by the greater distinctness of their nervous system and organs of sense, by the per¬ fection of the circulatory and breathing systems, and the first introduction and simplest evincement in the animal kingdom, of the dual or double arrangement of the struc¬ ture, called the “ bilateral system ,” in which all the chief organs of the body are arranged in pairs. Their general structure will present itself in the follow¬ ing review of Molluscan organization. Generation is bi-sexual, although both sexes are, among many species, combined in the same individual. In passing up to the Mollusca, we lose sight altogether of every trace of Gemmiparous and Fissiparous reproduc¬ tion. We have no illustrations of their increase by divi¬ sion into segments, or by budding. They are produced wholly by eggs, which, in a few cases, however, are de¬ posited within the body of the parent and retained within the shell until hatched, and thus living young are given forth. The eggs of some are distinct, and protected by a shell that is either albuminous and flexible, or calcareous and brittle. Others again connect their eggs into masses, or spread '.TYPE OF MOLLUSKS.—-NUTRITION. 59 them out in the form of a strap or ribbon, coiled up spirally like a watch-spring, attached by one of its edges. Some enclose their eggs in numbers together, within little tough capsules, which they string together on a chain, or cluster around a stem, like bunches of grapes, or arrange in radi¬ ating masses. Most usually, the young are passed from the egg in a form not in the least resembling the parent, and, after a series of changes, in the adult state alone are like their parents in all points. Such immature changes, through which the young of many animals pass, is called the larval state , and the young themselves the Larva Digestion , and its attendant functions of respiration and circulation, especially the last, are in a much higher degree of development among the Mollusca than in any of the preceding types. Among some, the mouth is unarmed, except by soft feelers, but among others, powerful and sharp jaws flank it on either side. We would observe, before speaking of what is called the lingual tube of some of these Mollusks, that among the invertebrates, the word tongue is used to designate any prehensile organ of the mouth, without reference to its being the organ of taste. Among many of the Mollusks, a tube extending back from the mouth, and lined its whole length with teeth, number¬ ing in some instances several thousands, is called the lin¬ gual-tube. These teeth assume all varieties of forms as well as numbers. ►Sometimes the tongue forms a short semicircular ridge 60 TYPE OE MOLLTTSXS.-NUTRITION. contained between the jaws, at others it is extremely elon¬ gated, and its folds extend backward to the stomach, while yet in others its folds are longer than the whole body. Some Mollusks are possessed of organs resembling the gizzard of a fowl, in which the food is received and ground. Others are yet further supplied with a crop, in which the food is received and held as it is swallowed until subjected to digestion. The liver is always large in the Mollusca; its secretion is derived from arterial blood, and is poured either into the stomach or the intestines. Many are even furnished with developed salivary glands, and some have a rudimentary pancreas, and also special glands for the secretion of colored liquids of the most delicate tints. MOLLUSKS.-RESPIRATION AND CIRCULATION. 61 CHAPTER XXIII. MOLLUSKS.-RESPIRATION AND CIRCULATION. Respiration among the Mollusks is of both kinds— aquatic and aerial; among those that live in the water it is accompanied by pairs of gills, called branchiae, which are exposed to the changing water, either by merely floating in it, or by having the water led in and out through a special pair of tubes, called siphons, which are drawn out to considerable length in those that live in the sand. Others breathe directly through their mantle or covering. The depth of water in which the Mollusks live is fixed partly by the amount of oxygen they need, and partly by the adaptation of the body to endure pressure. The air-breathing Mollusks are possessed of lungs, or air chambers, formed by the folding back of the mantle, in the interior of which are arranged the breathing organs, consisting of merely a permeable set of air bags, and a circulatory system to supply and remove the blood, the air being renewed with sufficient rapidity merely by the law of diffusion. The respiratory system is of very great importance in the economy of the Mollusks, and a fundamental point in their classification. Circulation is carried on among the Mollusks by a sys¬ tem of organs of much greater development than among either of the lower types. 62 MOLLUSKS.-NERVOUS SYSTEM. This type is not possessed of any distinct absorbent sys¬ tem, but the product of digestion (chyle), passes into the general abdominal cavity, and thence into the numerous veins of the body. They are possessed of a distinct heart of two chambers, arteries and veins, and so have a circula¬ tory system of much greater development than the Protozoa and Padiates, and, indeed, than even the larger number of the Articulates. The Nervous System. Among the Mollusca, the chief portion of the nervous system consists of a ring of nervous matter surrounding the throat, thickened out in places into nervous bunches, called ganglia, and from which proceed nerve-cords to all parts of the body. Organs of Sense. Every one of the organs of sense exist among some of the Mollusks, though in various degrees of perfection. Their eyes vary from mere spots on the sides of the mantle, that are scarcely capable of taking knowledge of light, as among such as the Oyster, to the most elaborate visual organs of the Cuttle Fish. The lowest of the type do not evince any organs of hearing, but among the higher orders the middle chamber chiefly is developed. The sense of Smell is evidently possessed by the Cuttle Fishes, and by the class to which the Snails and other uni¬ valve shells belong. Snails seek and select their food by this sense. Slugs are attracted by special odors, and many of the ocean Mollusks may be taken with animal baits. The sense of Taste is also indicated rather by the habits of the animal, and their choice of food, than by the struc¬ ture of a special organ, for, as we have already observed, MOLLUSKS.—OKGANS OF SENSE. 63 that which is called the tongue, is an organ intended for the seizing and preparing of the food, and not for the functions of taste. The sense of Touch , in the lowest class, is equally, dis¬ tributed over the entire covering of the body, called the mantle. Such is the case among the Clams, Oysters, and other bivalves; but, among the higher classes of the type, special organs, called tentacles, are appropriated to it. Thus all the Snails and other univalves have flexible feelers, or arms, from the head; two of which are for the purposes of touch, while the other pair support the eyes. Among members of the highest class there exist several arms around the head, which are endowed with organs of touch. Thus we find the special organs of sense, as well as the general structure of the nervous system, among the type of Mollusks is altogether on a higher plan than that of either of the two lower types, the Protozoa or Padiates. 64 MOLLUSKS.-INTEGUMENT. CHAPTEE XXIY. MOLLUSKS.-INTEGUMENT OR SHELL OF. We next proceed to consider the organs of locomotion and of protection, among the type of Mollusks. The muscles are mostly attached directly to the skin, or mantle, of the body, and are of rather a flaccid char¬ acter. A few muscles, however, are always of a more solid and compact character. Such are the strong muscles that, among the Oysters and Clams, connect together the two valves of the shell, and open and close them at will, and the projecting foot of the Snail, or the powerful prehensile arms of the Cuttle Fish and Nautilus. All the Mollusks are devoid of an endo-skeleton, except the Cuttle Fishes, where is found the first appearance of anything like an internal organ, of support and protection in the Cuttle Fish bone. Most of the Mollusks manufacture, however, an exo- skeleton , or shell, as it is called, which is constructed in a wonderful and delicate manner on the outside of the body, as it enlarges. The mantle, or skin covering of the body, is the organ that builds the shell. Its structure is commenced around, the young animal before it is passed from the egg-case MOLLUSKS.-INTEGUMENT. 65 and is enlarged and thickened as the animal grows larger. Molluscan shells are composed of three layers or parts : 1. Limy layer. 2. Nacreous layer. 3. Periostracum. The limy layer is the greater part of most shells. It is composed of carbonate of lime, and built along the outer side and edge as the animal increases in size. These layers are added one to the other, as may be seen in a transverse section of an onion, and each of these layers are composed of a series of closely fitting columns, usually hexagonal, which, themselves, are again composed of a series or pile of plates. The nacreous layer is deposited on the inside of the living shell by a liquid secreted by the body. It is com¬ posed of very delicate plates, laid partly over each other, and presenting sometimes the appearance of a coat of por¬ celain varnish, at others that of a sheet of burnished silvery metal, while yet others are ire descent, with a beautiful play of colors. This layer answers as a protecting cushion for the soft body that lies within it, and forms what we are so exten¬ sively acquainted with in the arts, under the name of mother-of-pearl, and is also used as a lining for any foreign object, as a grain of sand ; or a wound that may happen within the body of the animal, and this forms, by the addition of layer above layer, what we call a Pearl. The periostracum or epidermis is an outer coat of animal matter of various appearances; on some it is 66 MOLLUSKS.-INTEGUMENT. thin and transparent, on others again, quite thick and opaque. It is thick, leathery and olive-colored in all fresh water shells, and all Arctic sea shells. It gives color to a number of land shells. On some shells it is silky, or fringed with hairs ; on others it is thick and rough, like coarse cloth ; and on yet others, drawn out into long beard-like filaments. The periostracum is a living part of the shell, and protects it against the injury of water-wear and chemical action. After death, however, it soon cracks and crumbles away, often exposing beneath it a most exquisitely colored and variegated surface. This coloring of the shell is effected by little paint bags placed along the edge, from which the patches and lines of pigment are interwoven into the shell as it is built. Those shells, particularly of the tropics, that live nearer the surface of the water, are the most beautifully colored. Having thus reviewed the chief structural peculiarities of the type of Mollusks, we shall now proceed to say a few words in relation to their Natural History , or general economy. They are the first air-breathing animals we have yet, in our studies, reviewed. They have existed in the oceans from the earliest zoological ages, and have continued in immense numbers ever since, forming in some places, of their shells alone, layers of rocks hundreds of feet thick and scores and hundreds of miles in extent. MOLLUSKS.-INTEGUMENT. 67 They are more intimately connected with the economy of human life than either the Radiates or Protozoa. Their bodies are extensively used as food, and their shells for the manufacture of articles of use and orna¬ ment. 68 ARTICULATES.-ARRANGEMENT OF RINGS. CHAPTEE XXY. ARTICULATES-ARRANGEMENT OF RINGS. We will next proceed to consider the distinctive features of the type of Articulates. The type of which we are now to speak comprises by far the greatest number of species in the entire animal kingdom. They swarm in countless myriads in the ocean, on the land, under its waters, and in the air; and in places that seem absolutely untenable, even to the lowest kinds of animal life, as the interior of the bodies of other living animals, and the springs of boiling water. Nor is their structure less varied and interesting; according to their diverse habits and habitations, their forms assume an infinite variety of modifications, though all are built, as the other types, on one general plan. It is of this model, or general structure, that we will now proceed to speak. The Articulates are distinguished by the body being composed of a chain of rings. There are three plans, or methods, of arrangements of these rings: 1st. A uniform chain of rings, as among the worms. 2d. Eings arranged in cephalo-thorax and abdomen, as in Lobsters and Spiders. 3d. Eings arranged as among insects, in Head, Thorax, Abdomen. ARTICULATES.-ARRANGEMENT OF RINGS. 69 1. Those of the first plan merely have the rings of the body attached to each other, as a string of beads ; and, among the lowest of these, all the rings are possessed of an equal amount of vitality, so that, if they be severed into as many pieces as there are rings in the body, each piece might reproduce itself, or grow into a complete animal, such as the Tape Worm. 2. Those in which the rings are arranged into a ceph- alo-thorax and abdomen , have the rings of the front welded more or less together, into one piece, on vdiich are located the organs of the mouth, of the senses, and of locomotion. 3. The arrangements of the rings into the head, thorax and abdomen exists among the highest members cf the Articulates. The head is possessed of the largest nerve mass of the body, the organs of the senses and of the mouth. The thorax is axis of the body, composed usually of three rings, joined closely together, and to it are fastened the organs of locomotion, both wings and legs. And the abdomen is composed of rings, connected together with a flexible skin, or membrane, and possessed of the chief organs of digestion, respiration, circulation, generation and oviposition. t 70 ARTICULATES.-NUTRITIVE GROANS. CHAPTEE XXVI. ARTICULATES-NUTRITIVE ORGANS. We will now proceed to study the Articulates in relation to their structure. Reproduction is chiefly by means of eggs; the sexes in most cases are distinct; the young is passed from the egg usually in an immature condition, called larva, which often passes through many stages of transformation before reaching the adult state. Some will multiply themselves by breaking or dividing into parts— fissiparations. Such is the case with the Entozoa or intestinal worms. The Articulates multiply with extreme rapidity. Digestion among the Articulates has a wider range of action than among any other animals. As members of this type are found in nearly every place, they are also capable of taking and digesting every kind of food that any animal can live on. There is, accordingly, a great variation in the structure of the digestive organs; from a mere sack, without any special openings of entrance or exit—absorbing its nutri¬ ment like a plant cell from its entire surface, as in the intestinal worms—to an elaborate alimentary canal, con¬ sisting of mouth and prehensile organs, oesophagus, stom¬ ach, intestines, liver and other auxiliary organs, as among the insects, which we shall consider at length when we come to a reviewal of that class. Respiration among the Articulates is both aquatic and aerial. Myriads of them spend their entire brief existence ARTICULATES.-NUTRITIVE ORGANS. 71 in the water; as many others again, are exclusively deniznes of the air ; while yet others live in, and breathe water dur¬ ing their earlier life or larval condition, and are exclusively air-breathing during their adult state, as certain insects, such as the Dragon Fly, Mosquito, Gnat, etc. The respiratory organs are essentially of three kinds : 1st. Among the lower orders of the worms, the entire surface of the body acts the part of a breathing organ ; such are always inhabitants of the water, or of moist places. 2d. The body is provided with tubes, which are either— as among some of the higher worms—turned outward, and fastened about the body in tufts, or are turned inward, and branch as spiracles, through the rings of the body. Such we find to be the case among the insects. 3d. In the abdomen exist air-bags called lungs, as among the Spiders ; these are, however, different from the lungs of Vertebrates, in that they are simple sacks, and not collections of air and blood cells. Circidation is less complete in this type than among the Mollusks. The muscles and organs of the body are con¬ stantly bathed in blood, and even the insects have not any definite circulating tubes, as arteries and veins. A contractile tube along the back of the body answers to a heart, and moves the blood from one part of the system to another. The blood, however, among some of the active insects, as the Bee, is of a warmer temperature than among any of the lower types, and, indeed, even than among many of the higher types, as Fishes, and other cold¬ blooded Vertebrates. 72 ARTICULATES.-THE NERVOUS SYSTEM. CHAPTER XXYII. ARTICULATES-THE NERVOUS SYSTEM. Although the organs of circulation, among the Articu¬ lates, are of a lower grade than among the Mollusca, the greater development of their nervous system entitles them to a higher rank in the animal kingdom. The nervous system consists of a nerve cord or chain, extending the entire length of the body, along which, at regular distances, are fastened nerve-centres, or masses, called ganglia, and about each ganglionic mass is con¬ structed a ring or segment of the body, through which the nerves of feeling and motion branch from the centre gang¬ lion. The nerves of feeling, however, even among the Articulates, are but in a rudimentary condition. These animals, as well as all the other Invertebrates, seem almost destitute of the sensations of pain. It is true that in poetry we are told, “ The poor beetle “ that we tread upon, in corporeal sufferance, feels a pang “ as great as when a giant diesbut anatomy and true philosophy tell us differently: “ Pain, nature’s kind har- “ binger of mischief, is only inflicted for wise and import- “ ant purposes; either to give warning of the existence “ of disease, or as a powerful stimulus, prompting to “ escape from danger.” These animals, incapable of remedying the one or avoid¬ ing the other, not only ever exposed to destruction and death, but created for the purpose of preying on each other ARTICULATES.-ORGANS OF SENSE. 73 by tlie myriad, are kindly denied the feeling of acute pain. Their nervous organization still further denies the existence of it, save in a low degree. The central chain of ganglia, among the Articulates, can perceive external impressions and originate motion, bat not feel pain ; that is, a perception of the brain , and may be acute, as the body is under a special nerve mass located in the head, as among the Vertebrates. How far the feeling of pain is actively developed in these animals may be deduced from every-day observation. The Fly, seized by the leg, will leave its limb behind and alight, with apparent unconcern, to regale upon the nearest sweets within its reach ; the Caterpillar seems to enjoy a tranquil existence while the larvae of the Ichneumon hatched in its body devours its very viscera; and in the Crustacea, of so little importance is the loss of a leg, that the Lobster will throw off its claws if alarmed by the report of a cannon. The arrangement of the nervous sys¬ tem among the Articulates is the reverse of that among Man and the other Vertebrates. Organs of Sense. Organs of Sense , though of a lower grade than among the Vertebrates, are, in the highest orders, of a more elaborate and delicate structure than among any of the preceding types. There is, however, a great range in the development of these organs, from the lower worms to the insects; and as we shall treat of each of these, somewhat in detail, in connection with the classes of Articulates, our remarks in relation to them here shall be brief. The sense of Touch among the two higher classes is seated chiefly in feelers from the head, called Autense, Among 74 ARTICULATES.-ORGANS OF MOTION. the lowest class of worms, what there is of the sense is distributed over the entire surface of the body. Taste evidently exists among many of this type, for they evince a choice and selection in their food ; but, as among the Mollusks, the tongue is not usually invested with the sense of taste, but is moie generally a prehensile organ for the purpose of catching and retaining the food. The senses of Smell and Hearing are both in a rudi¬ mentary condition, and the organs, through which they act, undetermined, though usually referred to the antense or palpi of the head. Smell, however, seems to be more generally active than hearing, as it is active in the search and selection of food. The sense of Sight among the Articulates varies from the eye speck, scarcely capable of perceiving light, to elab¬ orate systems of simple and compound eyes, as in the heads of insects, while such as the Intestinal Worms are wholly destitute of organs of vision. Simple eyes, are single eyes placed on the back of the head, in numbers from one to about twenty; such are the eyes of Spiders, Lobsters and the young or larvae of insects; and they are also possessed, in connection with compound eyes, by many insects. Compound eyes consist of a pair of globes, one on each side of the head, composed on the surface of facets, or lenses, like a multiplying glass, varying in number from about fifty to ten or twenty thousand in each globe. The internal structure of these shall be observed in connection with the study of insects. ( )rgiins of Moti 4 COOPER UNION BUILDING, n NEW VORK. POPULAR LECTURES ON Illustrated with an immense collection of Zoological, Botanical, Mineralogical, Geolog¬ ical and Anatomical siREOinvnisnxrs, CASTS AND MODELS, jFrcc-Siaml lUacliboat'd fhauing, AND MAGNESIUM LIGHT AND Hydro-Oxygen Light Screen Microscopes, WOULD BE GIVEN BEFOBE LYCEUMS, LECTURE, LIBRARY AND CHRISTIAN ASSOCIATIONS, &c., ADRIAN J. EBELL, PLB., M.D. For particulars address, DR. EBELL, Room is. Cooper Union Building, NEW YORK. r UNIVERSITY OF ILLINOIS-URBANA 3 0112 042405529 A