A SYSTEM OF ANATOMY Foil THE USE OF '* STUDENTS OF MEDIC.- I BY CASPAR WISTAR, M. I). LATE PROFESSOR OF ANATOMY IN THE IT NITERS ITT -OF i'EJVNSYLTANI.'. FOURT H EDITION. WITH NOTES AND ADDITIONS- BY WILLIAM EDMONDS HORNY ;N M. X). AT -JIT^T PROFESSOR OF AWAT03I." IN THE TjSm-AS! ~ ■ ^ 7 ^A, MEMBER Ot THE A3lERICA:-r PHi'.':?0 T, ni' . Of — IN TWO VOLUMES — VOL. II. PHILADELPHIA: CAREY, LE 4 i- r ' P .THE l ; UT STREE 1827 M ■ as • ♦ EASTERN DISTRICT OF PENNSYLVANIA, TO WIT; BE IT REMEMBERED, that on the twentieth day of August in the forty-ninth year of the Independence of the United States of America, A. D. 1825, H. C. Carey & I. Lea, of the said district, have deposited in this office the title of a book, the right whereof they claim as Pro- prietors, in the words following, to wit: cc A System of Aanatomy for the use of Students of Medicine, By “ Caspar Wistar, M. D. Late Professor of Anatomy in the Universi- ty of Pennsylvania. Third edition. With notes and additions. By “ William Edmonds Horner, M. D. Adjunct Professor of Anatomy in the University of Pennsylvania, and member of the American Philosophical Society, &c. In two volumes. Volume II . 55 In conformity to the act of the Congress of the United States, entitled. An Act for the Encouragement of Learning, by securing the copies of Maps, Charts, and Books, to the authors and proprietors of such copies, 'during (lie times therein mentioned.” And also to the act, entitled, .' t supplementary to an Act, entitled “ An Act for the Encouraged^^^ of Learning, by securing the copies of Maps, Charts, and Books, ••.•‘hors and proprietors of such copies during the times therein mentionM^^ ml extending the benefits thereof to the arts of designing, engraving, and < r 1 ,ing historical and o,her print?.” D. CALDWELL, Clerk of the Eastern District of Pennsylvania. CONTENTS OF VOL. II. PART VI. OP THE NOSE— THE MOUTH— AND THE THROAT. CHAPTER I. Of the Nose. SECTION I. Op the External Nose - SECTION IL Of the Cavities of the Nose CHAPTER II. Of the Mouth and the Salivary Glands. C 7 the Mouth - = 14 Of the Salivary Glands ------ 25 CHAPTER III. Of the Throat. SECTION I. Of the Isthmus of the Fauces SECTION II. Of the Larynx - Of the Thyroid Gland - SECTION m. Of the Pharynx - 29 31 37 37 IV • •ox TEXTS. PART VII. OF THE THORAX. CHAPTER I. Of the General Cavity of the Thorax. SECTION I. Of the Form of the Cavity of the Thorax 44 SECTION II. Of the Arrangement. of the Pleurae Preparation of the Thorax - 45 47 CHAPTER II. Of the Heart and the Pericardium, and the Great Vessels connected with the Heart. SECTION I. Of the Pericardium - ■ - 49 SECTION II. \ Of the Heart ------ 50 SECTION III. Of the Aorta, the Pulmonary Artery and Veins, Venae Cavae at their commencement and the 59 CHAPTER III. Of the Trachea and Lungs. SECTION L Of the Trachea The Black Glands on the Bronchia; 62 64 SECTION II. Of the Lungs Thorax of the Foetus Physiological observations, &c. ... 65 69 71 OOMTENTSo V PART VIII. OF THE abdomen, CHAPTER I. A general view of the Abdomen and Pelvis , aim their Con- tents ; with an Account of the Peritoneum . SECTION I. Construction of the Abdomen - - - - SB Contents of the Abdomen - - - 85 SECTION II. Of the Peritoneum - - - - 90 CHAPTER II. Of the (Esophagus , the Stomach, and the Intestines . SECTION I. Of the (Esophagus - - - 94 SECTION II. Of the Stomach - - ' J 96 0.f the Gastric Liquor - - - ■ - 10; SECTION III. Of the Intestines - 104 Division of the Intestines . - - 109 The Small Intestines - . - 110 The Duodenum - - . - 111 Jejunum and lieon - - . - 113 The Mesentery - - - - ■ 115 Ot the Great intestines - - - -> 117 The Caecum and Colon ... - - 118 The Rectum - *, • 122 The Omentum - • . 125 CHAPTER III. Of the Liver , the Pancreas , and the Spleen. SECTION I. Of the Liver - _ „ _ , * „ iQS Vi CONTENTS. SECTION II. Of the Pancreas 141 SECTION III. Of the Spleen 145 CHAPTER IV. Of the Urinary Organs , and the Glandulse Renales. SECTION I. Of the Glandulse. Renales - - - - 154 SECTION II. Of the Kidneys and Ureters - - - - 155 SECTION III. Of the Urinary Bladder 162 CHAPTER V. Of the Male Organs of Generation . SECTION I. Of the Testicles and their Appendages = - * 171 SECTION II. *)f the Vesiculae Seminales and the Prostate Gland - 181 SECTION III. Of the Penis - ■■ -184 CHAPTER VI. Of the Female Organs of Generation. SECTION I. Of the External Parts of Generation - 200 SECTION II. Of the Vagina - 203 SECTION III. Of the Uterus, the Ovaries, and their Appendages - 205 SECTION IV. Of the Bladder and Urethra - 213 CONTENTS) Vii PART IS. OF THE BLOOD VESSELS. CHAPTER I. Of the General Structure and arrangement of the Blood Vessels, SECTION I. Of the Arteries ------- 226 SECTION II. Of the Veins - -- -- -- - 233 CHAPTER II. A Particular Aeeount of the Distribution of the Arteries. SECTION I. Of the Aorta, or the Great Trunk of the Arterial System 236 SECTION II. Of the Branches which go off from the Arch of the Aorta 23S SECTION III. Of the Branches which go off between the Arch and the Great Bifurcation of the Aorta - 2r2 SECTION IV. Of the Arteries which originate at and below the Great Bifurcation of the Aorta ----- 283 CHAPTER III. Of the particular Distribution of the Veins, SECTION I. Of the Superior, or Descending Vena Cava, and the Veins which communicate with it - 300 SECTION It Of the Inferior Vena Cava, and the veins connected with it - - - - - - 309 via CONTENl S. PART X. OF THE NERVES Nerves of the Brain - - - 3!23 Of the Cervical Nerves - _ - 343 Nerves of the Diaphragm 346 Brachial Plexus - - 347 Nerves of the Arm - - - 348 The Dorsal Nerves - . - 352 Of the Lumbar Nerves - - - 353 The Sacral Nerves - - - 356 Sciatic Plexus - - - 3 57 Great Sciatic Nerve . - 358 Great Sympathetic Nerve . - 361 Nerves of the Heart - - - 363 Nerves of the Abdominal Viscera - - - 368 PART XL OF THE ABSORBENT VESSELS. CHAPTER I. k Of the ./2b sorbents of the Lower Extremities — the Abdomen and the Thorax, SECTION I. i" e Absorbents of the Lower Extremities - - 376 SECTION II. The Absorbents of the Abdomen and Thorax - - 381 CHAPTER II. Of the Absorbents of the Head and Neel', of the Upper Ex- fremiti f \ and the Upper Part of the Trunk of the Body. SECTION I. Of the Absorbents of the Head and Neck - - 391 SECTION II. • >f the Absorbents of the Arm, and Upper Part of the Trunk - - - - - - SOS SYSTEM OF ANATOMY, PART VI. OF THE NOSE: THE MOUTH AND THE THROAT. CHAPTER I. OF THE NOSE. The prominent part of the 'face, to which the word nose is exclusively applied in ordinary lan- guage, is the anterior covering of two cavities which contain the organ of smelling. These cavities are formed principally by the upper maxillary and palate bones; and, therefore, to acquire a complete idea of them, it is necessary to study these bones, as well as the os ethmoides, the vomer, and the ossa spongiosa inferiora, which are likewise con- cerned in their formation. In addition to the description of these bones, in the account of the bones of the head, it will be use- ful to study the description of the cavities of the nose which follows it. See vol. I. page 64. After thus acquiring a knowledge of the bony struc- ture, the student will be prepared for a description of the softer part's Vn L , u . 1 3 The External Nose, SECTION i. Of the External Nose. The superior part of the nose is formed by the ossa nasi, and the nasal processes of the upper max- illary bones, which have been already described; (see vol. I. pages 42 — 46,) hut the inferior part, which is composed principally of cartilages, is much more complex in its structure. The orifice, formed by the upper maxillary and nasal bones, is divided by a cartilaginous plate, which is the anterior and inferior part of the sep- tum, or partition between the two cavities of the nose. The anterior edge of this plate projects be- yond the orifice in the bones, and continues in the direction of the suture between the ossa nasi. This edge forms an angle with the lower edge of the same cartilage, which continues from it, in a horizontal direction, until it reaches the lower part of the ori- fice of the nose, at the junction of the palatine pro- cesses of the upper maxillary bones; where a bony prominence is formed, to which it is firmly united. The upper part of the anterior edge of this carti- lage, which is in contact with the ossa nasi, is fiat, and is continued into two lateral portions that are extended from it, one on each side, and form a part of the nose : these lateral portions are sometimes spo- ken of as distinct cartilages: but they are really con- tinuations of the middle portion or septum. ^ Below the lower edge of these lateral portums are situated the cartilages which form the orifices of the nose, or the nostrils. Of these, there is one of con- siderable size, and several small fragments, on each side of the septum. Each of the larger cartilages forms a portion of an oval ring, which is placed ob- liquely on the side of the septum: so that the extre- 3 Of the Nose. jmty of the oval points downward and forward, while the middle part of the oval is directed upwards and backwards. The sides of this cartilage are flat, and unequal in breadth. The narrowest side is internal, and projects lower down than the cartilaginous sep- tum ; so that it is applied to its fellow of the other nostril. The external side is broader, and continues backward and upward to a considerable distance. The upper and posterior part of this oval ring is deficient; but the remainder of the nostril consists of several small pieces of cartilage, which are fixed in a ligamentous membrane that is connected by each of its extremities to the oval cartilage, and thus com- pletes the orifice. The anterior parts of the oval cartilages form the point of the nose ; and the ligamentous portions, the alae or lateral parts of the nostrils. When the external integuments and muscles are removed from the lower portion of the nose, so that the internal membrane and these cartilages only re- main, the internal membrane will be found attached to the whole bony margin of each orifice, and to each side of the whole anterior edge of the middle carti- lage, which projects beyond the bones. This mem- brane is afterwards continued so as to line the oval cartilages and the elastic membrane of the ala nasi, to the margin of the orifice of the nostril. The internal portions of the oval cartilages being situated without the septum, and applied to each other, they form the eternal edge of the partition between the nostrils, or the columna nasi ; which is very moveable upon the edge of the middle carti- lage. The orifices of the nostrils, thus constructed, are dilated by that portion of the muscle, called Levator Labii Superior is Alseque Nasi , which is inserted into the alas nasi. 4 Of the Cavities of the JYose. They are drawn down by the depressor labii supe- rioris alaeque nasi. They are pressed against the sep- tum and the nose by the muscle called Compressor JYaris , which has however an opposite effect when its upper extremity is drawn upwards by those fibres of the occipito-frontalis, which descend upon the nose, and are in contact with it. The end of the nose is also occasionally drawn down, by some muscular fibres which descend from it, on the septum of the nose, to the orbicularis oris : they are considered as a portion of this muscle by many anatomists, but were described by Albinus as a separate muscle, and called JYasalis Lahii Supe - rioris. When inspiration takes place with great force, the alse nasi would be pressed against the septum if they were not drawn out and dilated by some of the mus- cles above mentioned. SECTION II. Of the Cavities of the JYose. To the description of the osseous parts of the nasal cavities in vol. I. page 64, it ought now to be added that the vacuity in the anterior part of the osseous septum is filled up by a cartilaginous plate, connect- ed with the nasal lamella of the ethmoid bone above, and with the vomer below. This plate sends off those lateral portions already described, which form the cartilaginous part of the bridge of the nose. It should also be observed that at the back parts of these cavities are two orifices called the Posterior JYares, which are formed by the palate bones, the vomer, and the body of the sphenoidal bone, and are somewhat oval. The nasal cavities, thus constructed, are lined by a peculiar membrane, which is called pituitary front Schneiderian Membrane. 5 its secretion of mucus, or Schneiderian after an ana- tomist who described it with accuracy.* This membrane is very thick and strong, and abounds with so many blood vessels, that in the liv- ing subject it is of a red colour. It adheres to the bones and septum of the nose like the periosteum, but separates from them more easily. The surface which adheres to the bones has some resemblance to periosteum ; while the other surface is soft, spongy, and rather villous. Bichat seems to have considered this membrane as formed of two lamina, viz. perios- teum, and the proper mucous membrane ; but he adds, that it is almost impossible to separate them. It has been supposed that many distinct glandular bodies were to be seen in the structure of this mem- brane by examining the surface next to the bones ;f but this opinion is adopted by very few of the anato- mists of the present day. The texture of the mem- brane appears to be uniform ; and on its surface are a great number of follicles of various sizes, from which flows the mucous of the nose. These follicles appear like pits, made by pushing a pin obliquely into a surface which retains the form of the impression. They can be seen very distinctly with a common magnifying glass when the membrane is immersed in water, both on the septum and on the opposite surface. They are scattered over the mem- brane without order or regularity, except that in a few places they occur so as to form lines of various lengths, from half an inch to an inch. The largest of them are in the lower parts of the cavities. It may be presumed that the secretion of mucous is effected here by vessels which are mere continuations of arteries spread upon a surface analogous to the * Conrad .Schneider, a German professor, in a large work, “ De Ca- tarrhis,” published about 1660. t See Winslow, section X- No. 33 1 /, 6 Olfactory JVerves, exhalents, and not convoluted in circumscribed masses, as in the case of ordinary glands. The arteries of this membrane are derived from various sources : the most important of them is the nasal branch of the internal maxillary, which passes into the nose through the spheno-palatine foramen, and is therefore called the Spheno palatine Jlrtery. It divides into several twigs, which are spent upon the different parts of the surface of the nasal cavities. Two of them are generally found on the septum of the nose : one, which is small, passes forwards near the middle ; the other, which is much larger, is near the lower part of it. Two small arteries called the anterior and poste- rior ethmoidal, which are branches of the ophthal- mic, enter the nose by foramina of the cribriform plate of the ethmoidal bone. These arteries pass from the orbit to the cavity of the cranium, and then through the cribriform plate to the nose. In addi- tion to these, there are some small arteries derived from the infra orbital, the alveolar and the palatine, which extend to the Schneiderian membrane ; but they are not of much importance. The veins of the nose correspond with the arteries. Those which accompany the ethmoidal arteries open into the ocular vein of the orbit, which terminates in the cavernous sinuses of the head. The other veins ultimately terminate in the external jugulars. The nerves of the nose form an important part of the structure ; they are derived from several sources ; but the most important branches are those of the ol- factory. The olfactory nerves form oblong bulbs, which lie on each side of the crista galli, on the depressed por- tions of the cribriform plate of the ethmoid bone, within the dura mater. These bulbs are of a soft con- sistence, and resemble the cortical part of the brain Olfactory Nerves. 7 mixed with streaks of medullary matter. They send off numerous filaments, which pass through the fora- mina of the ethmoid bone, and receive a coat from the dura mater as they pass through it. These filaments are so arranged that they form two rows, one running near to the septum, and the other to the surface of the cellular part of the ethmoid bone, and the os turbinatum : and in addition to these are some intermediate filaments. When the Schneiderian membrane is peeled from the bones to which it is attached, these nervous fila- ments are seen passing from the foramina of the eth- moid bone to the attached surfaces : one row passing upon that which covered the septum, and the other to that of the opposite side ; while the intermediate filaments take an anterior direction, but unite to the membrane as soon as they come in contact with it. All of these can be traced downwards on the afore- said surfaces of the membrane for a considerable dis- tance, when they gradually sink into the substance of the membrane, and most probably terminate on the internal villous surface ; but they have not been traced to their ultimate termination. They ramify so that the branches form very acute angles with each other. On the septum the different branches are arranged so as to form brushes, which lie in contact with each other. On the opposite sides, the different ramifica- tions unite so as to form a plexus. Dr. Soemmering published last year some very elegant engravings of the nose, representing one of his dissections, which appears to have been uncom- monly minute and successful.* These represent the ramifications as becoming more expanded and deli- cate in the progress towards their terminations, and They are entitled, leones organovum liumanoruin olfactws. S Sphenopalatine and other Nerves of the Nose. as observing a tortuous course, with very short mean- dering flexures. It is to be observed that the i*am ideations of the olfactory nerve, thus arranged, do not extend to die bottom of the cavity. On the external side, they are not traced lower than the lower edge of the ethmoid, or of the superior spongy bone : and on the septum, they do not extend to the bottom, although they are lower than on the opposite side. On the parts of the membrane not occupied by the branches of the olfac- tory nerves, several other nerves can be traced. The nasal twig of the ophthalmic branch of the fifth pair, after passing from the orbit into the cavity of the cranium, proceeds to the nasal cavity on each side by a foramen of the cribriform plate ; and after send- ing off some fibrillse, descends upon the anterior part of the septum to the point of the nose. The sphe- no-palatine nerve, which is derived from the second branch of the fifth pair, and enters the nose by the spheno-palatine foramen, is spread upon the lower part of the septum and of the opposite side of the nose also, and transmits a branch through a canal in the foramen incisivum to the mouth. Several small branches also pass to the nose from the palatine and other nerves ; but those already mentioned are the most important. A question has been proposed, whether the olfac- tory nerve is exclusively concerned in the function of smelling, or whether the other nerves above men- tioned are also concerned in it. It seems probable that this function is exclusively performed by the olfactory nerve, and that the other nerves are like the ophthalmic branch of the fifth pair, with respect to the optic nerve. In proof of this, it is asserted that the sense of smelling has entirely ceased in some cases, where the sensibility to mechanical irritation of every kind has remained unchanged. If the ol- 9 Extent of the Schneiderian Membrane. factory nerve alone is concerned in the function of smelling, it follows, that this function must be confin- ed to the upper parts of the nasal cavities ; but it ought to be remembered, that the structure of the Schneiderian membrane, in the lower parts of these cavities, appears exactly like that which is above. The surface of the nasal cavities and their septum, when covered with the Schneiderian membrane, cor- responds with the osseous surface formerly descri- bed. The membrane covers the bones and cartilage of the septum, so as to make one uniform regular surface. From the upper part of the septum, it is continued to the under side of the cribriform plate of the ethmoid, and lines it ; the filaments of the olfactory nerve passing through the foramina of that bone into the fibrous surface of the membrane. It is continued from the septum, and from the cribri- form plate, to the internal surface of the external nose, and lines it. It is also continued backwards to the anterior surface of the body of the sphenoidal bone ; and, passing through the foramina or openings of the sphenoidal cells, it lines these cavities com- pletely ; but in these, as well as the other cavities, its structure appears somewhat changed ; it becomes thinner and less vascular. At the above mentioned foramina, in some subjects, it forms a plait or fold, which diminishes the aper- ture considerably. From the upper surface of the nasal cavities, the membrane is continued downwards over the surface opposite to the septum. On the upper flat surfaces of the cellular portions of the ethmoid, it forms a smooth uniform surface. After passing over the first turbinated bone, or that called after Morgagni, it is reflected into the groove, or upper meatus, immedi- ately within and under it ; the fold formed by the membrane, as it is reflected into the meatus, is rather Vol, ii. 2 10 Distribution of the Schneiderian Membrane . larger than the bone : and the edge of the fold there- fore extends lower done than the edge of the bone, and partly covers the meatus like a flap, consisting only of the doubled membrane. This fold generally continues backwards as far as the spheno-maxillary foramen, which it closes; the periosteum, exterior to the foramen, passing through it, and blending itself with the fibrous surface of the Schneide:ian mem- brane within. Here the spheno-palatine nerves and arteries join the membrane. Below this meatus, it extends over the middle, (formerly called the upper,) turbinated bone, and is reflected or folded inwards on the under side of this bone, and continued into the middle meatue below it. In the middle meatus, which is partly covered by the last-mentioned turbi- nated bone, there are two foramina: one communi- cating with the maxillary sinus ; and the other with the anterior cells of the ethmoid and the frontal si- nuses. The aperture into the maxillary sinuses is much less in the recent head, in which th Schnei- derian membrane lines the nose, than it is in the bare bones. A portion of the aperture in the bones is closed by the Schneiderian membrane, which is extended over it : the remainder of the aperture is unclosed ; and through this foramen, the membrane is reflected so as to line the whole cavity. As a por- tion of the foramen is covered by the membrane, and this portion as well as the other parts of the cavity is lined by the membrane, it is obvious that at the place where the membrane is extended over the foramen in the bone, it must be doubled ; or, in other words, a part of the aperture of the maxillary sinus is closed by a fold of the Schneiderian mem- brane. This aperture varies in size in different subjects, and is often equal in diameter to a common quill. It is situated in the middle of the meatus, and is covered Eustachian Tube. 11 by the middle turbinated bone : immediately above it, is a prominence of the cellular structure of the eth- moid bone, which has a curved or semicircular figure. Near this prominence, in the same meatus, a groove terminates, which leads from the anterior ethmoid cells and the frontal sinuses. From the middle meatus, the membrane proceeds over the inferior turbinated bone, and is reflected round and under it into the lower meatus. It appears rather larger than the bone which it covers; and therefore the lower edge of the bone does not extend so low as the lower edge of the membrane, which of course is like a fold or plait. The membrane then continues and lines the lower meaiuos here it appears less full than it is in the turbinated bone. In this meatus, near to its anterior end, is the lower orifice of the lachrymal duct : this is simply lined by the Schneiderian membrane, which is continued into it, and forms no plaits or folds that affect the orifice. Orifice of the Eustachian Tube. Immediately behind each of the nasal cavities, on the external side, is the orifice of the Eustachian Tube. It has an oval form, and is large enough to admit a very large quill. Its position is oblique; the upper extremity being anterior to the other parts of the aperture, and on a line with the middle mea- tus, while the centre is behind the inferior turbinated bone. The lower part of the oval is deficient. This tube is formed posteriorly by a cartilaginous plate. It is lined by the membrane continued from the nose. The cavities of the nose answer a twofold purpose in the animal economy: they afford a surface for the expansion of the olfactory nerves : and a passage for the external air to the windpipe, in respiration. The function of smelling appears to be dependent, to a certain degree, upon respiration. It has been as- 12 Observations respecting the Nose. serted that unless the air passes in a stream through the nose, as in respiration, the perception of odour does not take place; that in persons who breathe through wounds and apertures in the windpipe, the function of smelling is not performed. It is rather in confirmation of this proposition, that most per- sons, when they wish to have an accurate perception of any odour, draw in air rapidly through the nose. Although the ultimate terminations of the olfactory nerves cannot be demonstiated like those of the op- tic and auditory nerves, it is probable, from the ap- pearance of the fibres, while they are distinguisha- ble, that they are finally arranged with great delica- cy. It is certain that the impressions from whence we derive the perceptions of many odours must be very slight; as odorous bodies will impregnate the aii of a large chamber for a great length of time, without losing any sensible weight. With respect to delicacy of structure and sensibility, it is probable that the nose holds a middle rank be- tween the eye or ear, and the tongue : and on this account the mucous is necessary as a covering and defence of its surface. It has been ascertained, by the investigations of che- mists, that this mucous contains the same ingredi- ents as the tears already described, viz. animal mu- cous and water; with muriate of soda, and soda un- combined; phosphate of lime, and phosphate of soda. The animal mucous, which is a most important ingre- dient in the composition, resembles the mucilage formed by some of the vegetable gums in several particulars ; and differs from them in others. The mucous of the nose, if it remain there long after it is secreted, becomes much more viscid in con- sistence, and changes from a whitish colour to one which partakes more or less of the yellow. It is probable that an incipient putrefaction may occasion these changes in it. The use of the frontal, maxillary and other sinuses, communicating with the nose, has been the subject of some inquiry. As there can be no stream of air through them, and as the membrane lining them is neither so thick, villous nor flexible as that lining the nose, it may be concluded, a priori, that they are not concerned in the function of smelling. This 13 Use of the Sinuses of the Nose, opinion is strengthened by the fact, that very young children, in whom these sinuses scarcely exist, en- joy the sense of smelling in perfection. The fol- lowing fact is also in support of it. The celebrated Desault attended a patient, in whom one of the frontal sinuses was laid open by the destruction of the bone which covered it anteriorly. This patient was able to breathe a short time through the sinus when the mouth and nose were closed : At the re- quest of Desault he breathed in this manner when a cup of some aromatic liquor was held near the opening of the sinus : and had not the least percep- tion of odour. This experiment was repeated se- veral times. Many physiologists believe that these sinuses have an effect in modulating the voice. CHAPTER II. OF THE MOUTH. The general cavity of the mouth is formed ante- riorly and laterally by the connexion of the lips and cheeks to the upper and lower jaws : so that the teeth and the alveoli of both jaws may be considered as within the cavity. Above, it is bounded principally by the palatine processes of the upper maxillary and palate bones, and the soft palate, which continues backward from them in the same direction. Below, the cavity is completed by several muscles, which proceed from almost the whole internal circum- ference of the lower jaw, and, by their connexions with each other, with the tongue and the os hyoides, form a floor or bottom to it. The tongue is particular- ly connected to this surface, and may be considered as resting upon and supported by it. To acquire an idea of the parietes of this cavity, after studying the upper and lower maxillary bones, the orbicularis oris and the muscles connected with it, especially the buccinator, ought to be examined ; and also the digastricus, the mvlo-hyoideus, genio-hyoi- deus, and genio-hyoglossus. By this it will appear that the lips and cheeks, and the basis or floor of the mouth, are formed in a great measure by muscles. Upon the internal surface of these muscles, a por- tion of cellular and adipose substance is arranged, as well as glandular bodies of different sizes ; and to these is attached the membrane which lines the in- side of the mouth. This membrane passes from the skin of the face to the lips, and the inside of the mouth; and, al- though it is really a continuation of the skin, there Internal Surface of the Mouth. 15 is so great a change of structure, that it ought to be considered as a different membrane. At the orifice of the lips it is extremely thin, and so vascular that it produces the fine florid colour which appears there in health. It is covered by a cuticle, called by some anatomists, Epithelium , which has a proportionate degree of delicacy, and can be separated like the cuticle in other parts. When this cuticle is separa- ted, the lips and the membrane of the mouth appear to be covered with very fine villi, which are parti- cularly apparent in some preparations of the lips af- ter injection and maceration.* Under this membrane are many small glandular bodies' of a roundish form, called glandulse labiales, whose excretory ducts pass through it to the inner surface of the mouth, for the purpose of lubrifying it withjjieir secretion, which is mingled with the saliva. The membrane, which lines the inside of the lips and cheeks, is somewhat different from that which forms the surface of the orifice of the mouth : it is not so florid ; the blood vessels in its texture are larger and not so numerous. This change, however, takes place very gradually, in the progress of the membrane, from the orifice of the lips to the back part of the cheeks. Glandular bodies, like those of the lips, are situated immediately exterior to this membrane of the cheeks, between it and the mus- cles : their ducts open on its surface. These glands are called Buccales. This lining membrane is continued from the inter- nal surface of the lips and cheeks to the alveolar por- tions of the upper and lower jaws, which are in the cavity of the mouth, and covers them, adhering firmly to the periosteum. * Ruysch had a fine preparation of this structure. See Thesaurus VII. Tab. III. Fig. 5. 16 Gums. — Surface of the Hard Palate. The teeth appear to have passed through aper- tures in this membrane, and are surrounded by it closely at their respective necks. The portion of membrane, which thus invests the jaws, constitutes the gums; which have now acquired a texture very different from that of the membrane, from which they were continued. They are ex- tremely firm and dense, and very vascular. It is probable that their ultimate structure is not perfect- ly understood. In the desease called scurvy , they tumify and lose the firmness of their texture ; they acquire a livid colour, and are much disposed to hemorrhage. From the alveoli of the upper jaw, the lining membrane is continued upon the palatine processes of the upper maxillary and palate bones, or the roof of the mouth. This membrane of the palate is not quite so firm as that of the gums, and is also less florid : it adheres firmly to the periosteum, and thus is closely fixed to the bones. There is generally a ridge on its surface, immediately under thesature between the two upper maxillary bones ; and some transverse ridges are also to be seen upon it. On the internal surface of this membrane are small glandular bodies, whose ducts open on the surface of the palate. It is asserted, that this membrane has a limited de- gree of that sensibility which is essential to the func- tion of tasting ; and, that if certain sapid substances are carefully applied to it, their respective tastes will be perceived, although they have not been in contact with the tongue. The membrane is continued from the bones above mentioned to the soft palate, or velum pendulum pa- lati, which is situated immediately behind them. This soft palate may be considered as a continuation of the partition between the nose and mouth ; it is at- 17 Soft Palate.-- Uvula. tached to the posterior edge of the palatine processes of the ossa palati, and to the pterygoid processes of the sphenoidal bone. Its interior structure is mus- cular. The upper surface is covered by the mem- brane of the nose, the lower surface by the mem- brane which lines the month. The muscles, which contribute to the composition of this structure, are the circumflexi and the le- vatores palati above, and the constrictores isthmi faucium and palato-pharyngei below. (See Vol. I. page 190 — 191.) Thus composed the soft palate constitutes the back part of the partition between the nose and mouth. When viewed from before, with the mouth open, it presents towards the tongue an arched surface, which continues downwards on each side, until it comes nearly in contact with the edges of that organ. On each of the lateral parts of this arch, are two pillars, or rather prominent ridges, which project into the mouth. These ridges are at some distance from each other below, and approach much nearer above, so that they include a triangular space. They are called the lateral half arches of the palate. Each of them is formed by a plait or fold of the lining membrane of the mouth, and contains one of the two last mentioned muscles : the anterior, the constrictor isthmi faucium ; the posterior, the pa- lato-pharyngeus.^Tk a vities or. both sides of it. > This assu^ion of «. fact so difficult to reconcile with the general prin- ciples of the drculation. was received with great hesitation : and although The Great Vessels . 59 From the coarse of these different vessels round the basis of the ventricles of the heart, they are ge- nerally called Coronary Vessels: the arteries are denominated, from their position, Right and Left Coronary. The nerves of the heart come from the cardiac plexus, which is composed of threads derived from the intercostal or great sympathetic nerves, and the nerves of the eighth pair. SECTION III. Of the Aorta , the Pulmonary Artery and Veins , and the Vense Cavse ; at their commencement . The two great arteries, which arise from the heart, commence abruptly, and appear to be extremely dif- ferent in their composition and structure from the heart. They are composed of a substance, which has a whitish colour and very dense texture, and is very elastic as well as firm and strong. When the pericardium is removed, these arteries appear to proceed together from the upper part of the basis of the heart : the pulmonary artery being placed it was confirmed by some very respectable anatomists of the last century, it was denied by others. Some of the anatomists of the present day have denied the existence of these orifices, and some others have neglected them entirely. The subject has lately been brought forward in the London Philoso- phical Transactions for 1798, Part I by a very respectable anatomist, Mr. Abernethy, who states that he has often passed a coarse waxen injection from the proper arteries and veins of the heart into all the cavities of that organ, and particularly into the Left Ventricle. But it was only in sub- jects with diseased lungs that this was practicable. The existence of this communication between the coronary vessels and the great cavities of the heart seems therefore to be proved. The easy demonstration in such subjects is ingeniousl) referred by Mr. Abernethv. to the obstruction of the circulation in the lungs : and he regards the com- munication as a provision enabling the coronary vessels to unload them- selves, when the coronary vein cannot dtscha-ge freely into the right auricle. 60 Pulmonary Artery and Veins and Vense Cause. to the left of the aorta with the left auricle on the left side of it, and the right auricle on the right side of the aorta. The pulmonary artery arises from the most anterior, and left part of the basis of the right ventricle, and proceeds obliquely backwards and up- wards ; inclining gradually to the left side for about eighteen or twenty lines ; when it divides into two branches which pass to the two lungs. The aorta arises from the left ventricle, under the origin of the pulmonary artery, and immediately pro- ceeds to the right, covered by that vessel, until it mounts up between it and the right auricle : it then forms a great curve, or arch, which turns backward and to the left, to a considerable distance beyond the pulmonary artery. In this course, it crosses the right branch of the pulmonary artery ; and, turning down in the angle between it and the left branch, takes a position on the left side of the spine. The course of this artery, from its commencement at the ventricle, to the end of the great curve or arch, is extremely varied. The uppermost part of the curve is in the bottom of the chamber formed by the separation of the la- mina of the mediastinum when they join the first rib on each side. From this part of the curve three large branches go off, viz. one, which soon divides into the carotid and the subclavian arteries of the right side ; a se- cond, somewhat smaller, which is the left carotid ; and a third, which is the left subclavian artery. When the heart and its great vessels are viewed from behind, (after they have all been filled with injection ; and the pericardium, mediastinum, and windpipe have been removed,) the aorta appears first, descending behind the other vessels ; the pul- monary artery then appears, dividing so as to form an obtuse angle with its two great branches, each of Pulmonary Artery and Veins and Verne Cavse. 61 which divides again before it enters the lung to which it is destined. Under the main trunk of the pulmonary artery is the left auricle : its posterior surface is nearly of a square form, and each of the pulmonary veins pro- ceeds from one of its angles. These veins ramify in the substance of the lungs, at a very short distance from the auricle : the two uppermost of them are si- tuated rather anterior to the branches of the pulmo- nary artery. In this posterior view, the pulmonary vessels of the right side cover a great part of the right auricle, as it is anterior to them. The lower portion of the auricle, with the termination of the inferior cava, is to be seen below them. Above them the superior cava appears ; and in that part of it which is imme- diately above the right branch of the pulmonary ar- tery, is the orifice of the vena azygos. In its natural situation in the thorax, the superior cava is connected by cellular membrane to the right lamen of the mediastinum, and supported by it. At a small distance below the upper edge of the ster- num, it receives the trunk formed by the left subcla- vian and internal jugular vein, which passes oblique- ly across the sternum below its inner edge, in the upper space between the lamina of the mediastinum. CHAPTER III. OF THE TRACHEA AND THE LUNGS. Although the principal part of the windpipe is situated in the neck above the cavity of the thorax., it is so intimately connected with the lungs, that it is necessary to describe them together, SECTION I. Of the Trachea. Trachea is the technical name for. the windpipe, or tube which passes from the larynx to the lungs. This tube begins at the lower edge of the cricoid cartilage, and passes down the neck in front of the oesophagus as low as the third dorsal vertebra, when it divides into two branches called Bronchia , one of which goes to the right and the other to the left lung and ramifies very minutely in them. There is in its structure a number of flat cartila- ginous rings placed at small distances from each oth- er, the edges of which are connected by membrane so that they compose a tube. These cartilaginous rings are not complete, for they do not form more than three-fourths or four-fifths of a circle ; but their ends are connected by a mem- brane which forms the posterior part of the tube. They are not alike in their size or form ; some of them are rendered broader than others, by the union of two or three rings with each other, as the upper- most. The lowermost also is broad, and has a form which is accommodated to the bifurcation of the tube. Their number varies, in different persons from fif- teen to twenty. 63 Structure of the Trachea . These rings may be considered as forming a part of the first proper coat of the trachea ; which is com- posed of them, and of an elastic membrane that occu- pies all the interstice between them ; so that the car- tilages may be regarded as fixed in this membrane. A similar arrangement of rings exists in the great branches of the bronchia ; but after they ramify in the lungs, the cartilages are no longer in the form of rings : they are irregular in their figures, and are so arranged in the membrane that they keep the tube completely open. These portions of cartilage do not continue throughout the whole extent of the ramifi- cations ; for they become smaller, and finally disap- pear, while the membranous tube continues without them, ramifying minutely, and probably forming the air cells of the lungs. This membrane is very elastic : the lungs are very elastic also ; and it is probable that their elasticity is derived from this membrane. On the inside of this coat of the trachea is an ar- rangement of muscular fibres, which may be called a muscular coat. It is best seen by peeling off or removing the internal coat to be next described. On the membranous part of the trachea, where the cartilaginous rings are deficient, these muscular fibres run evidently in a transverse direction : in the spaces between the cartilages their direction is lon- gitudinal. There is some reason to doubt whether these longitudinal fibres are confined altogether to the spaces between the cartilaginous rings, and at- tached only to their edges, because their is a fleshy substance on the internal surface of the rings, which appears to be continued from the spaces between them. The internal coat of the trachea is a thin and de- licate membrane, perforated with an immense num- 64 Black Glands of the Bronchise. ber of small foramina; which are the orifices of mu- cous ducts. On the surface of this membrane there is an ap- pearance of longitudinal fibres which are not distri- buted uniformly over it, but run in fasciculi in some places, and appear to be deficient in others. These fasciculi are particularly conspicuous in the ramifica- tions of the bronchia in the lungs. On the posterior membranous portion of the tra- chea, where the cartilages are deficient, a consider- able number of small glandular bodies are placed, which are supposed to communicate with the mucous ducts that open on the internal surface. If these bodies are removed from the external surface of this portion, and the muscular fibres are also removed from the internal, a very thin membrane only re- mains, which is very different from that which is left between the rings, when the fleshy substance is re- moved from that situation. The reason of the deficiency in the rings, at this posterior part, is not very obvious.* It continues in the bronchia until the form of their cartilages is changed in the lungs : if it were only to accommo- date the oesophagus, during the passage of food, there would be no occasion for its extension to the bronchia. At the bifurcation of the trachea, and on the bronchia, are a number of black coloured bodies, which resemble the lymphatic glands in form and texture. They continue on the ramifications of the bronchia some distance into the substance of the lungs. Their number is often very considerable ; and they vary in size from three or four lines in dia- * Doctor Physic has advanced the opinion that it enables a person to ex- pel the raucous of the lungs by contracting the size of the trachea, and con- sequently increasing the velocity or impetus of the air.— Ed. 65 Root of the Lungs. meter to eighteen or twenty. As lymphatic vessels have been traced to and from them during their course to the thoracic duct, they are considered as lympha- tic glands. SECTION II. Of the Lungs. There are two of these organs: each of which oc- cupies one of the great cavities of the thorax. When placed together, in their natural position, they resemble the hoof of the ox, with its back part forward ; but they are at such a distance from each other, and of such a figure, that they allow the medi- astinum and heart to intervene; and they cover every part of the heart anteriorly, except a small portion at the apex. Each lung fills completely the cavity in which it is placed, and every part of its external surface is in contact with some part of the internal surface of the cavity ; but when in a natural and healthy state, it is not connected with any part except the lamina of the mediastinum. One great branch of the trachea and of the pulmonary artery passes from the mediastinum to each lung, and enters it at a place which is rather nearer to the upper rib than to the diaphragm, and much nearer to the spine than the sternum : at this place also the pulmonary veins return from the lungs to the heart. These vessels are enclosed in a membrane, which is continued over them from the mediastinum, and extended from them to the lung. Thus covered, they constitute what has been called the Root of the Lung. When their covering, derived from the mediasti- num, is removed, the situation of these vessels ap- pears to be such that the bronchia are posterior, Vol. ii. 9 66 Colour of the Lungs. the branches of the pulmonary artery are rather above and before, and the veins below and before them. Each of these vessels ramifies before it enters into the substance of the lungs: the bronchia and the branches of the pulmonary artery send each a large bianch downward to the inferior part of the lungs, from which the lower pulmonary veins pass in a di- rection nearly horizontal. In general, each of the smaller ramifications of the bronchia in the lungs is attended by an artery and a vein. Each lung is divided, by very deep fissures, into portions which are called Lobes. The right lung is composed of three of these lobes, and the left lung of two. The lungs are covered as has been already stated, with the reflected portion of the pleura continued from the mediastinum, which is very delicate and al- most transparent. They have, therefore, a very smooth surface, which is kept moist by exudation from the arteries of the membrane. The Colour of the Lungs is different in different subjects. In children they are of a light red colour; in adults they are often of a light grey ; owing to the deposition of a black pigment in the substance immediately under the membranes which form their external surface. Their colour is often formed by a mixture of red and black. In this case they are more loaded with blood, and the vessels of the internal membranes being distended with it, the red colour is derived from them. The black pigment sometimes appears in round spots of three or four lines in diameter : under the ex- ternal membrane it is often in much smaller portions, and sometimes is arranged in lines in the interstices of the lobuli, to be hereafter mentioned. It is also diffused in small quantities throughout the substance of the lungs. Structure of the Lungs . 67 The source of this substance, and the use of it, are unknown. The lungs are of a soft spongy texture ; and, in animals that have breathed, they have always a con- siderable quantity ol air in them. They consist of cells, which communicate with the branches of the trachea that ramify through them in every part. These cells are extremely small, and the membranes which compose them are so thin and delicate that if they are all filled by an injection of wax, thrown into the trachea, the whole cellular part of the lung will appear like a mass of wax. If a corroded preparation be made of a lung injected in this manner with force, the wax will appear like a concretion. These effects of injections prove that the membranes of which the cells are formed are very thin ; and, of course, that their volume is very small when compar- ed with the capacity of the cells. In those corroded preparations, in which the ra- mifications of the broncnia are detached from the wax of the cells, these ramifications become extremely small indeed. If the lungs of the human subjects, or of animals of similar construction, be examined when they are inflated, their cellular structure will be very obvious, although, their cells are so small that they cannot commonly be distinguished by the naked eye. Each of the extreme ramifications of the bronchia appears to be surrounded by a portion of this cellular sub- stance, which is gradually distended when air is blown into the ramification. This celiular substance is formed into small por- tions of various angular figures, which are denomi- nated Lobuli : these can be separated to a consider- able extent from each other. They are covered by the proper coat of the lungs, which is extremely de= 68 Structure of the Lungs. licate, and closely connected to the general covering derived from the pleura. Between the lobuli, where they are in contact with each other, there is a por- tion of common cellular substance, which is easily distinguished through the membrane covering the lungs. This is very distinct from the cellular struc- ture which communicates with the ramifications of the bronchia, and contains air ; for it has no com- munication with the air, unless the proper coat of the lungs be ruptured. If a pipe be introduced by * a puncture of the external coat of the lungs, and this interstitial cellular membrane be inflated, it will compress the lobuli. This cellular membrane is al- ways free from adipose matter : it may be easily ex- amined in the lungs of the bullock. Upon the membranes which compose the air cells, the pulmonary artery and vein ramify most minutely; and it seems to have been proved within the last thirty years, by the united labours of chemists and physio- logists, that the great object of respiration is to effect a chemical process between the atmospheric air, when taken into the air cells, and the blood which circu- lates in these vessels. In addition to the blood vessels which thus pass through the substance of the lungs, there are several smaller arteries denominated Bronchial , which arise either from the upper intercostal, or from the aorta itself: they pass upon the bronchia, and are distri- buted to the substance of the lungs. The veins which correspond with these arteries terminate ulti- mately in the vena azygos. The nerves of the lungs are small in proportion to the bulk of these organs. They are derived princi- pally from the par vagum and the intercostal nerves. The elasticity of the air cells of the lungs and of the ramifications of the bronchia which lead to them, appears by their rapid contraction after distention, 69 The Thorax of the Foetus. and by the force with which they expel the air which is used to inflate them when taken out of the thorax. The Thorax of the Foetus. In the cavity between the lamina of the medias- tinum, where they approach each other from the first ribs, is situated a substance which is denominated the Thymus Gland. This substance gradually diminishes after birth, so that in the adult it is often not to be found : and when it exists it is changed in its texture, being much firmer, as well as greatly diminished. In the foetus it is of a pale red colour ; and during infancy it has a yellowish tinge. It generally ex- tends from the thyroid gland, or a little below it, to the pericardium. From its superior portion two la- teral processes are extended upwards : below, it is formed into two lobes, which lie on the pericardium. If an incision be made into its substance, a fluid can be pressed out, which has a whitish colour, and coagulates upon the addition of alcohol. Although it is called a gland , no excretory duct has ever been found connected with it. The blood vessels of this body are derived from the thyroid branches of the subclavians, from the in- ternal mammaries, and the vessels of the pericardium and mediastinum. The Heart , And the great arteries which proceed from it, have some very interesting peculiarities in the foetus. In the septum between the two auricles, is a fora- men of sufficient size to permit the passage of a large quill, which inclines to the oval form, with its long- est diameter vertical when the body is erect. On the left side of the septum, a valve, formed by the lining membranes, is connected to this foramen ; and 70 The Thorax of the Foetus. allows a free passage to a fluid moving from the right auricle to the left, but prevents the passage of a fluid from the left to the right. This structure is evidently calculated to allow some of the blood which flows into the right auricle from the two venae cavse to pass into the left auricle of the heart instead of going into the right ventricle. As the contents of the left auricle pass into the left ventricle, and from thence into the aorta, it is obvious that the blood, which passes from the right auricle into the left through this foramen, must be transmitted from the sys- tem of the vena cava to the system of the aorta without going through the lungs, as it must necessa- rily do in subjects who do not enjoy the foetal struc- ture. The Pulmonary Artery and the Aorta Have a communication in the foetus, which is very analogous to the communication between the auricles of the heart. From the pulmonary artery, where it divides into the two great branches, another large branch conti- nues, in the direction of the main trunk, until it joins the aorta ; with which it communicates at a small distance below the origin of the left subclavian artery. In the young subject that has never respired, it ap- pears as if the pulmonary artery was continued into the aorta, and sent off' in its course, a branch on each side, much smaller than itself, to each lung. In subjects that have lived a few days, these branches to the lungs are much larger ; and then the main pulmonary artery appears to have divided into three branches : one to each lung, and one to the aorta ; but that which continues to the aorta is larger than either of the others. In the course of time, however, this branch of the aorta is contracted, so that no fluid passes through it; General Observations. 71 and it has the appearance of a ligament, in which slate it remains. The course of the blood from the right ventricle, through the pulmonary artery, to the aorta below its curve, is more direct than that from the left ventricle to the same spot, through the aorta at its commence- ment. The column of blood in the aorta below its curve is evidently propelled by the force of both ven- tricles : and this circumstance, although it seems to proceed merely from the state of the foetal lungs, is particularly calculated for the very extensive circula- tion which the foetus carries on, by means of the um- bilical arteries and vein in the placenta. The Lungs of the. Foetus Differ greatly from those of the adult. They appear solid, as if they were composed of the parenchymatous substance which constitutes the matter of glands, rather than the light spongy substance of the lungs of adults. They differ also in colour from the lungs of older subjects, being of a dull red. They have greater specific gravity than water ; but if air be once inspired, so much of it remains in them that they ever afterwards float in that fluid. The nature of the process of respiration, and its ef- fects upon the animal economy, particularly upon the action of the heart, appear to be much better un- derstood at this time than they were before the dis- covery of the composition of the atmosphere, by Dr. Priestly and by Mr. Scheele. The publications upon this subject, which have appeared since that period, viz. 1774, are therefore much more interesting to the student of medicine than those which preceded them. Two of these publications ought to be parti- cularly noticed by him : viz, an essay, by Dr. Ed- ward Goodwyn, entitled, “ The Connexion of life 72 Cases of Malformation. with respiration;” and — the “ Physiological Re- searches of M. Bichat upon Life and Death. Part Second.* The general doctrines respecting the oxygenation or decarbonation of the blood, and the absolute necessi- ty that it should take place to a certain degree in or- der to preserve life, are confirmed by a number of cases of malformation of the heart or the great ves- sels, in which the structure was such that a consider- able portion of venous blood passed from the right side of the heart to the aorta, without going through the lungs. In these different cases, notwithstanding the structure was somewhat varied, the symptoms produced were very much alike ; differing in the re- spective patients in degree only, and not in kind. The symptoms indicating this structure are blue colour of the face, (such as generally accompanies suffocation.) extending more or less over the whole body, and particularly apparent under the nails of the fingers and toes ; anxiety about the region of the heart; palpitation; laborious respiration; sensations of great debility, &c.: all of which are greatly aggra- vated by muscular exertion. These effects have gen- erally appeared to be proportioned to the quantity of venous blood admitted into the aortic system. t When these appearances take place immediately after birth, it is probable that they depend entirely upon malformation of the heart or great vessels; but when they commence at a subsequent period, they are * The student will derive much information respecting the publications on this subject, prior to 1804, from Dr. Bostock’s Essay on Respiration — Since the publication of that essay several interesting papers on respira- tion have appeared, viz. Two memoirs by the late Abbe Spalanzani ; — “ An Enquiry into the Changes induced on Atmospheric Air by the Ger- mination of Seeds,” &c. by Ellis ; — two very important communications by Messrs. Allen and Pepys in the Transactions of the Royal Society of Lon- don for 1808 and 1809; — and “ Further Enquiries into the Changes induced on Atmospheric Air.” Also by Ellis. f Cases of this kind are related in several of the periodical publications on medical subjects. Two of them were described by the late Dr. Wil- liam Hunter in the sixth volume of Medical Observations and Enquiries, by a Society of Physicians in London ; one, (quoted by Dr. Goodwyn,) is in the Observationes Anatomicse of Sandifort ; and another, by Dr J. S. Dorsey, has lately been published in the first oumber of the New England Journal of Medicine and Surgery. Cases of Malformation. 73 commonly the effect of a diseased alteration in the lungs. They sometimes occur near the termination of fcital cases of pneumonia or catarrh ; but a . diffe- rent cause, which has not latterly been suspected, appears to have produced them in the following case related by Dr. Marcet, in the first volume of the Edinburgh Medical and Physical Journal. The blue colour occurred in a young woman, twen- ty-one years of age, in whom it had never been ob- served before. — It came on during an affection of the breast, and was attended with great prostration of strength and difficulty of breathing ; as well as cough, oedema of the hands and feet, and several other symptoms. About seven weeks after the com- mencement of these symptoms she died $ when it was ascertained by dissection, that there was no unnatural communication whatever between the ca- vities of the heart, and that its valves were all in a perfect and natural state . The lungs were free from tubercles, or any other appearance of disease. Their substance seemed more compact than usual, especially the left lung, although it did not sink in water 5 but they adhered every where to the inner surface of the thorax, to the diaphragm , and to the pleura covering the pericardium . — This case is the more remarkable, because numberless instances have occurred in which very large, portions of the external surface of the lungs have been found upon dissection to adhere to the internal surface of the thorax without the occurrence of such symptoms during life. It may be inferred, from a statement published by M. Dupuytren, in a late volume of the Proceedings of the National Institute of France, that the oxygena- tion or decarbonation of the blood is much affected, in respiration, by an influence exercised by the nerves which are appropriated to the lungs. From his account it appears, that although the complete division of the eighth pair of nerves produces death after some time ; yet in the horse whose nerves are thus divided, life continues, and respiration goes an, from half an hour to ten hours ; but his arterial blood is in a state of great disoxygenation. or corbo- nation, during this time. This fact is more rem sxka- Vol, II„ W 10 74 Effects of Venous Blood on the Heart. hie, because venous blood, contained in a bladder exposed to the open air, will become oxygenated or decarbonated. It is also asserted in another Memoir, read to the National Institute by Dr. J. M. Provem^al, that ani- mals in whom the eighth pair of nerves has been divided, do not consume so much oxygen, or produce so much carbonic acid, by a considerable degree, as they did before the division of these nerves ; and that their temperature is considerably reduced.* The fact, that venous blood occasions death, when it is admitted into the left ventricle of the heart, and the aorta, is truly important. Dr. Goodwin explained it by suggesting that this blood was not sufficiently sti- mulating to produce the necessary excitement of the heart ; but on this occasion one of his friends propo- sed to him the following question: Why does venous blood affect the left side of the heart in this injurious manner, when it appears to exert no noxious effects whatever on the right side of tbatorgan ? His reply may be seen in a note at the 8 2d page of his Essay, in the first edition. Bichat has offered a solution which completely resolves this difficulty, viz. “ The effect of Venous blood upon the heart is produced by the presence of this blood in the proper, or coronary arteries of that organ, and not in its great cavities.” For the animation of the heart, like that of the other parts of the body, depends upon the state of the blood in the arteries which penetrate its fexture.t And while the heart acts, the blood of the coronary arte- ries will be the same with that of the left ventricle. See Bichat’s Researches, P. II. art. 6. § 2. The French Anatomists appear to entertain some pecu- liar opinions respecting the course of the blood in the foetus, which have a particular relation to the sub- ject last mentioned. Winslow, who paid great at- tention to the valve of Eustachius in the right auricle of the heart, was of opinion, that this valve was cal- These Memoirs are republished in the Eclectic Repertory of Phila- delphia for April and October, 1811. f It is probable that the contents of the great cavities of (.lie heart have no more effect upon its animation than the contents of the stomach and. bowels love upon the animation of those organs,. Sentiments of Sabatier , fyc. 75 dilated for some important purpose in the fetal eco- nomy.* 1 Although his hypothesis respecting its par- ticular use has not been retained by his countrymen, many of them have adopted his general sentiment $ and among others Sabatier. That learned anatomist believed that this valve, in the fetal state, serves to direct the blood of the inferior cava , after its ar- rival in the right auricle , through the foramen ovale into the left auricle ; while the blood of the upper cava passes directly into the right ventricle. His opinion seems to be supported to a certain degree. 1. By the direction in which the two columns of blood enter the auricle from the two vense cavse. 2. By the position of the Eustachian valve. 3. By the foramen ovale, when its valve is com- . plete ; as the passage through it, from the right to the left, is at that time oblique, and from below upwards. The theory of Sabatier appears to be this : the um- bilical vein brings from the placenta blood which has a quality essential to the animation of the fetus. If there were no particular provision to the contrary, a large portion of this blood, after passing from the umbilical vein by the inferior cava into the right • auricle of the heart, would proceed by the right ven- tricle, though the pulmonary pretry and arterial ca- nal, into the aorta, below the origins of the carotid and subclavian arteries ; and consequently none of it would pass to the head and upper extremities, but a considerable part would return again by the umbi- lical arteries to the placenta, without circulating through the body : while on the other hand, the blood which passed by the carotid and subclavian arteries to the head and upper extremities, return- ing from them to the heart by the superior cava, might pass from the right auricle to the left auricle and ventricle and the aorta, and so to the head and upper extremities again, without passing through the placenta. But by means of tins valve, the blood of the Sower cava, and of course of the umbilical vein, is directed to the left auricle and ventricle and the aorta, by which a considerable portion of it will necessarily pass to the head and upper ex- tremities : while the blood which returns from these See Memoirs of the Academy of Sciences for 1717 and 1725. 76 Sentiments of Sabatier , parts by the superior cava, must consequently pass from the right auricle into the right ventricle ami pulmonary artery ; from whence a large portion of it will proceed through the arterial canal into the aorta beyond the carotids and subclavians, and of this por- tion a considerable part will go to the placenta by the umbilical atteries. Sabatier compares the course of the blood in the foetus to the course of afluidin a tube which has the form of the numeral character 8.* — If this doctrine be true, the progress of the blood in the foetus and placenta is very analogous to that of the double circulation of the adult; the character 8 an- swering equally well in the description of either sub- ject. According to Sabatier, the blood of the placenta takes this peculiar course through the heart, in order that some of it may be carried to the head and upper extremities. But an additional reason may be sug- gested, which appears to be of great importance ; viz. the supplying of the coroijary or proper xesse\s of the heart with some of the same blood. The heart of the adult, as has been before stated, cannot act without its proper or coronary arteries are supplied with arterial blood. The heart of the foetus performs a more extensive circulation than that of the adult, and therefore is probably in greater need of such blood. But unless the blood of the placenta pas- ses through the foramen ovale into the left auricle and ventricle , and so to the aorta , it cannot enter the coro- nary arteries which originate at the commencement of the aorta; for the b.lood which flows from the right side of the heart through the arterial canal, passes into the aorta at so great a distance from the orifices of the coronary arteries, that it certainly cannot enter them. The whole of this doctrine seems to be supported by a fact, very familiar to accoucheurs, viz. the occur- rence of death in the foetus whenever the circulation through the umbilical cord issuspendedduringfifteen or twenty minutes: for as the placenta imparts to the foetal blood a quality essential to life, some arrange- ment seems necessary to provide for the equal distri- bution of the blood which comes from this organ, and " See Sabatier’3 Paper on this subject in the Memoirs of the Academy of Sciences, for 3774. Unusual Cases of Malformation . 77 especially for carrying the requisite proportion of it to the substance of the heart. Life has existed for some time with a structure very different indeed from that which is natural. In the series of elegant engravings relating to morbid ana- tomy, published by Dr. Baillie, is the representa- tion of a heart, in which the venae cavae opened into the right auricle, aftd the pulmonary veins into the left auricle, in the usual manner ; but the aorta arose entirely from the right ventricle, and the pul- monary artery as completely from the left. The canalis arteriosus, howevdr, passed from the pulmo- nary artery to the aorta, and the foramen ovale ex- isted. In this case it is evident that the pulmona- ry artery must have carried back to the lungs the arterial blood which came from them by the pulmo- nary veins, with a small quantity of venous blood that passed into the left auricle through the foramen ovale; add that the aorta must have returned to the body, the venous blood which just before had been brought from it by the venae cavae, with a small ad- dition of arterial blood that passed through the ductus arteriosuss Yet, with this structure, the child Ijjved two months after its birth, A case, which had a strong resemblance to the foregoing, occurred lately in Philadelphia, and was examined by the author of this work. The venae cavae terminated regularly in the right auricle, and the pulmonary veins in the same regular manner in the left ; but the pulmonary artery arose from the left ventricle, and the aorta from the right. There was no communication between these vessels by a ca- nalis arteriosus ; but a large opening existed in the septum between the auricles. It is very evident, that in this case also the pul- monary artery must have returned to the lungs the arterial blood as it came from them, and the aorta must have carried back to the general system the venous blood brought to the heart by the cavae ; ex- cepting only those portions of the arterial and venous blood which must have flowed reciprocally from one*auricle into the other, and thus changed their respective situations. 78 Foramen Ovale . The subject was about two years and a half old. Tiie heart was nearly double the natural size, and the foramen, or opening in the septum between the auricles, was eight or nine lines in diameter. The pulmonary artery was larger in proportion than the aorta or the heart. With this organization the child lived to the age above specified. His countenance was generally rather livid ; and this colour was always much in- creased by the least irregularity of respiration. His nails were alivays livid. He sometimes appeared placid, but more frequently in distress. He never walked, and seldom, if ever, stood on his feet. When sitting on the floor, he would sometimes push himself about the room; but this muscular exertion always greatly affected his respiration. He attain- ed the size common to children of his age, and had generally a great appetite. For some weeks before death his legs and feet were swelled. It is probable that the protraction of life depend- ed upon the mixture of the blood in the two auri- cles ; and that they really were to be considered as one cavity, in this case. There seems reason to -believe, thai^n adults of the common structure, there is no passage of blood from one auricle to the other, when the foramen ovale has remained .open ; because in several persons in whom it was found by dissection to have remained open, there were no appearances during life that in- dicated the presence of disoxygenated blood in the aortic system. It is probable, that the small size of the foramen ovale, the valvular structure which generally exists there, and the complete occupation of the left auricle by the blood flowing from the pul- monary veins, prevent the passage of blood from the right auricle to the left, in such persons ; whereas, in the case in question, the opening between the auricles was very large indeed, and there was no appearance of a valve about it. Although it be admitted, that* in adults with the foramen ovale pervious, there is no transmission of On the Source of the motion of the Heart. 79 blood from the right to the left auricle ; there is every reason to believe that this transmission goes on steadily in the fetus. To the arguments, derived from the structure and the nature of the case, it may be added, that the pulmonary veins, in the fetal state, carry to the left auricle a quantity of blood, not sufficient .to fill it; while the venae cavse carry to the right auricle not only the whole blood of the body, but of the umbilical cord and placenta : some of which must flow into the unfilled • .left auricle, when the right auricle becomes fully distended. The question how far the functions of the heart and lungs are dependent upon the brain is very impor- tant, and has often been agitated with great zeal. In favour of the opinion that the motions of the heart are independent of the brain, may be stated • the numerous cases in which the brain has been de- ficient in children, who have notwithstanding lived the full period of utero-gestation, and even a short time after birth, and have arrived at their full size, with every appearance of perfect vigour and action in the heart. In support of the doctrine, that the action of the heart is immediately dependent upon the brain, it may be observed, that no organ of the body appears to be so much influenced by passions and other mental affections as the heart. These contradictory facts have occasioned this question to be considered as undecided, if not incapable of so- lution ; although Cruikshank and Bichat* have stated circumstances very favourable to the opinion that the motions of the heart are independent of the brain. This question seems now to be settled by the ex- periments of Dr. Legallois, a physician of Paris, * See Cruikshank’s Experiments on the Nerves anc! Spinal Marrow of living animals ; London Philosophical Transactions for 1795. The eighth experiment has a particular relation to this subject. Bichat’s researches, part 2, article 9. The Abbe Fontana has considered this subject in bis Treatise on the Venom of the Viper, vol. ii. page 194, English translation ■ and also in same of his other works. 80 Humboldt and others on Legallois* Paper . which prove, that in animals who have suffered de- capitation , the action of the heart does not cease as an immediate consequence of the removal of the head ; but its cessation is an indirect effect , induced by the suspension of respiration. That respiration is immediately affected by decapitation, and depends upon the influence.of the brain transmitted through the eighth pair of nerves. That the action of the heart will continue a long time after decapitation, * if inflation of the lungs, or artificial respiration, be performed ; but, on the contrary, if the spinal mar - ' row be destroyed, the action of the heart ceases ir- recoverably. The inference from these experiments seems very conclusive, that the Spinal Marrow , and not the brain, is the source of the motions of the heart. It appears also by some of the experiments, that the power of motion in the trunk of the body is de- rived from the spinal marrow; and that when this or- gan is partially destroyed, the parts which receive nerves from the destroyed portion soon cease to live. By particular management of the spinal marrow, one part of the body can be preserved alive for some time after the other parts are dead. These experiments of Dr. Legallois commenced in 1806 or 1807, and were communicated to the Im- perial Institute of France in 1811. The committee of that body to whom they were referred, viz. Messrs. Humboldt, Halle and Percy, reported that the experiments had been repeated before them, at threfe different meetings of several hours each ; and that' to allow themselves sufficient time for reflec- tion, they suffered an interval of a week to take place between the meetings. The committee be- lieve these experiments to have proved, 1st. That the principle upon which all the move- ments of inspiration depend, has its seat about that part of the medulla obiongatafrom which the nerves of. the eighth pair arise. 2d. That the principle which animates each part of the trunk of the body is seated in that portion of the spinal marrow from which the nerves of the part arise. 3. That the source of the life and strength of the heart is also in the spinal marrow; not in any distinct portion, but in the whole of it. Brodie on the Source of the Motion of the Heart . 81 4th. That the great sympathetic nerve is to be consi- dered as originating in the spinal marrow, and that the particular character of this nerve is to place each of the parts to which it is distributed under the immedi- ate influence of the whole nervous power. The interesting memoir of Dr. Legal lois is confirmed to a certain degree by a communication of B. C. Brodie to the Royal Society of London in 1810, in which are detailed many very interesting experiments which in- duced the author to conclude, — That the influence’ of the brain is not directly ne- cessary to the action of the heart ; and That when the brain is injured or removed, the action of the heart ceases, only because respiration is under its influence ; and if under these circumstances respira- tion is artificially produced, the circulation will still continue. These various experiments apply particularly to the cases in which the brain is deficient. The effects of mental agitations on the heart are likewise reconcilable to the theory which arises out of them. But they throw no light on the question why the motions of the heart are so perfectly free from the influence of the will : and although they seem to prove incontestibly that the mo- tion of the heart is independent of the brain, it ought to be remembered that in certain diseased states of the brain, where that organ appears to be compressed, the action of the heart- is often very irregular, and its contractions less frequent than usual. VOL. II. 11 SYSTEM OF AX ATOMY. PART VIII. OF THE ABDOMEN. The lowermost of the two great cavities of the trunk of the body is called Abdomen. The pelvis may be considered as a chamber of this cavity, al- though its structure is very different. CHAPTER I. A GENERAL VIEW OF THE ABDOMEN AND PELVIS AND THEIR CONTENTS, WITH AN ACCOUNT OF THE PE- RITONEUM. , SECTION I. Of the Abdomen. This great cavity occupies more than half of the space enclosed by the ribs, and all the interior of the trunk of the body below the thorax. It is formed by the diaphragm, supported by the lower ribs ; by a portion of the spine ; by the vari- ous muscles which occur between the lower margin of the thorax and the upper margin of the ossa in- nominata ; and by the ossa innominata, which con- tribute, for the purpose, the costas of the ossa ilia, as well as the pelvis. S3 Construction of the Abdomen. The general figure of this cavity partakes of the figure of the lower part of the trunk of the body ; with these exceptions, that the diaphragm makes it arched or vaulted above, that the spine and psose muscles, &e. are rather prominent on the posterior surface, and that the lower part corresponds with the costas of the ossa ilia and with the pelvis. To acquire a precise idea of this cavity, it is ne- cessary first to study the bones concerned in its structure, in their natural situation in the skeleton ; and then the muscles, which form so large a part of it. The arrangement of the tendons of some of these muscles, with a view to complete the cavity, is par- ticularly interesting ; as that of the external oblique where it forms the crural arch.* The ligaments of the pelvis and the levatores ani muscles, as they also contribute to the formation of the cavity, and have an influence upon its figure, should likewise be at- tended to. In the walls of the cavity, thus constructed, there are many foramina by which the viscera and other contained parts communicate externally ; but few of them pass directly into the cavity ; for like the tho- rax, there are no vacuities in- it exterior to the con- tained organs. Three of these foramina are in the diaphragm. One for the transmission of the aorta, another for the vena cava, and a third for the oesophagus. Below, there is an aperture at each of the crural arches, for the transmission of the great femoral vessels ; in each of the ligamentous membranes, which close the fora- men thyroideum, for the obturator vessels and nerve; and at the sacro sciatic notches, for nerves and blood vessels. * See the account of this tendon, vol. i. in the description of the “ Ob* liquus Decendens Externus.” 84 Construction of the Abdomen, There are also two apertures at the bottom of the pelvis, for the orifice of the rectum and of the ure- thra. In the tendons of the external oblique mus- cles are two orifices, covered by the integuments, for the spermatic cords ; and, in the foetal state, one for the umbilical cord. The apertures in the tendons, and under their edges, for the transmission of the spermatic cords, and the blood vessels, &c. are not to be considered as simple perforations made abruptly; but the edges of these foramina are formed by tendinous mem- branes turned inwards and continued so as to com- pose a cylindrical tube, which becomes gradually so thin that it cannot be readily distinguished from the cellular membrane with which it is connected.* The blood vessels, &c. pass along this tube before they go through the apertures. It is evident from the construction of this cavity that it is essentially different from the thorax. It has no power of spontaneous dilatation whatever : it yields passively to the distention of the stomach and intestines, during deglutition, and when air is extri- cated from the aliment, &c. ; but it is particularly calculated for compressing its contents by the con- traction of the muscles* which compose it. The di- minution of its capacity, which is thus effected, not only takes place to a great degree, but occasionally with great force. The diaphragm and the abdomi- nal muscles may be considered in some measure as antagonists of each other. When the diaphragm descends, if the abdominal muscles are passive, they are distended by the contents of the abdomen, which are forcibly pressed from above ; but if the abdomi- nal muscles act at the same time, an effort to dimi- * The [student of anatomy, ■when engaged with this subject, will be gratified by the examination of Mr. Astley Cooper’s plates relating to hernias. Contents of the Abdomen. 85 msh the cavity in every direction takes place, and the contained parts are compressed with more or less force according to the exertion made. This will be very evident upon examining the situation of the diaphragm and of the abdominal muscles. When their force is considered it will also be very obvious that the vari- ous outlets of the cavity are constructed most advan- tageously ; otherwise hernia or protrusion of its con- tents would be a daily occurrence. The abdomen contains, 1st. The Stomach and the whole Intestinal Tube, consisting of the small and the great intestines. 2d. The Assisting Ckylopoietic Viscera , — the Liver, the Pancreas and the Spleen. 3d. The Urinary Organs , — the Kidneys, the Ureters, and the Bladder. To which should be add- ed the Glandulae Renal es. 4. The Organs of Generation in part: those of the female sex being almost wholly included in the pelvis ; and those of the male being situated partly within and partly without it. 5th. The Peritoneum and its various processes. The Mesentery, Omentum, &c. 6th. A portion of the Aorta, and almost the whole of the Inferior Cava, and their great ramifi- cations ; with such of' their branches as are appro- priated to the Viscera of the Abdomen and Pelvis. 7th. Those portions of the Par Vagum and In- tercostal Nerves which are appropriated to the cavi- ty ; and portions of some of the nerves destined to the lower extremities. . 8th. The lower part of the Thoracic Duct, or the Great Trunk of the Absorbent System, with the large branches that compose it, and the glands con- nected with them ; and also those absorbent vessels called Lacteals, and their glands. As the cavity of the abdomen has no natural divi- 86 Regions of the Abdomen . sions, anatomists have divided it by imaginary lines into various regions, with a view to precision in their accounts of the situation of the different contained parts. Thus, They have, very generally, agreed to apply two transverse lines to form three great divisions ; viz. the Upper , Middle, and Lower : and they have also agreed that each of these divisions shall be subdivid- ed into three regions. The three regions of the uppermost division are defined with some precision. Those on each side, which are called the Right and Left Hypochondriac regions, occupy the spaces immediately within the lower ribs and their cartilages ; while the middle space, included within the margins of these cartila- ges, and a line drawn from the lower edge of the thorax on one side to that on the other, is denomi- nated the Epigastric region. The boundaries of the regions below are less pre- cisely defined. Many anatomists have fixed the two transverse lines above mentioned at an arbitray distance above and below the umbilicus : some choosing for this pur- pose two inches, and others a hand’s breadth. As these distances will occupy different proportions of the cavity in persons of different stature, other ana- tomists, with a view to avoid this inconvenience, have proposed to connect these lines With certain fixed points of the skeleton. It is of importance that the boundaries of these regions should be fixed, and therefore the proposi- tion of Sabatier may be adopted ; viz. To draw the upper transverse line from the most inferior part of the lower margin of the thorax, on one side, to the corresponding part on the opposite side ; and the lower transverse line from the uppermost part of the 87 Regions of the Abdomen. spine of one ilium to the same part of the other. These lines will mark the three great divisions. If then two parallel lines are drawn directly upwards, one from each of the superior anterior spinous pro- cesses of the ilium until it touches the lower margin of the thorax, they will divide each of the two lower divisions of the abdomen into three regions. The centre of the middle division is the umbilical, and on each side of it is the right and left lumbar region. The middle of the lower division is the hypogastric ; and on each side of it the right and left iliac region. It is true, that the three middle regions of the ab- domen will be made very small by the vicinity of the transverse lines to each other ; but the advantages derived from a principle which is similar in its ap- plication to all subjects fully compensates for this in- convenience. There are therefore nine of these regions : viz. The Epigastric and the two Hypochondriac: the Um- bilical, and the two Lumbar : the Hypogastric, and the two Iliac regions.* And it should be added, that the space immediately around the end of the ster- num is sometimes called the Scrobiculus Cordis ; and the space immediately within the os pubis, the Regio Pubis. These different regions are generally occupied in the following manner. The liver fills nearly the whole of the right hypochopdriac region, and ex- tends through the upper part of the epigastric region into the left hypochondriac. The stomach occupies the principal part of the epigastric region, and a considerable portion of the left hypochondriac. The spleen is also situated in the left hypochondriac re- * It is to be observed that the lateral regions of the middle and lower di- visions of the abdomen are named differently bv different writers. 38 Situation of the Viscera, 6,'C. in the gion. That portion of the intestinal tube, which is composed of small intestines, is generally found in the umbilical, the hypogastric, and the iliac regions ; and when the bladder is empty, in the pelvis. But the duodenum, or first of the small intestines, which proceeds immediately from the stomach, is situated in the epigastric and umbilical regions. The great intestine commences in or near the right, iliac region, and ascends through the right lumbar* to the right hypochondriac region. It then crosses the abdomen, passing through the lower part of the epigastric, 01 s upper part of the umbilical to the left lumbar region ; from this it continues into the left iliac region, and curves in such a manner that it finally arrives at the middle of the upper part of the os sacrum, when it descends into the pelvis, and, partaking of the cur- vature of the last mentioned bone, continues to the termination of the os coccygis. In the back part of the epigastric region, and very low down in it, is situated the pancreas. The kid- neys lie in the most posterior parts of the lumbar re- gions, and from each of them is continued a tube or duct, called Ureter, that passes into the pelvis to con- vey the urine to the bladder. This viscus, in males, is in contact with the last portion of the great intes- tine called the Rectum, and with it occupies almost all the cavity of the pelvis ; while in females, the uterus and its appendages are situated between this intestine and the bladder. In the posterior part of the abdomen, in contact with the spine is the aorta. This great blood ves- sel passes from the thorax between the crura of the diaphragm, and continues down the spine until it approaches towards the pelvis, when it divides into two great branches called the Iliac Arteries. Each of these great branches divides again, on the side 89 Cavity of the Abdomen. of the pelvis, into two ; viz. the External Iliac , which passes under the crural arch to the thigh, and the Internal Iliac, or Hypogastric , which descends into the cavity of the pelvis. Soon after the arrival of the aorta in the abdomen it gives off two large branches. The first, which is called the Cceliac, is distributed to the liver, the sto- mach, and the spleen : the second, called the Supe- rior Mesenteric, is spent upon the intestines. Lower down, in the abdomen, it also sends off a small branch for the intestines, called Inferior Mesen- teric. Besides these vessels for the chylopoietic vis- cera, the aorta sends off a large branch, called Emul- gent, to each kidney. The inferior or ascending vena cava is situated on the right of the aorta, in front of the spine. It is formed below by the union of the iliac veins, and in its progress upwards it receives the emulgent veins, which correspond to the arteries of the kidneys ; but it receives in its course no veins which correspond directly with the cceliac and mesenteric arteries. The smaller veins, that answer to the branches of these arteries, unite and form one large vein, which goes to the liver, and is called, (from the part of that viscus at which it enters,) Vena Portarum. From the liver three large veins pass into the vena cava, and deposit there the blood of the vena portarum, after it has furnished materials for the secretion of bile. The vena cava, in its passage upwards, is in close contact with the posterior .thick edge of the liver : it often passes along a deep groove in this edge, and sometimes it is completely surrounded by the liver in its course. The veins of the liver enter the vena cava at this place, and of course they are not to be seen without dissection. Immediately af- ter leaving the liver the vena cava passes through an Vol. ii. 12 90 The Peritoneum. aperture in the tendinous centre of the diaphragm to unite itself to the right auricle of the heart. SECTION II. Of the Peritoneum. The abdomen, thus constructed and occupied, is lined by a thin firm membrane called Peritoneum , which is extremely smooth on its internal surface, and is immediately connected with the cellular sub- stance exterior to it. This membrane adheres close- ly to the anterior, lateral, and superior portions of the surface of the abdomen ; and is extended from the posterior surface so as to cover, more or less com- pletely, the viscera of the cavity. Those viscera which are in close contact with the posterior surface of the abdomen, as some portions of the large intes- tine, are covered only on their anterior surfaces, and are fixed in their precise situation by the peritone- um ; which extends from them to the contiguous surface of the cavity, and adheres where it is in con- tact, so as to produce this effect. Other viscera, which are not in close contact, but moveable to a distance from the posterior surface of the abdomen, are covered by this membrane, which is extended to them from the surface ; and this ex- tended portion forms an important part of the con- nexion between the viscus and the cavity in which it lies. This connecting part is called Mesentery , when it thus passes to the small intestines; Mesoco- lon ., when it goes to the colon, one of the larger in- testines ; and Ligoment , when it passes to some of the other viscera. The peritoneum is a complete but empty sac, which is fixed in the abdomen anterior to the viscera. The anterior portion of this sac forms the lining to the anterior and lateral parts of the surface of the The Peritoneum. 91 abdomen : the posterior portion covers the viscera,, and forms the mesentery, mesocolon, and ligaments above described. * It necessarily follows that the mesentery and the other similar processes are mere plaits or folds of the sac, which invests the viscera ; and that they must consist of two lamina ; and as the blood vessels, nerves, and absorbents, are all posterior to the peri- toneum, they naturally pass between these lamina of the mesentery. Some of the viscera are much more completely in- vested with the peritoneum than others. The sto- mach, liver, and spleen, are almost completely surrounded by it ; and it is said to form a coat for each of these viscera. That portion of the smaller intestinal tube, which is called jejunum and ileum, and the transverse portion of the large intestine, call- ed the arch of the colon, are invested by it in the same way. But a considerable portion of the duo- denum and the pancreas is behind it. The lateral portions of the colon are in close contact with the posterior surface of the abdomen, and the peritone- um only covers that portion of their surfaces which looks anteriorly towards the cavity of the abdomen, and is not in contact with its posterior surface. The urinary organs are not much connected with the peritoneum. The kidneys appear exterior to it, and behind it: the bladder of urine is below it, and has but a partial covering from it, on its upper por- tion. The peritoneum, which covers the stomach, is ex- tended from the great curvature of that organ so as to form a large membrane, which descends like an apron before the intestines. This process of perito- neum is composed of two lamina, so thin and deli- cate as to resemble cellular membrane, which, after extending downwards to the lower part of the abdo- 92 The Peritoneum. men, are turned backwards and upwards, and pro- ceed in that direction until they arrive at the colon, which they enclose, and then continue to the back of the abdomen, forming the mesocolon. The part of this process which is between the stomach and the colon, is called Epiploon or Omentum. This extension of a membrane, from the surface of a cavity, which it lines to the external surface of a viscus in that cavity, is called, by some anatomists, “ reflection and the technical term reflected mem- brane is therefore applied to a membrane distributed like the peritoneum. It must be evident that this distribution of the pe- ritoneum is very complex, and that* it is not easy to form an accurate conception of it from description, but it can be readily understood by demonstration ; therefore no further account of its arrangement will now be attempted, but each of its processes will be considered with the organs to which they are parti- cularly subservient. That portion of the peritoneum which lines the abdomen and covers the viscera, is thin and delicate, but very firm. It yields to distension, as in preg- nancy, ascites, &c. and again recovers its dimen- sions. It was formerly thought to be composed of two lamina, but this cannot be proved. The internal surface of this membrane is very smooth, and highly polished ; and from it exudes a liquor which is well calculated for lubrication, and barely sufficient to keep the surface moist during health ; but sometimes it is very abundant, and occasions the aforesaid dis- ease — ascites. This fluid appears to exude from the surface of the peritoneum when it is compressed in a living animal, or in one recently dead. It is probably efl'used from the extremities of arteries, for an effusion takes place when water is injected into these vessels. The Peritoneum . 93 The peritoneum abounds with absorbent vessels, and therefore possesses the power of absorption to a great degree. This power may be inferred, not only from the spontaneous removal of the fluid of ascites, but if milk and water be introduced into the abdomen of a living animal, through a puncture, it will also disappear. The blood vessels of the peritoneum are derived from those which supply the neighbouring parts. Nerves have not yet been traced into it, and it has little or no sensibility. This membrane supports the viscera of the abdo- men in their proper situations ; and also forms a sur- face for them, and for the cavities which contains them, so smooth and lubricated, that no injury can arise from their friction. The cellular substance, by which the peritoneum is connected to the contiguous parts, is very different in different places. It is very short indeed between this membrane and the stomach and intestines, and also between it and the tendinous centre of the dia- phragm. Between the peritoneum and the muscles generally, it is much longer. When it covers the kidneys and the psoas muscles it is very lax and yielding. About the kidneys a large quantity of adeps very commonly collects in it. On the psoas muscle it yields with but little resistance to the pas- sage of pus, or any other effused fluid, as in the case of the psoas abscess. CHAPTER II. OF THE OESOPHAGUS, THE STOMACH, AND THE IN- TESTINES. SECTION I. Of the (Esophagus . The (Esophagus is a muscular tube which passes from the pharynx to the stomach, and is so intimate- ly connected with the stomach, that it will be advan- tageous to the student to attend to its structure im- mediately before he engages in the examination of that important organ. The pharynx has been lately described* as com- posed of a varied stratum of muscular fibres, lined by a membrane which is continued from the internal sur- face of the nose and mouth. From the pharynx the oesophagus passes downwards between the trachea and the vertebrae. After the bifurcation of the tra- chea, it proceeds in contact with the spine, between the lamina of the mediastinum, to the diaphragm, which it passes through, and then terminates in the stomach. The oesophagus is a flexible tube, which, when distended, is nearly cylindrical. It consists of a muscular coat externally, and an internal tunic evi- dently continued from that of the pharynx. These coats are connected by a cellular substance called the Nervous Coat , which is remarkably loose, and allows them to move considerably upon each other. The muscular coat which is very distinguishable from that of the pharynx, consists of two substantial strata of fibres ; the exterior of which is nearly lon- * See page 37. The (Esophagus. 95 • gitudinal in its direction, and the interior circular or transverse. The internal coat of the oesophagus, resembling that of the fauces, is soft and spongy. It is covered with a very delicate cuticle, which Haller supposed to be to tender to confine the matter of variolous pustules, as he had never found these extending into the oesophagus. It is very vascular, and abounds with the orifices of mucous follicles, from which is constantly poured out the mucous that is spread over this surface. When the oesophagus is not distended, many longitudinal plaits are found in this membrane by the contraction of the circular or transverse fibres exterior to it. These plaits are calculated to admit readily of the distention which is requisite in deglu- tition. This tunic is continued from the lining mem- brane of the pharynx above, and terminates below in the villous coat of the stomach ; from which, howev- er, it is very different. The blood vessels of the oesophagus come from those which are in the vacinity. The nerves are derived from the eighth pair. The lymphatic ves- sels are very abundant. In the neck, the oesophagus inclines rather to the left of the middle line. As it proceeds down the back between the lamina of the mediastinum, it pre- serves the same course to the fourth dorsal vertebra, when it assumes the middle portion and proceeds downwards, with the aorta to its left, and the peri- cardium before it. About the ninth dorsal vertebra it inclines again rather to the left, and somewhat for- ward, to arrive at the aperture in the diaphragm through which it passes. Throughout this course it is connected by cellular membrane to the contiguous parts ; and this investi- ture of cellular membrane has been called its Ex- ternal Coat, 96 Form of the Stomach. While the oesophagus is in the posterior mediasti- num, it is in contact with several small absorbent glands, especially when it first assumes a situation to the right of the aorta. These glands were for- merly believed to be particularly connected with this tube, but they are now considered as belonging to the absorbent system. They are sometimes great- ly enlarged. SECTION II. Of the Stomach. This most important organ which occasionally exerts a powerful influence upon every part of the body, appears very simple in its structure. It is a large sac, which is so thin when much in- flated that at first view it seems membranous, but upon examination is found to be composed of several lamina or coats, each of a different structure. It is of considerable length, but incurvated. It is much larger at one extremity than the other, and changes so gradually in this respect, that it would appear conical if it were straight. It is not, however, strictly conical, unless it is greatly distended ; for when moderately distended, a transverse section is rather oval than circular. It is therefore considered as hav- ing two broad sides or surfaces, and two edges, which are the curvatures. It has been compared by the anatomists of different nations to the wind sac of the musical instrument called the bagpipe.* The orifice in which the oesophagus terminates is at a small distance from its largest extremity, and is called Cardia. The orifice which communicates * The student ought not to attempt to acquire an idea of the form of the stomach without demonstration, for a view of one moment will be more serviceable than a long description. Position of the Stomach . 97 with the intestines is at the termination of its small incurvated extremity, and is called the Pylorus, The two ends of the stomach being thus very dif- ferent in size, are denominated the great and small extremities. The two curved portions of the surface are also called the great and small curvatures. The two flat portions of the surface, or the broadsides, are called the anterior and posterior surfaces. The situation of the stomach in the abdomen is nearly transverse : it lies principally in the left hy- pochondriac and epigastric regions, immediately be- low the liver. The great extremity of the stomach is in the left hypochondriac region, and the lesser extremity in the epigastric region, under the left lobe of the liver. The upper orifice, or Cardia, is nearly opposite to the body of the last dorsal ver- tebra ; and owing to the curved form of the stomach, the other orifice, or Pylorus , is situated at a small distance to the right of that bone, and rather lower and more forward than the cardia ■: both orifices be- ing in the epigastric region. The position of the stomach is oblique in two respects ; it inclines in a small degree from above downwards, from the left to the right ; and it also inclines downwards and forwards, from behind. Its two orifices are situated obliquely with respect to each other ; for, if the sto- mach, when placed with its small curvature upwards, were divided into two equal parts by a vertical plane passing lengthways through it, they would be found on different sides of the plane. As the oesophagus terminates in the stomach im- mediately after it has passed through an aperture of the diaphragm, it is evident that the stomach must be somewhat fixed at that place ; but it is more move- able at its other orifice ; for the extremity of the duo- denum, into which it is continued, is moveable. Vol. ix. 13 98 External Coat of the Stomach. The stomach is connected to the concave surface of the liver by the reflexion or continuation of the peritoneum which forms the lesser omentum. This membrane, after extending over each surface of the stomach continues from its great curve in the form of the large omentum, and connects it to different parts, especially to the colon. There are likewise folds of the peritoneum, as it passes from the dia- phragm and from the spleen to the stomach, which appear like ligaments. Notwithstanding these various connexions, the stomach undergoes considerable changes in its posi- tion. When it is nearly empty, and the intestines are in the same situation, its broad surfaces are pre- sented forwards and backwards ; but when it is dis- tended, these surfaces are presented obliquely up- wards and downwards, and the great curvature for- wards. When its anterior surface is presented up- wards, its urifices are considerably influenced in their direction, and the oesophagus forms an angle with the plane of the stomach. The stomach is composed of four dissimilar lamina, which may be demonstrated by a simple process of dissection. There is first a coat or external covering conti- nued from the peritoneum : within this, and connect- ed to it by delicate cellular substance, is a coat or stratum of muscular fibres : contiguous to these fibres, internally, is a layer of dense cellular substance, call- ed a nervous coat ; and last is the internal coat of the stomach, called villous or fungous, from the structure of its surface. The external or first coat of the stomach, as has been already stated, is continued from the concave surface of the liver to the lesser curve of the sto- mach in two delicate lamina, which separate when they approach the stomach, and pass down, one on i Muscular and Nervous Coats of the Stomach . 99 each side of it, adhering firmly to it in their course : at the opposite curve of the stomach they again unite to form the great omentum. The stomach is there- fore closely invested by the peritoneum on every part of its surface except two strips, one at the lesser and the other at the greater curvature. These strips or uncovered places are formed by the separation of the lamina above mentioned, which includes a triangular space bounded by the stomach and these two lamina. In these triangular spaces, at each curvature of the stomach, are situated the blood vessels which run along the stomach in those directions, and also the glands which belong to the absorbent vessels of this viscus. The peculiar arrangement of the lamina at this place is particularly calculated to permit the dila- tation of the stomach. When it is dilated the lamina are in close contact with its surface, and the blood vessels being in the angle formed by the adhesion of the two lamina to each other, are so likewise : when it contracts, the blood vessels appear to recede from it, and the lamina are then applied to each other. Where the peritoneum thus forms a coat to the sto- mach, it is stronger and thicker than it is between the liver and stomach. In a recent subject it is very smooth and moist, but so thin that the muscular fibres, blood vessels, &c. appear through it. If it is care- fully dissected from the muscular coat, it appears somewhat flocculent on that surface which adhered to the muscular fibres. It seems to be most abundant- ly furnished with serous vessels ; but it has been as- serted by Mascagni and Soemmering, that a large proportion of its texture consists of absorbent ves- sels. The cellular substance which connects this to the muscular coat appears no way different from or- dinary cellular membrane. The Muscular Coat of the stomach has been de- scribed very differently by respectable anatomists ; a 00 Internal Coat of the Stomach. some considering it as forming three strata of fibres? and others but two. If the stomach and a portion of the oesophagus attached to it be moderately distended with air, and the external coat carefully dissected away, many longitudinal fibres will appear on every part of it, that evidently proceed from the oesophagus; these fibres are particulary numerous and strong on the lesser curvature of the stomach. — Beside the lon- gitudinal fibres there are many that have a circular direction, and these are particularly numerous to- wards the small extremity ; but it has been doubted whether there are any fibres in the muscular coat of the stomach that go directly round it. The whole sur- face of the stomach, when the peritoneal coat is re- moved, appears at first view to be uniformly covered by muscular fibres ; but upon close examination, there are interstices perceived, which are occupied with firm cellular membrane. In contact with the internal surface of the mus» cular coat is the cellular stratum , which has been called the JYervous Coat of the stomach. It is dense and firm, of a whitish colour, resembling condensed cellular membrane. It was considered as different from ordinary cellular membrane ; but if air be in- sinuated into its texture, by blowing between the muscular and villous coats, while it connects them to each other, it exhibits the proper appearance of cel- lular substance. It however adds greatly to the ge- neral strength of the stomach, and the vessels which terminate in the villous coat ramify in it. The internal coat of the stomach in the dead sub- ject is commonly of a whitish colour, with a tinge of red. It is named villous, from, its supposed resem- blance to the surface of velvet. It has also been called fungous, because the processes analogous to the villi are extremely short, and its surface has a Gastric Liquor. 101 granulated appearance 5 differing in these respects from the internal surface of the intestines. It is con- tinued from the lining membrane of the oesophagus, but is very different in its structure. Many very small vessels seem to enter into its texture, which are derived from branches that ramify in the nervous coat. It is supposed by several anatomists of the highest authority, to have a cuticle or epithelium ; and it is said that such a membrane has been sepa- rated by disease. It ought however, to be remem- bered, that the structure of the villous coat of the stomach and intestines, is essentially different from the structure of the cuticle. The internal coat of the stomach is generally found covered, or spread over, with mucous, which can be readily scraped off. This mucous is certainly effus- ed upon it by secreting organs, and it has been sup- posed that there were small grandular bodies exte- rior to the villous coat, which furnished this secre- tion ; but the existence of such bodies is very doubt- ful, as many skilful anatomists have not met with any appearance that could be taken for glands, ex- cept in a very few instances, which would not be the case if those appearances had been natural. Pores, perhaps the orifices of mucous follicles, and also of exhalent vessels, are very numerous, but no proper glandular masses are attached to them. Glands, as have been already said, are found in the triangular spaces between the lamina of the peritoneum at the great and small curvatures of the stomach, but these evidently belong to the absorbent system. Besides the mucous above mentioned, a large quantity of a different liquor, the proper Gastric Juice , or fluid of the stomach, is effused from its surface. It nas been supposed that this fluid is furnished by the small glandular bodies believed to exist between the coats 102 The Pylorus. of this origin ; but, admitting the existence of these glands, they are not sufficiently numerous to produce so much of it as is found, and it is therefore probable that this fluid is discharged from the orifices of ex- halent vessels in the internal surface. Much information respecting the gastric liquor has been obtained within a few years past by the researches of physiologists, and they are generally agreed that it is the principal agent in the effects produced by the stomach upon alimentary substan- ces.* - As the muscular coat of the stomach frequently varies its dimensions, trie villous and nervous eoats, which have no such power of contraction, cannot ex- actly fit it. They therefore generally appear larger, and of course are thrown into folds or rugae. These folds are commonly in a longitudinal direction ; but at the orifices of the stomach they are arranged in a radiated manner, and sometimes they are observed in a transverse direction. They depend upon the contraction of the muscular fibres, and disappear en- tirely when the stomach is laid open and spread out. At the lower orifice is a circular fold, which is permanent, and constitutes the valve denominated * On this subject the student may consult with advantage. M. Reaumur. In the Memoirs of the Academy of Sciences for 1752. John Hunter. London Philosophical Transactions for 1772 ; and also his observations on the Animal Economy, 1786. Dr. Edward Stevens. Inaugural Thesis de Alimentorum Concoctione. Edinburgh, 1777. The Abbe Spalanzani. Dissertations relative to Natural History, Szc. The first volume of the English translation contains the author’s disserta- tions on digestion, and also the first paper of Mr. Hunter, and the Thesis of Dr. Stevens, as well as an account of the experiments of Mr. Gosse of Geneva. In addition to these, there are several interesting essays in the French, German, and Italian languages, a compilation of which is to be found in Johnson’s “ History of the progress and present state of Animal Chemistry.” See Vol. I. page 180. Lymphatics and Nerves of the Stomach. 1Q3 Pylorous. It appears like a circular septum with a large foramen in its centre, or like a flat ring. The villous and nervous coats of the stomach contribute to this, merely by forming the circular fold or ruga ; and within this fold is a ring of muscular fibres, evi- dently connected with the circular fibres of the mus- cular coat of the stomach, the diameter of which at this place is not larger than that of an intestine : the fibres of this ring seem a part of the muscular coat projecting into the cavity of the stomach and duode- num. If a portion of the lesser extremity of the sto- mach and the adjoining part of the duodenum be de- tached, and laid open by a longitudinal incision, and then spread out upon a board, the internal coat can be very easily dissected from the muscular, and the py- lorous will- then appear like a ridge or narrow bundle of muscular fibres, which runs across the extended muscular membrane. It is evident that when the parts are replaced so as to form a cylinder, this nar- row fasciculus will form a ring in it. Thus arrang- ed, the circular fibres can readily close the lower ori- fice of the stomach. The pylorus separates the stomach from the intes- tine duodenum ; and this separation is marked exte- riorly by a small circular depression, which corres- ponds exactly with the situation of the pylorus. The arteries of the stomach are derived from the Coeliac, the first branch which the aorta sends off to the viscera of the abdomen. This great artery, im- mediately after it leaves the aorta, is divided into three branches, which are distributed to the stomach, the liver, and the spleen, and are called the Superior Cm'onary or Gastric, the Hepatic , and the Splenic. Beside the first mentioned branch, which is distri- buted principally to the neighbourhood of the cardia and to the lesser curvature, the stomach receives a considerable, branch from the hepatic, which passes 104 The Intestines in general. along the right portion of its great curvature, and has been called the right gastro-epiploic, and another from the spleen, which passes along the left portion of the great curvature, and has been called the left gastro-epiploic. In addition to these branches, the splenic artery, before it enters the spleen, sends off several small arteries to the great extremity of the stomach, which are called vasa brevia. These vasa brevia generally arise from the main trunk of the splenic artery, but sometimes from its branches. •The veins which receive the blood from these ar- teries have similar names, and pursue corresponding courses backwards ; but they terminate in the vena portarum. The absorbent vessels of the stomach are very nu- merous and large : they pass to the glands which are on the two curvatures, and from thence to the thoracic duct. It is an important fact relative to the history of digestion, that there are good reasons for doubting whether chyle commonly passes through them, not- withstanding their number and size.* The nerves of the stomach are derived principally from the two great branches of the par vagum, which accompany the oesophagus and are mostly spent upon this organ. It also receives branches from several plexus, which are derived from the splanchnic por- tions of the intercostal nerves. SECTION III. Of the Intestines. The intestines form a Continued canal from the pylorus to the anus, which is generally six times ' Sabatier, however, in one subject observed white lines on the stomach, which he suspected to be lacteals, See his account of the absorbents of the stomach. 105 Villous Coat of the Intestines, the length of the subject to which they belong. Al- though the different parts of this tube appear some- what different from each other, they agree in their general structure. The coats or lamina of which they are composed, are much like those of the stomach, but the peritoneum which forms their external coat does not approach them in the same manner ; nor is it continued in the form of omentum from the whole tube, there being only a certain portion of intestine, viz. the colon, from which such a process of perito- neum is continued. The Muscular Coat, like that of the stomach, consists of two strata, the exterior of which is com- posed of longitudinal fibres, which adhere to the external coat, and do not appear very strong. The other stratum, consisting of circular or transverse fibres, is stronger, as the fibres are more numerous. It is observable that they adhere to the longitudinal fibres: and they seldom if ever form complete cir- cles. The cellular substance immediately within the muscular fibres resembles the 'nervous coat of the stomach in its firmness and density. It is likewise so arranged as to form many circular ridges on its in- ternal surface, which support to a certain degree the permanent circular plaits of the internal coat, called valvulze conniventes. The inner surface of the internal coat has been commonly compared to that of velvet, and the coat is therefore called villous ; but there is certainly a considerable difference between these surfaces ; for if a portion of the small intestine be inverted, and then suspended in perfectly transparent water, in a clear glass, and examined with a strong light, it will appear like the external surface of the skin of a peach, on which the down or hair-like processes are not so close as those on velvet. On this surface, Vol* ii. 14 106 Lieberkuhn on the Villous Coat. between the villi, there are many orifices of mucous follicles and of exhaling vessels.* Exterior to the villous coat, many very small glandular bodies are sometimes found, which are called after their descri- bers Glandulse Brunneri and Peyeri. The internal coat of the upper portion of the in- testinal tube is arranged so as to form a great num- ber of transverse or circular folds or plaits^ called Valvulse Conniventes, which do not generally extend round the intestine, but are segments of circles; they are so near each other that their internal edges, which are very moveable, may be laid upon the folds next to them, like tiles or shingles. It is evident that this arrangement of the internal coat must add greatly to its length. This coat is extremely vascu- lar, so that in the dead subject it can be uniformly coloured by a successful injection. The minute struc- ture of it has been the subject of very diligent inquiry. There can be no doubt but that an immense number of exhaling and of absorbent vessels open upon it; but there are many different opinions respecting the termination of one set of vessels and the commence- ment of the other. A very interesting account of the Villous Coat was published in 1744, by Lieberkuhn, who was con- sidered by his contemporaries as a most expert prac- tical anatomist, and was also very skilful in micro- scopical examinations, for which he was particularly calculated, as his natural powers of vision were un- commonly strong. In his essay he refers to his preparations, which were at Berlin, and which ap- pear to have excited great surprise in the minds of * It appears clearly, from the account of Lieberkuhn, that the ori- fices or terminations of the arteries on the intestines, are distinct from the follicles ; for he forced injection from the arteries into the cavity of the intestines, and found the follicles still filled with mucous. He then urged the injection further, and filled the follicles, or forced the mucous out of tfeeat. Hewson and others on the Villi. 107 the members of the Academy of Sciences of Prussia, at a time when one of the first anatomists of Europe, the celebrated Meckel, was of their number. According to this account, the internal surface of the small intestines abounds with villi, and with the orifices of follicles. These villi are ^about the fifth part of a line in breadth. In each of them is a cavity filled with a soft spungy substance, which has one or more orifices communicating with the intestines, and from which also proceeds a lacteal vessel. On the membrane which forms this cavity, blood vessels are most minutely ramified. This cavity he calls an ampullula, and supposes it to constitute the principal part of the villous. By injecting the arteries of the intestine, he was able to pass a fluid through the ampufluia into the cavity of the gut ; he kept a stream of air in this way passing through the am- pullula until it was nearly dry and stiff, and then laid it open with a fine instrument. From the ap- pearances which then presented, he inferred that the cavity of the ampullula was occupied with a spongy or cellular substance. Around each villus he found a number of mucous follicles, which often were filled with a tenacious mucous: and distinct from these must be the exhalent orifices, which discharged a fluid injected by the arteries without passing through the mucous follicles. Lieberkuhn died early, and left but one essay on this subject, which was originally published in Hol- land, in 1744, but has been republished by the Aca- demy of Berlin, in their Memoirs ; and also by Mr. John Sheldon, of London. This account of Lieberkuhn appears to have been admitted by Haller ; but it has been rigidly scrutinized by some of the anatomists of London, who were particularly interested with the subject; as they had paid great attention to the absorbent 108 Fyfe and others on the Villi . system, and were very successful in the investiga- tion of it. The late Mr. Hewson, whose opinion is entitled to the greatest respect, rejected the idea of the ampul- lula, and believed that the villi are composed of net- works of lacteals, as well as arteries and veins ; al- though he added that (i this is the only circumstance concerning these parts in which he should differ from this very acute observer.”* Mr. Sheldon agrees with Lieberkuhn : but Mr. Cruikshank asserts, that, “ in some hundred villi, be has seen the lacteals originate by radiated branches, w hose orifices were distinct, on the surface of the villus.” The villus being transparent, when the intestine was immersed in water, these branches, filled w ith chyle could be seen passing into the lac- teal. Mr. Cruikshank therefore supposes that Lie- berkuhn was mistaken, and that the spongy cavity, or ampullula, was the common cellular membrane, connecting together all the arteries, veins, nerves, and lacteals. It seems probable, from Mr. Cruikshank’s state- ment, that Dr. William Hunter held the same opinion with himself. And there is also reason to believe that Monro the second, who studied anato- my at Berlin, held a different opinion from Lieber- kuhn. Mr. Fyfe, who has bedn much employed in the investigation of the absorbent system, and must be perfectly acquainted with the'preparations of Monro, asserts that each lacteal takes its rise upon one of the villi by numerous short radiated branches, and each branch is furnished with an orifice for imbibing chyle. bieveral of the late French writers adopt the opi- * See Hewson’s Experimental Inquiries, vol, 2, page 171. 109 Division of ihe Intestines „■ nion of Lieberkuhn ; but his countryman Soemmer- ing gives a different account of the subject. He says, that,, besides the blood vessels, each villus consists of a fine net-work of absorbent vessels, whose orifices may be distinctly recognised ; and that from six to ten of these orifices are sometimes discovered. Mascagni, who has published the most extensive work upon the absorbent system that has yet appear- ed, supposes Lieberkuhn to have been mistaken, and confirms the description of Hewson : but he also agrees with Hewson in his opinion of the general ac- curacy of Lieberkuhn. Notwithstanding their differences respecting the origin of thelacteals, all these observers have agreed, that the orifices which communicate with the lacteals are on the villi ; and that these villi contain also very fine ramifications of blood, vessels. They have also agreed, that the surface of the intestines in the intervals of the villi seems occupied with the orifices of duets or of exhalent vessels.* Division of the Intestines . Although there is a considerable degree of uni- formity in the structure of the intestinal canal, dif- ferent parts of it are very distinguishable from each other by their exterior appearance, by their size, their investments, and their position. The first division is into two great portions, which are very different from each other in their diameter and length, as well as their situation : the first por- tion being much smaller in diameter, and near four times the length of the other. * On this subject the student will consult with advantage, Hewson’s Experimental Inquiries, vol. 2 ; Sheldon’s History of the Absorbent System, part 1st ; Cruikshank on the Anatomy of the Absorbing Ves- sels ; and the Historia Yasorum Lymphaticorum Corporis Hvtmani, of Mascagni. 110 Division of the Intestines. These portions are therefore known by the names of Great, and Small Intestines , and the line of sepa- ration between them is very strongly marked ; for they do not gradually change into each other, but the al- teration in size and in exterior appearance is very abrupt, and their communication is not perfectly di- rect. A considerable portion of the Great Intestine is fixed immoveably in the abdomen, while a large part of the Small Intestine is very moveable. Each of these great portions of the intestinal tube is subdivided into three parts. Thus, in the Small Intestine , there is a piece at the commencement call- ed Duodenum , a great part of which has no coat from the peritoneum, and is immoveably fixed in one situa- tion ; while all the remainder of the small intestine has a uniform covering from the peritoneum, and is very moveable. This last piece, notwithstanding its exterior uniformity, is considered as forming two parts. The uppermost two-fifths form one part* which is called Jejunum ; and the remainder is called Ileum. The Great Intestine commences in the lower part of the right side of the abdomen, and after proceeding up that side crosses over to the left, along which it descends to the lower part again, when by a peculiar flexure it proceeds to the cen- tre of the posterior margin of the pelvis, from which it passes down to the anus. A short portion of this intestine, which is above its junction with the ileum, is called Cxcum ; the part which proceeds from this, round the abdomen, is called Colon ; and the por- tion which is in the pelvis is called Rectum. Of the Small Intestines. Previous to the description of the small intestines, it is necessary to observe, that the Mesocolon, or process of the peritoneum connected to the trans- verse portion of the colon, forms a kind of moveable Commencement of the Small Intestines . til and incomplete septum, which divides the abdomen into an upper and lower apartment. Above this sep- tum are the stomach, with the commencement of the duodenum, the liver, and the spleen ; below it. that portion of the small intestine which is called jejunum and ileum, makes its appearance. The portion of the intestine which passes from the stomach to the jejunum, and is called Duodenum , is so much in- volved by the mesocolon, that the greatest part of it cannot be seen without dissecting the mesocolon from its connexion with the back of the abdomen. — For the duodenum proceeds backwards from the pylorus, and passing down behind the peritoneum, enters a vacant space between the two lamina of the mesoco- lon ; it proceeds for some distance in this space, and then emerges on the lower side of the mesocolon. Here the duodenum terminates, and the small intes- tine then is invested by the peritoneum in such man- ner as to form the mesentery, which continues with it throughout its whole course to the great intestine. This portion of the intestine, although very uniform in its exterior appearance, as has been observed be- fore, is divided into Jejunum and Ileum : the jeju- num being the upper’ portion, which begins at the mesocolon ; and the ileum the lower portion, which opens into the great intestine. '> Of the Duodenum. The length of this intestine is equal to the breadth of twelve fingers, and hence its name. It is very different from the rest of the small intestine, not only as respects its position and investment by the perito- neum, but on account of its connexion with the liver and pancreas, by means of their excretory ducts, which open into it. From this connexion with these glands, probably, all the peculiarities of its position are to be deduced. 112 Situation of the Duodenum. When the stomach is in its natural situation, the pylorus is at some distance from the back of the abdomen. The duodenum proceeds backwards from this point, and passes near the neck of the gall-bladder, being here connected with the small omentum ; it then curves downwards, and descends before the right kidney, sometimes as low as the lower part of it; then it curves again, and passes over to the left: after it has arrived at the left side of the spine, at the second or third lumbar vertebra, it projects forwards and downwards to form the je- junum. The only portion of this intestine which is moveable, is that which is in sight as it proceeds immediately from the pylorus, being about an inch and a half or two inches in length. The remain- der is connected to the back of the abdomen, and lies between the two lamina of the mesocolon. In its progress its passes before the aorta and the vena cava, but the principal branch of the vena portarum is be- fore it. The duodenum is larger in diameter than any other part of the small intestines, and has a stronger mus- cular coat. Its general situation admits of great dilata- tion, and it has been called a second stomach. Its in- ternal coat is strictly villous, in the anatomical sense of the word ; and its folds, the valvulse conniventes, begin at a small distance from the pylorus. The ori- fices of many mucous ducts are to be seen on its sur- face. It is supposed that some of these are the ter- minations of ducts from the glands of Brunner, which sometimes appear in the villous coat, or very close to it exteriorly ; being small flat bodies, with a depres- sion in the centre, and a foramen in the depression. They are sometimes very numerous at the upper ex- tremity of this intestine, and diminish gradually to- wards the other extremity. The biliary and pancreatic ducts open posteriorly 113 Jejunum and Ileum . into the duodenum, rather above the middle of it. The orifice of these ducts is generally surrounded by a small tubercle, which is oblong, somewhat rounded at one extremity, and pointed at the other. Some- times this orifice is in a plait, like one of the valvulse conniventes. Most commonly the two ducts unite before they perforate the coat, so as to form but one orifice ; and sometimes they open separately, but al- ways very near to each other. Absorbent vessels, which contain chyle, are found on the duodenum. The Jejunum and Ileum Are situated in the abdomen very differently from the duodenum. When the cavity is opened, and the omentum raised, they are in full view ; and every portion of them, except the two extremities and the parts near them, can readily be moved. This free- dom of motion is owing to the manner in which they are invested by the peritoneum ; or in the technical language of anatomy, to the length of their mesen- tery. They agree in their structure with the general description of the small intestines, but their muscular coat is rather weaker than that of the duodenum. The valvulze conniventes are very numerous and large in the upper part of the tube, or the jejunum ; and gra- dually diminish in number, until they finally disap- pear, in the lower part of the ileum. The villous coat is in perfection in the jejunum, the villi being more conspicuous there than in any other part of the intestinal tube. There are frequently found, exterior to this coat, but intimately connected with it, many small glandular bodies of a roundish form, which are often clustered together at that part of the intestine which corresponds with the interstice of the lamina of the mesentery. They are called Peyer’s glands, Vol. ii, 15 114 Distinction between Jejunum and Ileum after the anatomists who first described them ; and are supposed, like the glands of Brunner, to secrete mu- cus. If a portion of the jejunum be inverted, and moderately distended with air, these bodies appear very distinctly in it, dispersed at small distances from each other. In the ileum they appear in small clus- ters, which often have the appearance of disease. No natural line of separation for distinguishing the jejunum and ileum from each other, is to be found; but these names are still retained ; and therefore a rule laid down by Winslow is generally adopted, viz. to name the first two-fifths of the tube jejunum, and the remainder ileum. There are, however, some important differences between these portions of the intestine. In the jejunum, the valvulae conniventes are so nu- merous, that they lie in contact with each other, as shingles on the roof of a house ; in the ileum they gradually diminish in number, and finally disappear. In the jejunum the villi are much stronger than they are in the ileum. It is very difficult to acquire a precise idea of the arrangement of this part of the intestinal tube, while it is in the abdomen, especially if it be much distend- ed ; but if it be separated at each extremity from the intestine with which it is connected, and the mesen- tery cut off from the back of the abdomen, and the whole then spread out upon a flat surface, it will appear, as has been already said, that the intestine is arranged so as to form a semicircle or large curve; the concavity of which is opposite to the back of the abdomen, while the convexity presents forward. It will also appear, when thus placed upon a table, that the intestine, while connected with the mesentery, is laid into many folds. It has been supposed, that the middle portion of the mesen- tery, and the intestine connected with it, is generally Construction of 'the Mesentery . 115 itn the umbilical region ; and the two portions on the sides of it are in the iliac regions ; but their situation in the abdomen varies considerably at different times. When the viscera of the pejvis are empty, a large portion of the small intestine is in the pelvis; but when those viscera are filled, the intestine is in the general cavity of the abdomen. The Mesentery Is a process of the peritoneum, which is formed in the manner of a plait or fold, and of course consists of two lamina. These lamina proceed from the back part of the abdomen, and are so near to each other, that they compose one substantial process ; having cellular and adipose substance, blood vessels and nerves, with absorbent or lacteal vessels and their glands, between them. The form of this process, when it is separated from the back, and the intestines are detached from it, is somewhat semicircular : that portion of its mar- gin or edge which corresponds to the diameter of the semicircle, is connected to the back of the abdomen, and called the root of the mesentery; the edge, which is the circumference of the semicircle, is connected with the intestine. The edge connected with the back of the abdomen is commonly about five or six inches in length -. the semicircular edge, instead of ex- tending fifteen or eighteen inches, the ordinary pro- portion, is attached to a portion of intestine some- times twenty-four feet in length. The mesentery on account of this great difference between its diameter and circumference, has been compared to the ruffle of a shirtsleeve; its roots being taken for the plaited edge of the ruffle, and the circumference for its loose edge. But the comparison is not precisely accurate; for the mesentery is not plaited at its root, but perfectly 116 Root of the Mesentery . smooth, and free from every kind of fold. It begins to enlarge towards its circumference, and enlarges to that degree, that it falls into plaits or folds : precisely such as would exist in a semicircular piece of mem- brane about six inches in diameter, if a number of simple incisions, of about an inch and a half in length, were made in a radiated direction from its circumfer- ence, and if portions like a sextant or quadrant were taken from a circular membrane three inches in di- ameter, and united by their edges to these incisions, so that their circumference might be continuous with the circumference of the large semicircular piece. In this case, the portions like quadrants or sextants would assume a folded position like the edge of the mesentery, while the middle of the semicircular piece would preserve its regular form without folds ; as is the case with the mesentery at some distance within its circumference. By many additions of this kind, the circumference of a membrane, which was original- ly a semicircle of five or six inches, may be extended so as to exceed greatly that of the mesentery. It seems of course impossible to form an accurate mo- del of the mesentery with a single piece of membrane or paper ; but it may be easily made with clay, or any ductile substance. A model of this kind must necessarily be folded after the manner of the mesen- tery ; and its circumference, like the mesentery, would appear as if formed of portions of the circumference of smaller circles united to each other.* The root of the mesentery commences with the jejunum on the lower side of the mesocolon, at the left of the spine, and extends downwards near to the right iliac region $ crossing the spine obliquely. * A model, upon the plan first mentioned, was invented by Dr. J. G. Shippen. It has been proposed, I believe by M. Gavard, to make one with a single piece of buckskin, of a semicircular form f by stretching it at the circumference! Appearances of the C cecum and Coton. 117 When it is examined in its natural situation, the peri- toneum is found continued from the back of the ab- domen to the intestine ; it then surrounds the intes- tine, and continues from it to the back of the abdo- men again. There must therefore be two lamina of peritoneum in the mesentery, and there must be a small portion of intestine answering to the interstice between these lamina, which is not covered by the peritoneum. The blood vessels, and absorbent or lacteals, pass most commodiously to the intestines between these lamina ; for they are connected with large trunks that lie on or near the spine, and the root of the mesentery commences there. The glands connected with the lacteals or absorb- ents are very conspicuous in the mesentery, and are commonly called mesenteric glands. They are of different sizes, from more than half an inch to one or two lines in diameter. They are very numerous, and scattered irregularly, but are seldom observed very near to the intestine. They are often enlarged in consequence of disease, especially in children. The nerves of the small intestines which are de- rived principally from the superior mesenteric plex- us, are also to be found here. The adipose matter between the lamina of the me- sentery is very often in a large quantity, but varies, in proportion to the general quantity of adeps in the subject. OF THE GREAT INTESTINES. The Ceecum and Colon Are very different from the small intestines in many respects. They are much larger in diameter. Their external surface is marked by three longitudi- nal bands of a light colour, which extend the great- est part of their length, and are placed nearly at 118 Position of the Caecum, equal distances from each other. The spaces be- tween these bands are marked by transverse inden- tations, which pass from one band to the other at short but unequal distances. At these indentations the coats of the intestine are pressed inwards, as if a fine thread had been drawn round it externally? while the spaces between them are full and tumid? and on this account are called cells. The great intestine, with these appearances? be- gins, as has been already observed, in the right iliac region, by a rounded end which rests on the fossa or concave surface formed by the costa of the ileum ; from this it is continued upwards in the right lum- bar region, anterior to the kidney, until it arrives near the liver, when it forms a curve? and passes directly across the abdomen to the left side. In this course it approaches so near to the under side of the liver, that it is often in contact with it, and with the gall-bladder, which, after death, tinges it with a yellow colour. On the left side it passes dowm the lumbar region, before the kidney, to the left iliac region ; here it is curved so as to resemble the Roman letter S, inverted ; this curve generally car- ries it to the right side of the spine, and then brings it back to the centre of the sacrum. Here the in- testine changes its course, and passing into the pelvis, continues downward, in contact with the sacrum and coccygis, and partaking of the curvature of those bones, until it terminates at the anus, where it is connected with the sphincter and levator ani muscles. About two inches from the commencement of the great intestine the ileum opens into it laterally ; and all that portion which is between its commencement and the insertion of the ileum is termed. Caecum, or the blind intestine: that part of the great tube? which is included in its course from the insertion of 119 Structure of the Colon . the ileum to the posterior part of the brim of the pelvis, is called Colon ; and the remainder, or the part which is contained in the pelvis, is termed i?ee- turn. The Csecurn is nearly as wide as it is long ; it is fixed in the right iliac fossa by the peritoneum, which invests it so that the great body of the intes- tine projects from the surface of the fossa covered by the peritoneum ; but a portion is in close con- tact with the surface, and connected to it by celhi" lar membrane. Its external surface, covered by the peritoneum, is marked by two of the bands or stripes before mentioned, which proceed on it length- ways. These bands are in full view, but the third band is generally on that part of the intestine which rests on the iliac fossa, and is therefore out of sight. At the rounded extremity of the csecurn, situated anteriorly and internally, is a small process resem- bling an earth-worm in form and size : this is there- fore called Jippendicula Vermiformis. It is hollow and communicates with the cavity of the csecurn at the place of junction ; and like the csecurn, has its other extremity closed up. It is composed of the same number of coats and has the same structure as the great intestine : its length varies from two to four inches. The longitudinal bands above mentioned commence at the junction of this appendix with the csecurn, and continue throughout the extent of the colon. They appear to be formed by some of the longitudinal fibres of the muscular coat, which are arranged close to each other. These fibres seem to be shorter than the coats of the intestine, and the interior coats adhere firmly to them. Thus are produced the indentations and cells ; for if the bands are divided transversely, the indentations disappear, and the surface of the intes- 120 Valve of the Colon. tine becomes uniform. One of these bands is cover* ed by the mesocolon. The circular or transverse fibres of the muscular coat of the csecum and colon are very delicate, and not numerous. The internal coat differs materially from that of the small intestines, although at first view they seem to resemble each other : for if a portion of the ileum and of the colon be inverted and suspended in water, no villi can be seen with the naked eye on the internal coat of the colon, while those of the ileum are very visible. The glands exterior to this coat are larger than those on the small intestines. Instead of valvulse conniventes, are the ridges made by the indentations or depressions above described, which separate the incomplete cells from each other. These ridges differ essentially from the valvular con- niventes, because all the coats of the intestine are con- cerned in their formation, whereas the valvulse conni- ventes are formed by the villous coat only ; they also project into the cavity of the intestine, while the val- vulse are laid on its surface. They pass only from one longitudinal band to another, and, in consequence of this, the cells are small, and the position of each band is very evident when the intestine is laid open. The communication of the ileum with the great intestine has been already stated to be on the left side of it, about two inches from its commencement. The aperture is so constructed, that it is considered as a valve, and is called the Valve of Bauhin, or of Tulpius, after the anatomists who have described it.* The appearance of the aperture is as follows: If the csecum, with a small portion of the ileum and * Posthius in 1566; Vidus Vidius about 1569; Alberti in 1581, and Varo- lius who died in 1575, each lay claims to the discovery of it. Bauhin’s claims are in 1579. Em Structure of the Valve of the Colon. • 121 of the colon, be separated from the other intestines, and kept in an inflated state until it be so dry as to preserve its form when opened, and then if the cae- cum and colon be laid open opposite to the aperture of the ileum, a large transverse ridge, resembling some of the ridges or folds just described, will be seen projecting into the cavity of the intestine. In the internal edge of this fold is a long slit or opening, which forms the communication between the two in- testines. It is obvious that the form of this fold must be that of a crescent ; and that its two surfaces with the slit between them, must have the appearance of two lips, which would readily permit a fluid or sub- stance of soft consistence to pass from the ileum into the great intestines, but must impede, if not prevent, its passage back ; especially if the large intestines were distended, as then the lips would be pressed against each other. When the peritoneal coat is dissected from each of the intestines at their place of junction, and this structure is then examined from without , it appears as if a transverse or half circular indentation had been formed by the villous coat of the great intestine, and that the internal coat of the extremity of the ileum was pressed into this indentation, and united to the internal coat of the great intestine which form- ed it; while there was a slit, both in. the indentation and in the end of the ileum, which formed a commu- nication between the eavityof the great intestine and the ileum. The longitudinal fibres of both intestines, as well as their external coats, seemed to be united, so as to form a common cover for them ; while the circular fibres were blended in the two portions of the indentation which form the lips of the orifice. This orifice is, of course, transverse with respect to the intestine. It has been observed, that there was a difference in the thickness and strength of the VoLj ii, 16 ' 122 Position of the Rectum . two lips or valves ; that the lower valve was the strongest, and appeared to have the largest propor- tion of muscular fibres in its composition. At the extremities of the orifice, and near each end of the fold or ridge, are tendinous fibres, which give strength to the structure ; they are called the Reti- nacula of Morgagni, as they were first described by that anatomist. There is great reason for believing that this valve cannot prevent the retrograde motion of the contents of the intestines in all cases ; for in some instances of hernia and of colic, matter perfectly stercoraceous has been vomited, and the probable inference from such a state of the ejected matter is, that this matter has been in the large intestines. It is also said, that suppositories and enemata have been discharged by vomiting. On the right and left sides of the abdomen, the colon is in close contact with the posterior surface of the cavity. The peritoneum, which covers this sur- face, extends over the intestine also, and thus retains it in its position . The great arch of the colon, which is loose and moves far from the back of the abdomen, is invested by the two lamina of the omentum, which, after surrounding it, unite again and form the meso- colon. Connected with the exterior surface of the colon are many processes, composed of adipose mem- brane, varying in length from half an inch to an inch and a half : these appear, to be of the nature of the omentum, and are therefore generally denominated Appendices Epiploicsc. The Rectum. After forming the sigmoid flexure, the colon ter- minates; and the rectum begins opposite to the lower Surface of the last lumbar vertebra, and nearly in contact with it ; from this it proceeds downwards, 123 Structure of the Rectum. forming a curve like the sacrum, until it terminates at the anus, where it is invested with the muscles called the sphincter, and levator ani. It is called rectum, because in this course it is supposed not to incline to either side ; but it is often found on one side of the middle line.* This intestine being in contact with the posterior surface of the pelvis, is covered, on its anterior sur- face only, by the peritoneum which lines the poste- rior surface of the pelvis ; and it is fixed in this situ- ation by the peritoneum, as the colon is on the right and left sides of the abdomen, but more loosely; and therefore the term Mesorectum has sometimes been applied to that portion of the peritoneum which is analogous to the mesentery and. mesocolon. The pe- ritoneum does not extend to the end of th'e rectum ; for it is reflected at the lower part of the pelvis from the rectum to the bladder, or uterus, and does not line the bottom of the pelvis ; so that the lower part of this intestine, as well as of the other viscera of the pelvis, is below the peritoneum, and not connect- ed with it. The muscular coat of the rectum is much thicker and stronger than that of any other intestine. The strata of longitudinal and circular fibres which com- pose it are very distinct from each other. The lon- gitudinal fibres are most numerous, and terminate at the insertion of the fibres of the levator ani muscle. The lower circular fibres are intimately connected with the sphincter ani. The internal coat is very vascular, but the villous structure is not apparent. Mucous follicles are also very numerous; and there are likewise some distinct glandular bodies exterior to this coat, which vary in size in different subjects. * Morgagni and Haller supposed it to be commonly on the left of the middle line: and Sabatier on the right. 124 Absorbents and JYerves of the Intestines. The quantity of mucous discharged from the rec- tum in certain cases of disease, is sometimes very great. The internal coat, in consequence of the con- traction of the circular fibres exterior to it, some- times forms longitudinal folds, which have been called its columns ; these often disappear when the intestine is opened lengthways and spread out. By the contraction of the longitudinal fibres, the internal coat is often thrown into folds or doublings, that must assume a transverse or circular direction ; they occasionally pass down through the sphincter, and form the prolapsus ani. The rectum is most plenti- fully supplied with blood vessels, to be described hereafter; and it may be observed, that, on the low- er part of the internal coat, the veins are particular- ly numerous. The internal coat of the rectum terminates ab- ruptly just within the anus, and is united to a pro- duction of the skin, which, like the covering of the lips, is very delicate and vascular, and has an epi- thelium, or very thin cuticle* spread over it. The levator and sphincter ani muscles, with which the termination of the rectum is invested, are described in the first volume. , The Absorbents of the Intestines are commonly denominated Lacteals .* They originate on the in- ternal surfaces of these viscera, as has been already described. After passing through the lymphatic glands, which are so numerous on the mesentery, they generally unite and form one of the great trunks which compose the thoracic duct. It is asserted, that some of the absorbent vessels of the lower in- testines unite to the lymphatics of the loins. * The lacteals were first observed by Erasistratus and Herophilus, of the school of Alexandria, during the reign of the Plotemies ; and subse- quently by Aselli, of Pavia, in 1622, the knowledge of them having been lost for 1900 years. — E d. Origin and Arrangement of the Omentum. 125 The Nerves of the Intestines are principally de- rived from the intercostals, or great sympathetics From each of these nerves, while they are in the tho- rax, an important branch, called the ramus splanch- nieus, arises. These splanchnic branches pass through the diaphragm, and are the chief contributors to the ganglions and plexuses formed in the abdomen. A plexus, derived from this source surrounds the supe- rior mesenteric artery, and another the inferior me- senteric ; and from these proceeds the nerves of the intestines. The Omentum Requires a separate description, although several circumstances connected with its structure have been already noticed. It often varies in its position; but when it is rendered firm by a quantity of adipose mat- ter, it is spread over the intestines like an apron, ex- tending from the lower edge, or great curvature of the stomach, towards the bottom of the abdomen. As has been already said, it is an extension of the peritoneum, in two lamina, from the concave surface of the liver to the lesser curvature of the stomach ; and these lamina, after surrounding the stomach, come in contact with each other near its great curva- ture. From this portion of the stomach, from the commencement of the duodenum, and also from the spleen, the Omentum , composed of two lamina de- scends over the colon and the small intestines more or less low into the abdomen ; it is then folded back- wards and upwards, and is continued. until it meets the great arch in the colon : here the lamina again separate and enclose that portion of the intestine, on the posterior side of which they again approach each other, and form a membrane like the mesentery, of two lamina, which passes from the concave or pos- terior surface of the colon to the back of the abdo- 126 Origin and Arrangement of the Omentum . men, where it is continued into the membrane which lines that surface. This last portion is the Mesoco- lon: the portion between the liver and stomach is called the Omentum of Winslow , or the lesser omentum; and the great portion between the sto- mach and colon is called the Great Omentum , or the omentum gastro colicum. There is also a process of peritoneum continued from that portion of the colon which is on the right side of the abdomen, and from the caecum, which extends to some distance ; it is formed of two lamina, that compose a cavity of an angular form. This has been called the Omen- tum Colicum. The great and small omentum, with a portion of the peritoneum on the back of the abdomen, form a sac, which encloses a distinct cavity in the abdomen. The anterior part of this sac is composed of two lamina, and between these lamina are the stomach and the great arch of the colon. This cavity, form- ed by the two omenta, communicates with the gene- ral cavity of the abdomen by a foramen of a semicir- cular form, called the Foramen of Winslow, which is behind the great cord of the vessels that go to the liver. The omentum is so delicate in structure, that when free from fat, it is very liable to laceration merely by adhering to the fingers, if they are dry. Winslow therefore advised that some unctuous sub- stance should be rubbed on the hands, before they were applied to it. The appearance of the great omentum is very dif- ferent in different persons. In the emaciated, it appears like a delicate transparent membrane ; in the corpulent, it is like a broad mass of adeps, which sometimes is very thick. When it is thus loaded with adeps, it is most commonly spread over the small intestines : when it is free from fat, it is often Varieties in the Appearance of the Omentum. 127 compressed together, so as to form a small mass near the arch of the colon, on the left side. The principal blood vessels of the omentum are derived from those of the stomach, and are called gas- tro epiploic arteries and veins. The use of this membrane in the animal economy has not been ascertained with certainty. It seems probable that one of its principal objects is to protect the small intestines, and lessen the friction consequent upon their motion ; but it has been supposed to an- swer several other important purposes.* * See Halleri Elementa Physiologic, vol. vi. page 381. Gavard. Traite de Splanchnologie, page 350. Dr. James Rush’s Inquiry into the use of the Omentum. CHAPTER III. OF THE LIVER, THE PANCREAS, AND THE SPLEEN. SECTION L Of the Liver. This largest viscus of the abdomen, when in a healthy condition, is of a reddish-brown colour. If it is taken out of the subject, and laid on a flat surface, it is flat, but in the abdomen it is convex and concave. It is situated in the right hypochondriac region, which it occupies entirely ; and extends through the upper portion of the epigastric into the left hypochon- driac region. Being placed immediately under the diaphragm, and in close contact with it, as well as with the inner surface of the right hypochondriac region, it partakes of their form, and is convex above and concave below. When thus situated, it is of an irre- gular figure, between the circular and the oval, but it is broader at the right extremity than at the left, and very irregular in thickness. The edge or margin which is in contact with the posterior part of the right hypochondriac region, is very thick. It gradually be- comes thinner towards the left, and also towards the front; so that the right margin, and a large portion of the posterior margin, is very thick, while the left and the anterior margin is thin. The upper convex surface of the liver, when in its natural situation, is smooth : the lower concave surface is marked by several grooves or fissures and eminences. One of these, called the Umbilical or the great fissure, commences at a notch in the ante- 129 Lobes and Fissures of the Liver. rior edge of the liver, to the left of the middle, and continues to the posterior edge. At the commence- ment of this fissure the umbilical ligament enters ; and at the termination, or near it, the vena cava is situated. Opposite to this fissure, on the upper or convex surface, is a ligament passing from the dia- phragm to the liver, which is called the falciform. The fissure and the ligament divide the liver into its two great lobes, the Right and Left . Another great fissure , called the transverse or principal, commences in the right lobe and extends to the left, crossing the first mentioned fissure at right angles, and extending a very short distance beyond it. It is very deep, and rather nearer to the posterior than the anterior edge of the liver. In this fissure, near to its right extremity, the great vein, called vena portarum, and the hepatic artery enter, and the excretory duct of the liver, commonly called the hepatic duct, comes out. About the mid- dle of the fissure are two prominences, one on each side ; these were called the portae, or gates of the liver, and hence the great vein was called vena por- tarum. This vein has two very large rectangular branches, which constitute what is called the sinus of the vena portarum, and they occupy the principal extent of the fissure. The liver is in close contact with the vena cava behind; and there is either a groove in it for the pas- sage of the vein, or this great vessel is completely en- closed by it. There is also an excavation on the lower surface of the liver, which is occupied by a portion of the gall bladder. Besides the great lobes above mentioned, there are also two or three prominent parts on the concave surface, which are denominated lobes. One of these, called Lohulus Spigelii, is oblong, with two sides, and an angle continued along its whole length, Vol, II. 17 130 Ligaments of the Liver „ which extends from the transverse fissure to the pos- terior margin of the liver. It is situated between the posterior part of the transverse fissure, or ductus venosus, and the vena cava. The anterior extremity of this lobe, which forms one of the margins of the transverse fissure, is some- what bifurcated, and has been called lobulus caudatus. The largest portion of the bifurcated end forms a pro- cess like a papilla, and is one of the portae. Between the umbilical fissure and the depression for the gall bladder is a protuberant space, which varies from an inch and a quarter to two inches in breadth. This has also been called a lobe, Lobulus Quarlus or Anonymous , its posterior point opposite the papilla of the lobulus spigelii forms the other porta of the liver. The peritoneum is extended from the surface of the abdomen to the surface of the liver, in such man- ner as to cover it, and to form ligaments, which have a great effect in retaining it in its proper situation. The whole posterior edge of the liver is in contact with the back of the abdomen. The peritoneum above the liver is reflected to the upper surface of it, and the peritoneum below it to the lower surface ; so that two lamina of the peritoneum pass from the lower part of the diaphragm at the back of the abdo- men to the posterior edge of the liver. These pro- cesses of the peritoneum are considered as forming two ligaments, which are called the right and left lateral ligaments. A portion of the posterior surface of the liver, uncovered by the peritoneum, is often in contact with a portion of the tendon of the diaphragm, also uncovered by peritoneum : around this place of contact, the peritoneum is extended from the dia- phragm to the liver, and thus forms what has been called the coronary ligament of the liver. The peritoneum of the right side of the diaphragm, and of the abdominal muscles, as far down as the 131 Mode of supporting the Liver, umbilicus, is extended to the liver, and joins it on the convex surface immediately opposite to the um- bilical fissure. The peritoneum from the left side of these parts does the same j and as these reflections ©f the peritoneum are continued from so low a part as the umbilicus, they are extended not only to the convex surface of the liver, but also to the great notch, and along the umbilical fissure. From the umbilicus proceeds a round cord-like lig- ament, which in the foetal state was a vein, that passes to the great fissure of the liver, and along it. The process of the peritoneum above mentioned is so connected with this cord, that it encloses it in its lower edge, and the whole is called the falciform ligament of the liver. The cord, when named se- parately, is the umbilical or the round ligament : and the membrane or lamina of the peritoneum forms the suspensory ligament. Besides these, the peritoneum on the lower side of the liver is so arranged, that it not only extends to the stomach, but to the duodenum and the colon. By these ligaments the position of the liver must be fixed to a great degree ; and there is one additional connexion, which must have a great effect in retain- ingit in its proper situation. The vena cava receives two or three great veins from the liver, at the place where it is in contact with the posterior edge of that viscus : these veins of course pass directly from the substance of the liver into the cava, and connect if to that vessel. As the cava is supported by the heart, and also by the diaphragm, it must afford a consider- able support to the liver. When the stomach and intestines are distended, they must also contribute in a considerable degree to the support of the liver. The liver has a strong tendency, when we are erect, to change its situation % and some considerable 132 Acini of the Liver. — Proper Coat of the Liver , support is necessary to counteract this tendency. It would move to the right, when we lie on the right side, if it were not in contact with the ribs ; and it inclines to the left, for want of such support, when we lie on the left side. It has been computed, that the liver descends about two inches, when the position of the subject is changed from the horizontal to the erect. As it is in contact with the diaphragm, it is obvious that it must be influenced by the motions of that muscle, and that it must descend when the diaphragm con- tracts. The liver is composed of a substance which has some firmness of consistence, although it is yielding; and is also somewhat brittle or friable.* When cut into, the sections of many tubes, or vessels of dif- ferent diameters, appear on the cut surface. When the texture of this substance is more closely ex- amined, it appears somewhat granulated, or com- posed of very small bodies, which were called acini by the anatomists who first described them. The whole substance is enclosed by the peritoneum, which is extended to it from the surface of the abdomen in the manner that has been already described. It has also a proper coat or capsule ; and on the posterior edge, where the lamina of the lateral ligaments pass from the diaphragm to the liver, at some distance from each other, a portion of the liver, covered by this coat and by cellular substance, is in contact with the diaphragm. The same thing occurs likewise at the coronary ligament, f The liver holds the first place among the glands * It has been fractured in the living body by external violence. f Many anatomists deny the existence of this coat ; but if one of the lamina of the ligaments be carefully peeled off from the surface of a liver which is slightly affected by putrefaction, it will be apparent, al- though very thin. It was described by M. Laennec, in Le Journal de Medecine for 1803. Vessels of the Liver.-— Hepatic Artery „ 133 of the body for size, but it is still more remarkable for some other circumstances in its economy. In addition to an artery, which passes to it as arteries do to other glands, there is a large vein which also enters it as an artery; and after ramifying through- out the liver, communicates, as does the artery, with other veins, which carry the blood from this gland into the vena cava and the general circulation. There are therefore three species of blood vessels in the liver; and with these are found the vessels which carry out of the gland the fluid secreted by it, or the bile. The artery of the liver is denominated the Hepa~ tic Artery. The vein which goes to the liver is call- ed the Vena Portarum , from the place at which it enters. The veins which carry to the vena cava the blood brought to the liver by the hepatic artery and the vena portarum, are called the Hepatic Veins ; and the duct through which the bile flows out of the liver, is called the Hepatic Duct. Three of these vessels, the Hepatic Artery , the Vena Portarum , and the Hepatic Duct , enter the liver at the great fissure, at the spot where the prominences exist call- ed the porta 2 ; hence the name vena portarum was applied to the vein. These vessels ramify in the manner presently to be described; and it is ascertained by minute anatomical investigation, that the liver is entirely composed of the ramifications of these vessels and of the hepatic veins, with absorbent vessels and nerves, which are connect- ed together by cellular membrane. It has been already observed, that the first great branch sent off by the aorta in the abdomen, the Cce- liac, divides into three branches, which go respective- ly to the stomach, the liver, and the spleen. The Hepatic is generally the largest of these branches. In its progress towards the liver it sends 134 Vena P or forum - off an artery to the stomach, called the gastrica dex- tra. At the great fissure it divides into two branches"; the right branch, which supplies the right lobe of the liver, is of course the largest. Tins branch sends off oat to the gall-bladder, which is called the cystic ar- tery ; and also some smaller branches : it passes un- der the hepatic duct, and ramifies through the great lobe of the liver. The left branch is distributed through the left lobe of the viseus. It can be proved by injection, that the hepatic artery communicates not only with the hepatic veins, but with the biliary duct, and the vena portarum also. It has been disputed whether the size of this artery is greater than would he requisite for the nourishment and animation of the liver- The Vena Portarum , the great peculiarity of the liver, originates from all the chylopoietic viscera ex- cept the liver, and is of course formed by the union of the veins which correspond to all the branches of the coeliae and mesenteric arteries, as they are dis- tributed to the stomach and intestines, the spleen, the pancreas, and the omentum. The veins from the intestines generally form two great trunks, which are denominated the greater and lesser mesenteric veins. The great mesenteric vein is situated to the right, and rather before the mesenteric artery.— After it has approached the origin of the artery it separates from it, and passes behind the pancreas i at this place, nearly in front of the spine, it is joined by the great vein of the spleen, which forms almost a right angle with it, and these constitute the great trunk of the vena portarum- The lesser mesenteric vein, which corresponds to the inferior mesenteric artery, and brings blood from the pelvis and from the left part of the colon, becomes finally a large vessel, and commonly unites with the splenic about an inch and a half before its junction with the supe- 135 Vena For iarum . —Hepatic Duct . sior mesenteric vein. The vena portarum, thus formed, proceeds towards the liver, inclining to the right, and is generally about three inches in length: in its course it sometimes receives small veins, which in other cases pass to its splenic and mesenteric branches. When it has arrived at the great trans- verse sinus of the liver, it divides into two large branches, each of which forms nearly a right angle with it. Their size is so great, that, when distend- ed with injection, they appear like an independent vessel, into which the vena portarum enters ; and on this account they are called the great Sinus of the vena portarum. They do not adhere firmly to the glandular substance of the liver, but are united to it by cellular membrane. The right branch is the widest and shortest. It generally divides into three branches; an anterior, a posterior, and a lateral branch; which ramify minutely, and extend them- selves in the right lobe. The left branch is much longer, and continues to the extent of the transverse fissure. Near its termination it is joined by the urn- bilical ligament, which has been already mentioned. This branch is generally in contact with a branch of the hepatic artery and of the hepatic duct ; and ra- mifies, like the right branch, into the contiguous parts of the liver. The Hepatic or excretory duct originates, by very small vessels, from the acini or corpuscles of which the liver is composed, and into which the minute ra- mifications of the vena portarum and hepatic artery extend. They accompany these vessels, increasing as they increase, although the fluid they contain moves in an opposite direction ; and two large branches which they ultimately form are situated at the portae of the liver, in contact with the great branches of the vena portarum and the hepatic artery. These three vessels are in contact with each other 136 Hepatic Veins. — Nerves of the Liver . before they enter the liver. The biliary duct is an-^ terior, the vena portarmn posterior, and the artery to the left of them. They are accompanied by nerves and lymphatic vessels, and are surrounded by a con-\ siderable quantity of cellular substance, and thus ar- ranged are partially covered with peritoneum. The cellular substance which invests them continues with them into the liver, and is more particularly con- nected with the vena portarum. It is called Glis- son’s Capsule, and was supposed to have some con- tractile power, which assisted the circulation of the vena portarum ; but that idea is now altogether aban- doned, The hepatic veins, which receive the blood of the hepatic artery and the vena portarum, open into the anterior part of the vena cava, where it is in contact with the liver. Generally there are three of these veins, but sometimes there are only two ; in which case one of them is formed by two others, which unite immediately before they open into the vena cava. It is to be observed, that the various branches of these veins do not accompany those branches of the vena portarum or hepatic artery to which they correspond, but form very large angles with them. This is probably owing merely to their termination in a part so distant from that in which the artery and the vena portarum originate ; but it is very different from what occurs in other glands. The Nerves of the Liver are derived from the sisnilunar ganglions of the splanchnic nerves. From these many nerves proceed, which form a net-work denominated the solar plexus. From this plexus many threads are sent off, which form a net-work that is divided into the right and left hepatic plexus. These plexuses surround the hepatic artery and the vena portarum, and accompany them in their ramifi- cations throughout the liver, being enclosed by Glis- Lymphatics . 137 son's capsule. They receive some threads from the stomachic plexus, formed by the par vagum. A1- though the number of nervous fibres is very conside- rable, their bulk, compared with that of the liver, is very small. The Lymphatics of the Liver are extremely nu- merous; and those in that portion of the peritoneum which invests the liver may easily be rendered con- spicuous : for by pressure the injected fluid can be forced from the trunks and large branches into the smtll ramifications, in opposition to the valves. When all the surface is injected in this manner, it has the colour of the substance injected; as is the case with parts which are very vascular, when the blood ves- sels are injected. The deep-seated lymphatics are also very numer- ous in the liver, and communicate freely with the su- perficial. The superficial lymphatics which are on the upper surface, proceed through the diaphragm into the tho- rax in their course to the thoracic duct. Those which are deep-seated emerge from the liver at the portae, where the great vessels enter, and unite with the tho- racic duct in the abdomen, after passing through se- veral glands. The lymphatics of the lower surface unite with the deep-seated. The glandular or parenchymatous substance of the liver is of a reddish-brown colour, and moderately firm consistence. When it is cut into, the cut surface exhibits the sections of the branches of the different blood vessels above mentioned, and of the excretory ducts. These vessels are often distinguishable from each other. The section of the biliary duct appears the thickest ; that of the artery next ; the vena por- tarum is next in order ; and, last of all, the vense hepatic^o VOL. n. 18 138 Glandular Substance of the Liver. The branches of the vena portarum are surrounded by cellular substance, or Glisson’s capsule ; and there- fore adhere less to the substance of the liver than the branches of the hepatic veins. The sections of the hepatic ducts have often bile in them, and are there- fore termed pori biliarii. The branches of the arte- ry are also very distinguishable. When the internal substance of the liver is brought into view, and examined accurately, it appears to be formed of small bodies, or acini, which are distin- guishable from each other. If the liver happens to be torn or lacerated, the lacerated surfaces are rough and irregular, owing to the separation of these acini from each other. It is asserted by several microscopical observers, that a minute branch of each of the aforesaid vessels can be traced into each of the aeini. It is also de- clared, that if each of these vessels be injected sepa- rately with mercury, oil of turpentine coloured, or a saturated aqueous solution of gutta gamba, there is no part of the glandular mass as large as a grain of mus- tard seed in which these vessels will not be found. Several anatomists of the first character have like- wise declared, that a fluid properly injected into one of these vessels, will occasionally pass into all of them. Thus an injection will not only pass from the vena portarum to the biliary duct, but to the hepatic artery and veins also. It will likewise pass, in a re- trograde course, from the biliary ducts to the vena portarum, and to the hepatic artery and the hepatic veins ; or from any one of the four orders of vessels into the three others.* The great peculiarity of the liver is, that venous blood, instead of arterial, is brought to it for the purpose of secretion. Thus, the great vein of the * I have tried the experiment and find the assertion to be correct— E d. Peculiarity of the Liver. — Gall Bladder. 139 ehylopoietic viscera, instead of passing to the cava, enters the liver by the transverse fissure, and takes on the office of an artery ; its coats, on this account, being much thicker and stronger than those of the hepatic veins.* The Biliary or Hepatic Duct is formed of very minute vessels, which originate in the acini above described ; these unite together like veins until they form considerable branches, which finally compose the great ramifications of the biliary duct. This duct is very strong and firm, and on its internal sur- face are the orifices of many mucous follicles or ducts. It passes from the transverse fissure of the liver, with the hepatic artery, as before described, and at the distance of an inch and a half or two inches from the fissure, it unites with a duct from the gall bladder, which is called the Cystic Duct. This duct is nearly equal in length to the hepatic, and af- ter running almost parallel to it, at length unites so as to form an acute angle with it. The cystic duct is smaller than the hepatic, and they unite much like two branches of an artery. The Gall Bladder , from which the cystic duct arises, has the shape of a pear, with a very long neck, curved in a way to be hereafter described. It is situated in a superficial pit or cavity in the con- cave surface of the right lobe of the liver ; and its fundus, or basis, often projects a small distance be- yond the anterior edge of the viscus. Its position is such, that it extends from before backwards, and inclines rather to the left ; of course, therefore, when the subject lies on his back, the bottom of the blad- * A case is related by Mr. Abernethy, in the London Philosophical Transactions, in which the vena portarum terminated in the vena cava be- low the liver, without commmunicating with it. The hepatic artery was the only vessel which carried blood to the organ, and was unusually large; the liver being nearly of the natural size. Some bile was in the gall bladder, but it was less acrid tha» usual. 140 Gall Bladder. der is the uppermost part of it ; when he lies on the left side, it is also higher than the neck ; and when he lies on the right side it is the lowermost. The gall bladder consists of an internal coat, and one that is cellular or nervous, and has somewhat of a fibrous appearance. This coat connects the gall bladder to the surface of the pit nr cavity in which it lies. The peritoneal coat of the liver is extended from the surface of the viscus over that part of the surface of the gall bladder which is not in contact with it. The internal coat has a peculiar structure, with a faint resemblance to that of the villous membrane. It is so arranged as to form very fine folds, which have various directions : in some places they make a net-work ; in others, as the neck of a bladder, they are longitudinal. Many mucous follicles exist on its internal surface. The neck of the gall bladder is suddenly bent down or curved upon itself, and twisted, so that it resem- bles the neck of the swan, when the head of that bird is applied to one side of its breast. A branch of the hepatic artery, which leaves it be- fore it enters the liver, is appropriated to the gall bladder, and is therefore denominated the cystic artery. The veins corresponding to this artery empty them- selves into the vena portarum .* The lymphatic ves- sels are united to those which are found on the lower surface of the liver, and the nerves are derived from the hepatic plexus. The gall bladder appears to be merely a reservoir, into which bile passes through its duct in a retro- grade direction. If air be blown through the hepa- tic duct from the liver, it will pass to the gall blad- der almost as freely as it passes to the duodenum. * It has been justly observed by John Bell, that the veins would not ter- minate thus, if bile were secreted by the gall bladder. Ductus Communis Choledochus.— The Bite . 141 The biliary duct from the liver, after receiving the duct from the gall bladder, takes the name of Ductus Communis Choledochus. It is wider than either of the other ducts, and near three inches in length. It passes down before the vena portarum, and on the right of the hepatic artery, to the posterior surface of the right extremity of the pancreas. It passes through a small portion of that gland, and then per- forates the muscular coat of the duodenum ; after which it proceeds from half an inch to an inch be- tween this coat and the villous, and opens into the cavity of the intestine. The orifice forms a tubercle which extends lengthways of the intestine, and is rounded above and pointed below, with a slit in it. While this duct is in contact with the pancreas, a duct from that gland generally opens into it, so that the biliary and pancreatic fluids enter the duodenum by the same orifice ; but sometimes the pancreatic duct opens into the duodenum, by a distinct orifice, very near to that of the biliary duct. The Bile, or fluid secreted by the liver, appears to an- swer a two-fold purpose in the animal economy. It produces a chemical effect upon the alimentary mix- ture which passes from the stomach through the intes- tines ; and it increases the peristaltic motion of those important organs. By an inverted action of the duodenum, some of this fluid is frequently carried upwards into the stomach : it then often produces only slight derangement of the functions and sensations connected with that viscous; but sometimes violent vertigo, and even convulsions , seem to have arisen merely from the presence of a large quantity of bile in the stomach : for they have gone off completely upon the discharge of bile by vomiting. Notwithstanding these effects of bile in certain cases, in which a great deal of it exists in the sto- mach, it is often carried into the mass of blood in large quantities, and appears to be mixed with the 142 The Pancreas. serum, and to circulate through the body, without pro- ducing any very sensible effect : thus many persons who are deeply tinged by bile in their blood, experi- ence but few effects that can be imputed to the mixture of it with the circulating fluids : and neither the brain nor the heart appear to be much influenced by the cir- cumstance. Bile is miscible with water and with alcohol, and also with oily substances ; and it often assumes a green colour, when mixed with acids. The colour of the alvine discharges is derived from the bile, and they are therefore sometimes very green, when the acetous fermentation takes place in the contents of the sto- mach and bowels. It is asserted by some chemists, that ten parts in eleven of the human bile consists of water ; that albu- minous matter composes about one forty-sixth part of it ; and that there is nearly an equal quantity of re- sinous matter in it. There is also a small quantity, (one part in 244,) of uncombined soda dissolved in it, and a smaller quantity of neutral salts, consisting of soda combined with the phosphoric, sulphuric and mu- riatic acids. In addition to these is a very small quantity of phosphate of lime and of oxide of iron, and some yellow insoluble matter. The bile in the Gall Bladder is generally more viscid than that which is found in the Hepatic Duct . SECTION II. Of the Pancreas. The pancreas is a glandular body, which has a strong resemblance to the salivary glands in seve- ral particulars. It is seven inches in length, and is irregularly oblong in its form, one extremity being much larger than the other. Its large extre- mity is in contact with the duodenum, and it ex- tends from this intestine in a transverse direction to the spleen, to which it is connected by the omentum and by blood vessels. It is not invested by the peritoneum, but is situated in the space which exists The Pancreas . 143 between the two lamina of the mesocolon, as they proceed from the back of the abdomen, before they come in contact with each other. It is anterior to the aorta and vena cava, and to the mesenteric vein, or main branch of the vena portarum ; being connect- ed to these parts by cellular membrane. At the right extremity, which is connected with the duodenum, is a process of the gland that extends downwards in close contact with the intestine. — This is called the head of the pancreas, or the lesser prancreas. The position of the pancreas is such, that one of its surfaces looks forwards and rather upwards, and the other backwards and downwards ; one edge is of course posterior and superior, and the other ante- rior and inferior. The posterior of these edges is much thicker than the other, and has a groove or excavation which is occupied by the splenic blood vessels. This gland differs from the other large glands of the abdomen, inasmuch as it has not a large artery particularly appropriated to it ; but instead of this, it receives branches from the contiguous arteries. The arterial blood of this gland is partly supplied by the splenic artery, which, in its course from the main trunk of the cceliac to the spleen, while it is in the groove at the edge of the pancreas, sends off into the gland one considerable branch called the great pancreatic, and a number of small branches, which go off in succession. In addition to these, the pan- creas receives vessels from one of the branches of the hepatic artery, before it sends off its great rami- fications, as well as small twigs from several other contiguous arteries. The veins correspond with the arteries, but ultimately are discharged into the vena portarum. The pancreas resembles the salivary glands in 144 Pancreatic Duct . colour, and also in texture ; for it it of a dull white colour with a tinge of red, and it appears to consist of small bodies of a granulated form, which are so arranged as to compose small masses or lobes that are united to each other by cellular membrane. — , Each of these granulated bodies receives one or more small arterial twigs, and from it proceeds not only a vein but a small excretory duct, which, uniting with similar ducts from the adjoining granu- lated portions or acini, forms a larger duct in each lobe or mass; these open into the great duct of the gland, which proceeds through it lengthwise from the left extremity, in which it commences, to the right. This duet is situated in the body of the gland, which must be dissected to bring it into view. It is thin and transparent, like the ducts of the salivary glands, and is rather larger in diameter than a crow’s quill. In its progress towards the right extremity of the gland it gradually enlarges, and commonly receives a branch from the part called the lesser pancreas. It most commonly unites with the biliary duct before it opens into the duodenum : sometimes these ducts open separately, but very near to each other. They penetrate the coats of the intestine rather obliquely, and between four and five inches from the pylorus. This canal is sometimes called Dustus Virsungii , after an anatomist who published a plate of it. The pancreas has an irregular surface, and no coat which covers it uniformly. It is invested by cellular membrane, which also connects its different lobes to each other. Absorbent vessels and nerves are traced into it. The portion called the lesser pancreas adheres to the duodenum, and when it is enlarged by disease., 145 Size of the Spleen. the passage of aliment through that intestine is much impeded, and sometimes completely obstructed.* It is now generally believed that the fluid secreted by the pancreas is similar to that which is produced by the salivary glands. SECTION III. Of the Spleen. The Spleen is a flat body of a bluish colour, and an irregular oblong form, with thick edges, which are indented in some places. It is various in different subjects, both in size and form. Its most common size is between four and five inches in length, and about three or four inches in breadth ; but it has often been found of more than four times this size ; and it has also been seen not much longer than an inch. Its ordinary weight is be- tween six and nine ounces ; but it has varied in dif- ferent subjects from eleven pounds to one ounce. It is supposed, by many physiologists, that it frequent- ly varies in size in the same individual. It is situated in the left hypochondriac region, in contact with the diaphragm, below the eighth rib. The position of the spleen is somewhat oblique,— one extremity being directed downwards and rather forwards, and the other upwards and backwards ; but when the stomach is distended, the lower end of it is pushed forward by the great extremity of that viscus. In general it is so deeply seated in the left hypo- * In several cases where examination after death evinced that the pan- creas had become enlarged and indurated, particularly at the right ex- tremity, the principal symptoms wer e jaundice; great uneasiness after ta° ing food; vomiting some time after eating , but not immediately ; extreme acidity of the matter rejected Vol. lie- " 19 146 Enlargement of the Spleen chondriac region, that it is out of view when the sub- ject is opened in the ordinary way : but in some cases of enlargement, after the intermitting fever, it has extended downwards, nearly as low as the pelvis ; and towards the right side beyond the um- bilicus. The external surface of the spleen is convex, in conformity to the surface of the diaphragm, with which it is in contact. The internal surface of the spleen is irregularly concave, having a longitudinal fissure which divides it into two portions. The spleen is invested by the peritoneum, one process of which is often extended from the dia- phragm, above and behindit, in the form of ligament. Another process of the same membrane is extended to it from the great extremity of the stomach. The peritoneum is also continued from the spleen in the form of omentum. Within this peritoneal covering is the proper coat of the spleen, which is so closely connected to it, that many anatomists have considered them as one mem- brane : they are, however, very distinct at the great fissure, but the external coat is extremely thin. The proper coat of the spleen is not very thick : it is dense and firm, and somewhat elastic* but not much so. Iris partly transparent. The spleen has a large artery, which is one of the three great branches of the cceliac. This vessel runs in an undulating manner, in a groove in the upper edge of the pancreas, and in this course sends off many small branches to supply that gland. The splenic artery, before it arrives at the spleen, divides into five or six branches, which are also undulating in their progress, and penetrate into the body of the viscus at the above mentioned fissure. These branches are distributed to every part of the viscus, and ram- ify minutely. 147 Vessels of the Spleen , From these branches, or from the main trunk be- fore it ramifies, three or four smaller branches pro- ceed to the left extremity of the stomach. They are called vasa brevia or arterise breves. The arteries which enter the spleen are accom- panied by veins that emerge from it, and unite to form a great trunk. This trunk observes a course corresponding to that of the splenic artery, and re- * ceives veins from the stomach and pancreas, which * correspond with the arterial branches sent to those organs. The splenic vein is one of the principal branches of the vena portarum. The splenic artery is very large in proportion to the viscus to which it is sent, and the vein is unusu- ally large in proportion to the artery. The vein is also very tender and delicate in its structure. The absorbent vessels of the spleen are very nu- merous. It has been asserted, that when those of the external coat of the spleen are injected, they are sufficient to form a fine net-work on it. The absorb- ents of the deep-seated parts unite to the superficial at the*fissure where the blood vessels enter. They terminate in the thoracic duct, after passing through several lymphatic glands. The nerves of the spleen are derived from the solar .plexus : they form a plexus round the vessels and accompany them through the viscus. The spleen consists of a substance which is much softer than that of any other viscus of the abdomen. This substance is made up either wholly or in great part of the ramifications of the splenic artery and vein, which are demonstrated by injections to be very minute and numerous in this body. There are also many fine white cords, like threads, which pass from the internal surface of the inner coat of the spleen into its soft substance, in which some of them ramify. These cords connect the substance of the spleen 148 Malpighi on the Structure of the Spleen „ pretty firmly to its coat, and they seem to have the effect of rendering the exterior part of the substance more firm and dense than the internal. They are particularly conspicuous if the spleen be immersed in water, and the coat pulled off while it is in that situation. The spleen has a strong resemblance to the glan- dular organs, but has no excretory duct, and its par- ticular function is not very obvious : for these reasons the structure of this organ is a subject of very inte- resting inquiry. Malpighi, who took the lead in researches of this nature, before injections of the blood vessels with wax were in use, after investigating the structure of the spleen by long maceration, by boiling, by infla- tion, by the injection of ink or coloured fluids, and by examination with microscopes, declared that its structure was cellular ; that the cells communicated more freely with the veins than the arteries ; and that they might be considered as appendices of the veins. He also asserted, that a large number of white bodies or vesicles were to be found in those cells and throughout the whole substance of the spleen, which were in bunches like grapes, and pre- served their whitish colour although the vessels around them were injected with a coloured fluid. This description of Malpighi appears to have been admitted by some of the very respectable anatomists who were cotemporary with him ; but it was most zealously opposed by Ruysch, who exhibited the spleen so completely injected with wax, that it ap- peared to be composed entirely of vessels.* Ruysch appears to have paid great attention to * Two plates, taken from drawings of these preparations, are published in Ruysch’s works. One is attached to Epistola Problematica Quarta, in the second volume ; and the other to Thesaurus Septimus, in the third volume. 149 De La Sone on the Spleen „ this subject, and to have made many preparations of the spleen. From these he derived the opinion, that the substance of this organ was entirely composed of arteries, veins, absorbent vessels and nerves ; and that if it were properly injected before it was dis- sected, no other structure would be found. He stated, that the minute ramifications of the blood vessels appeared to have acquired a peculiar quality, and were so soft and delicate, that their texture was destroyed by the least friction ; and that by the slightest degree of putrefaction they appeared to be reduced to a fluid state. He also denied the exist- ence of cells, or of the whitish bodies described by Malpighi. The question thus at issue between these great masters of their art, was very carefully examined by M. De La Sone, a French physician, whose observa- tions are published in the Memoirs of the Academy of Sciences for 1754. After repeating the proces- ses of each of these anatomists, and instituting some others in addition, he adopted the opinion that there was in the texture of the spleen a pulpy substance which was not a mere coagulum, but which, howev- er, could not be injected. He derived his opinion from this fact among others. After macerating the spleen a considerable time, and injecting water into the vessels until it returned co- lourless, he injected ink, and confined it some time in the vessels by tying them : he then allowed the ink to flow out of the vessels, and made various sections of the spleen, but no ink appeared in the pulpy sub- stance, although it was visible in many small vessels which ramified in that substance. He observes that this could not have been the case, if the pulpy sub- stance had been composed entirely of vessels, as was supposed by Ruyseh. He also examined the spleen after it had been in- 150 Haller and the French Anatomists on the Spleen . jected with wax, according. to the manner of Ruyseh, and believed not only that the pulpy matter remain- ed uninjected, but that Ruyseh himself, in his own preparations, removed this substance, supposing it to exist for the mere purpose of connecting the vessels to each other. To see the blood vessels in the same state of de- tention in which they were during life, he tied the splenic vessels in a living animal, and removed the spleen with the ligatures on the vessels. In this situation he boiled it, and then examined the ap- pearance of the vessels and the pulpy substance. — From these, as well as his other observations, he decided, that the pulpy substance did not consist en- tirely of vessels, but was an additional and different structure. He also suggested, that as the brain and the mus- cular fibres were so covered by blood vessels in the injected preparations of Ruyseh, that they appeared to be composed entirely of vessels, when in fact they consisted of a different substance, so the pulpy sub- stance of the spleen was covered ar obscured by the blood vessels which passed through it, without con- stituting its whole substance. He confirms the account of Malpighi respecting the Whitish Vesicles or Follicles ; and states, that, in a majority of cases they are not to be discovered with- out a particular preparation ; but that they are gene- rally made obvious by long maceration of the spleen in water. In his opinion they are the most essential part of the organ. Notwithstanding these investigations of M. De La Sone, the question respecting the structure of the spleen remains not completely decided even to this day. Haller, who was perfectly well acquainted with the subject, inclined to the opinion of Ruyseh ; 151 British Anatomists on the Spleen, while Sabatier adopted completely the opinion of De La Sone; It appears from the statement of Gavard, that De- sault did not admit the existence of the transparent bodies; although he believed that the pulpy substance of the spleen, consisted of cells which resembled those of the cavernous bodies of the penis. Boyer, whose descriptions of the animal structure appear to have been formed with scrupulous exacti- tude, admits the existence of transparent bodies; sometimes so small as to be scarcely visible, and sometimes as large as the head of a pin.’ He ob- serves, that the best method of examining them is to place a very thin slice of the spleen between the eye and a strong light, when the transparency of these bodies occasions the slice of the spleen to appear as if perforated. As to the general structure of the pulpy substance, he avows himself unable to decide respecting if; but observes, that upon examining the cut surface of the spleen, you perceive black liquid blood flow from the vessels ; if you then scrape this surface, you may express easily a species of sanies different from that which flows from the vessels, which, after exposure, becomes red, and resembles coagulated blood ; whe- ther this is contained in the capillary vessels, or in the cavities of this organ, he acknowledges himself unable to determine. Notwithstanding the sentiments of these French gentlemen, many of the British Anatoihists, who are entitled to great attention on account of their skill in minute injections, have adopted the ideas of Ruysch. Among these are to be mentioned the late Dr. F, Nicholls, and many of- the anatomists of London, as well as the second Professor Monro, of Edinburgh. There are, however, two remarkable exceptions to this account of the British anatomists* The late 152 British Anatomists on the Spleen* Mr. Faleonar, who wrote a dissertation on the situa- tion and structure of the spleen, which contains the sentiments of the late truly respectable Mr. Hewsoq,* after stating that the organ was extremely vascular, so that when injected it appeared like a mere conge- ries of vessels, makes this unequivocal assertion — that there are innumerable cells dispersed through- out the whole substance of it, which are so small that they are only to be discovered by the aid of a micro - scope ; and are to be seen after steeping a thin piece of spleen, the blood vessels of which have been mi- nutely injected, in clear water during a day, and changing the water frequently. He also adds, that the ultimate branches of the arteries and veins form a beautiful net-work on each cell ; and that these cells are sufficiently distinguished from the irregular inter- stices of the cellular substance, by their round figure and their great regularity. Mr. Everard Home, in his papers on the structure and uses of the spleen, confirms the account of the ve- sicles in this organ ; and adds that these vesicles are occasionally seen in a distended and in a contracted state. That when distended they are twice as large as when contracted, and are distinguishable by the naked eye ; whereas, when contracted, they require a magnifying glass to be distinctly seen. These ob- servations appear to have been made upon quadru- peds.! Professor Soemmering appears to unite in the ge~ neral sentiment of the British anatomists, that the spleen is simply vascular. He says, that the tuber- culi which sometimes appear in it, when examined with a magnifying glass appear to be composed en- tirely of vessels. * See Experimental Inquiries, voi. iii. f See the London Philosophical Transactions for 1808. Questions relating to the Structure of the Spleen. 153 There are therefore two questions not perfectly de- cided respecting the spleen. First. Whether its general structure is simply vas- cular, or whether there is any other structure either cellular or more substantial, which composes its ge- neral bulk. Second. Whether the small transparent vesicles, originally described hy Malpighi, are to be regarded as essential parts of the structure of the spleen. With respect to the first question, the injections of Ruysch, and of the British Anatomists in general, and even of Mr. Hewson, as well as of Haller and Soemmering, seem to afford positive facts in opposi- tion to those of a negative kind adduced by M. De La Sorie, and render it highly probable that the ge- neral structure is simply vascular. But the second question stands on differentgrounds* The existence of small transparent vesicles, although denied by Ruysch, and neglected by the British Ana- tomists in general, was asserted as a positive fact by Malpighi and De La Sone ; and their assertions have been confirmed, not only by most of the French Ana- tomists, but also by Hewson and Home among the British. The sentiments of physiologists respecting the func- tions of the spleen, are more discordant than those of anatomists respecting its structure ; although the sub- ject has been considered by many authors of great in- genuity.^ * See M. Lieutaud. Elementa Physiologic?, Hewson’s Experimental Inquiries, 'vol. iii. Dr. Rush. Medical Museum, vol. iii.' Haller. Elementa Pbysiologis, tom. vi. page'414. Yol, n» 20 CHAPTER IV* OF THE URINARY ORGANS AND THE GLANDULE RENALES. The urinary organs consist of the Kidneys, which are situated in the lumbar regions ; of the Bladder, which is in the pelvis ; of the Ureters, which are flexible tubes or canals that pass from the kidneys to the bladder; and of the Urethra, or tube through which the urine is discharged from the bladder. These organs have but little connexion with the peritoneum. The kidneys are behind it ; and a con- siderable quantity of cellular membrane is placed between them and it. The ureters are also behind it; and but a part of the bladder is invested with it. The Glandulx Benales are described with the uri- nary organs, on account of their contiguity to'the kid- neys : and to avoid a derangement of the natural or- der of description they are considered first. The urethra pertains to the organs of generation as well as to the urinary organs, and can be descri- bed most advantageously with them. SECTION I. Of the Glandulx Benales . These are two small bodies, situated on the psoas muscles, one on each side of the spine, behind the peritoneum and above the kidney, being in contact with its upper and anterior edge. They have an ir- regular semilunar figure with three sides, one of The Kidneys and Ureters . 155 which is accommodated to the convexity of the kid- ney. Their colour is commonly a dull yellow. The appearance and texture of these bodies have some resemblance to those of glands, and hence their name, but they have no excretory duct. When they are laid open by an incision, a cavity often appears, which is somewhat triangular, and from the lower part of it a small thin ridge arises.^ A small quantity of fluid is generally found in it, which has a very dark colour in adults, is yellowish in young subjects, and red in infants. These bodies have not a single artery appropriated to them, as the spleen has, but receive small branches from several contiguous sources ; viz. from the arte- ries of the diaphragm, from the coeliac artery or the aorta, and from the arteries of the kidneys. There is generally one principal vein, as well as some that are smaller, belonging to each of these bodies : the large vein on the right side generally opens into the vena cava, and on the left into the left eraulgent vein. These bodies were first described by Eustachius, and have been regarded with attention by many ana- tomists since that period. They exist in a great num- ber of animals ; but their nature and functions are altogether unknown. SECTION II. Of the Kidneys and Ureters. The kidneys are two glandular bodies which se- crete the urine. They are of a dull red colour, and their form has a strong resemblance to that of the bean which bears their name. They have a peeu- * The cavity in these bodies has sometimes been sought for in vain. Haller found it in sixteen cases out of nineteen. 156 The Kidneys . ' liar texture, which is uniform, and not granulated or composed of acini ; and they are covered by a thin delicate tunic, which has no connexion with the pe- ritoneum. They are situated in the lumbar regions of the ab- domen, one on each side of the spine. They are opposite to the two last dorsal and the two first lum- bar vertebrae. They rest principally upon the psoas and quadratus lumborum muscles, and their posi- tion is oblique ; the concave edge presenting inwards and forwards, the convex edge backwards, and the upper extremity approaching nearer to the spine than the lower. The Right Kidney is situated rather low'er than the left : it is below the posterior part of the right lobe of the liver, and behind the duodenum and the colon. The Left Kidney is below the spleen, and behind the descending portion of the colon. Each of the kidneys is below' and very near to one of the glandulae renales. They are surrounded with a large quantity of lax adipose membrane, which in corpulent persons forms a very large mass of adeps around them ; while in the emaciated they are surrounded with a membrane almost free from fat. Each kidney has two broad sides, two extremities, and two edges. The side or surface which is posterior, when the kidney is in its natural situation, is rather broader than the other. The upper extremity, or portion, is also broader and larger than the lower. The edge which is pos- terior and external is regularly convex ; the anterior edge is concave ; but the concave edge, or margin, is not very regular. In the middle it is largely in- dented ; in this indentation is a deep fissure, which separates the two broad surfaces or sides of the gland from each other; and here the breadth of the posterior surface is evidently greater than the anterior. 157 . The Kidneys. Each of the kidneys receives a large artery, which proceeds immediately from the aorta, nearly in a rectangular direction. A vein, which opens into the vena cava, accompanies the artery. It is obvious, from the situation of the kidneys with respect to the great vessels, that the artery on the right side must be longer than that on the left, and that the reverse of this must be the case with the veins; the veins are also anterior to the arteries. At the great fissure these vessels divide into several branches, w T hich enter the kidney at that place. The branches of the vein are before and above ; those of the artery are below, and in the middle. Surrounded more or less by the branches of those vessels, is a membranous sac, the breadth of which extends from above downwards. This sac terminates in a tube that proceeds from the lower part of the fissure down to the bladder. The sac is denominated the pelvis of the kidney, and the tube a ureter: each of these parts will soon be more particularly described. The substance of the kidney, as has been already said, is uniform in its texture, and of a reddish brown colour. When it is divided by an incision made lengthways, and from its convex to its concave edge, there appears to be a small difference in the different parts of it. The exterior part, which is called corti- cal, is rather more pale in colour and softer in con- sistence than the internal part. It varies in thickness, so that some writers have described it as equal to two lines, and others to one-third of the kidney. In a majority of subjects it will be. found between the two statements. The interior part is called medullary 7 , or tubular, and appears to be composed of very fine tubes. These tubes are so arranged, that a number of pa- pillae or cones are formed by their convergence, and project into the fissure of the kidney. These pa- 158 The Kidneys . 4 pillae have been supposed to consist of a substance different from either of the two above mentioned, but they appear to be formed merely by the tubular part. The arteries, accompanied by corresponding veins, and by nerves and absorbent vessels, after ramifying in the fissure of the kidney, proceed into its substance, and continue their aborescent ramifications until they have arrived very near the exterior surface. They are so uniformly distributed to the different parts of the organ, that when the blood vessels are injected with wax, and the substance of the kidney is removed from the injected matter, as is the case in corroded preparations, the injection exhibits accurately the form of the kidney. The large branches of the blood vessels occupy the vacuities between the papillae in the fissure of the kidney. When they penetrate the substance of the kidney, they are enclosed by sheaths which are derived from the cqat of the gland, and are sur- rounded by membrane, which frequently contains adeps. There are commonly ten or twelve papillae in the fissure of each kidney, but there are sometimes more and sometimes less than this number. These pa- pillae are surrounded by a membranous sac of a cor- responding form ; the papilla being a cone, and the sac resembling the upper part of a funnel. The sac is therefore called an infundibulum, or calyx. Sometimes there are two papillae in each infundibu- lum, and then the form of the sac is not so regular. The infundibulum adheres to the base of the papilla, but lies loose about the other parts of it. Each in- fundibulum communicates, at its apex, with the pel- vis of the kidney. The Pelvis, as has been already mentioned, is a membranous sac which terminates in the ureter, ex- Cortical and Tubular Portions of the Kidney . 159 terior to the kidney. This sac generally divides it- self, in the fissure of the kidney, into three large ir- regular branches, each of which very soon terminates in three or four of the infundibula above described. That portion of the sac which terminates in the ure- ter is exterior to the kidney. When the interior parts of the kidney are exposed, to view, by the section above mentioned, after the arteries and veins have been minutely injected, the cortical part will be found to consist almost entirely of the minute ramifications of these vessels. Among them are. some small bodies, which are dispersed through the substance, like berries on a bush : these are asserted also to be composed of vessels. The tubular part certainly proceeds from this vas- cular cortical substance : for Ruysch, and after him several other injectors, have filled these tubes with injection thrown into the arteries. The tubuli, of which this part is composed, seem to arise obscurely from the cortical part. They soon assume somewhat of a radiated direction, and are finally arranged so as to form the papillae or, cones above described. On these papilla or cones some of them can be traced, uniting with each other, to form larger tubes, which terminate on the surfaces of the papillae, in orifices large enough to be seen distinctly. From these orifices urine may be forced out by compress- ing the papillae. On this account the tubes have been called tubuli uriniferi. , In the foetal state the kidney is formed of a num- ber of distinct lobuli, each of which consists of a papillae with the cortical matter connected to its base. Soon after birth these lobuli coalesce ; and in two or three years the substance of the kidney appears uniform, as above described. In some ani- mals this lobulated structure continues during life ; 160 Nerves and Lymphatics.— Coat of the Kidney. in them, and also in the foetus, each lobe appears like a distinct organ. Although in the adult kidney this structure has disappeared, the papillae and the tubular part connected with them are somewhat de- tached from each other, in a manner corresponding to their original arrangement. . The Nerves of the Kidneys originate from the se- milunar ganglion, formerly mentioned. They form a plexus round the blood vessels, and go with them into the gland. The kidneys have internal and external absorbent vessels, although the external vessels are very small. These absorbents pass through glands in the lum- bar region to the thoracic duct. The proper coat of the kidney is said, by some anatomists, to consist of two lamina ; but this can- not be shown in common cases. It appears simple in its structure, and very flexible. It is but slightly, connected to the glandular substance which it en- closes, and may be easily peeled off. It is reflected inwards at the fissure of the kidney, and can be traced inwardly to some distance, forming sheaths for the vessels. In this internal situation it is very thin. The Ureters. The pelvis of the kidney terminates exteriorly in the Ureter , which is a membranous cylindrical tube, rather flattened, and between three and five lines in diameter, with some variations in this respect. The Ureters descend from the pelvis of the kid- ney so as to pass obliquely across the psoas muscle and the great iliac vessels. They are behind the peritoneum, but in contact with it. They approach the pelvis near the junction of the os ilium with the sacrum, and thence descend forwards and inwards, surrounded with loose cellular membrane, to the Structure of the Ureters. 161 lower part of the bladder, into which they are in- serted at its external and posterior part. They first penetrate obliquely the muscular coat, and then pro- ceed between the muscular and internal coats, from half an inch to an inch, in an oblique direction, from without inwards and forwards, when they terminate by small orifices in the internal coat, each of which is at an equal distance, (rather more than an inch,) from the orifice of the urethra, thus forming a triangle with it. The ureters are said to have three coats. The ex- terior appear to be derived from the cellular sub- stance : within it is another, which has been regarded very differently by different anatomists ; some consi- dering it as merely membranous, and others as mus- cular. If the ureter be laid open and the internal coat peeled off, the muscular structure of this coat is often very perceptible. The internal coat is called villous, or mucous, and is continued from the internal coat of the bladder. Over this coat mucous is constantly spread, which defends it from the acrimony of the urine. It is very difficult to separate the two last mentioned coats from each other. The ureters receive blood vessels and nerves from those of the neighbouring parts. Their internal coat is very vascular, and is also very sensible of ir- ritation. The passage of a small urinary calculus can be traced from the pelvis of the kidney to the bladder, by the exquisite pain and the spasmodic af- fections which it often excites. Vol, II. 21 162 Situation of the Bladder . SECTION III. Of the Urinary Bladder. The urinary bladder is a large sac, of a muscular and membranous structure, which occupies the ante- rior part of the cavity of the pelvis, immediately within the ossa pubis. The size of the bladder is in a continued state of variation, according to the quantity of urine secreted. When moderately distended, it is of an irregular oval form, but rather more flat at its lower extremity than above. It varies in form according to the dif- ferent circumstances of the pelvis to which it has been subjected. It is fixed firmly and immoveably to the pelvis immediately within the symphysis pubis ; so that it is always to be found there of a larger or smaller size. This fixture is produced by the attachment of the lower portion of fundus of the bladder to the parts beneath it, but principally by the anterior liga- ments of the bladder which proceed one on each side from the lateral surfaces of the prostate gland, and are inserted into the pubis of the corresponding side at the lower part of the symphysis. These ligaments are in fact the extension of a membrane, (called by the French Anatomists, the pelvic aponeurosis,) which proceeds from the upper part of the pelvis to the side of the prostate gland and bladder, and which may be seen by turning off the peritoneum from the levator ani muscle.* It is sometimes completely empty, and occupies no more space than the thickness * See Thesis on Femoral Hernia, &c. by Gilbert Breschet. Paris, April, 1819. Colies’ Surgical Anatomy, Dublin, 1811, for a more minute account of this membrane. Ed-. 163 Situation of the Bladder. of its coats requires. When moderately distended, it occupies a considerable portion of the pelvis : when distention increases, it presses the parts poste- rior to it against the sacrum, and extends itself above the brim of the pelvis into the general cavity, rising not only to the umbilicus, but in some eases to? the epigastric region. In males the relative situation of the bladder and rectum is such, that the upper and middle part of the rectum is behind the bladder ; but the lower part of the rectum, following the curve of the os sacrum and coccygis, is below the posterior part of the bladder. In females the vagina and uterus are situated be- tween the bladder and rectum ; so that the connex- ion of these last mentioned parts is very different in the two sexes. The peritoneum is reflected at the anterior part of the brim of the pelvis from the abdominal muscles, which it lines, to the upper part of the bladder, which is generally contiguous to the brim of the pel- vis. It continues over to the posterior side of the bladder, and passes down upon it some distance to- wards the lower part ; but before it has arrived at the bottom, it is reflected towards the sacrum. In males it extends from the bladder to the rectum, and in females to the vagina and uterus ; so that there is a considerable portion of the lower part of the bladder which is not invested by the peritoneum. It also follows that when the bladder is extended into the abdomen, and rises above the brim of the pelvis, that part of it which presents anteriorly, and is in contact with the abdominal muscles, is without a co- vering of peritoneum, being below it. The bladder is composed of a coat consisting of muscular fibres, of a stratum of cellular substance immediately within this, and of an internal lining 1 64 Muscular Coal of the Bladder. membrane, which lias been called villous, but, as there are no villi perceptible on it, may be more properly denominated mucous. It should be observed, that, in addition to these coats, the bladder has a peculiar investment of the peritoneum, as has been already described; and also of the common cellular membrane, which is placed between it and every part to which it is con- tiguous. The Muscular Coat of the Bladder consists of fibres which are not spread over it of a uniform thick- ness, but are thin in some places, and in others are collected in fasciculi. They run in every direction: some appear longitudinal, others circular, and some oblique ; and there are interstices between them which are occupied by cellular membrane. The longitudinal fibres originate from the lower part of the bladder ; and as this is the fixed part of that viscus, it is the place from which these fibres must necessarily act. These fibres are generally exterior. There is no arrangement of muscular fibres to which the term of sphincter can properly be applied ; but many anatomists have thought that the fibres near the neck of the bladder, by their separate contrac- tion, might prevent the escape of urine ; this senti- ment, however, is contrary to that of several very respectable writers. The direction of the fibres, taken collectively, is such, that, when they all contract, the cavity of the bladder is completely obliterated. The cellular substance between the muscular and internal coats is dense. It yields in a remarkable manner to distension, and recovers its original dimen- sions very easily. From its analogy to a similar coat in the intestines, it is called the Nervous Coat. The Internal Coat of the bladder is of a light co- lour in the dead subject, when it has been free from 165 Internal Coat of the Bladder . disease. It had been called villous improperly ; for the villous structure is not apparent upon its surface. Being continued from the integuments of the body which are extended along the urethra, it has been in- ferred, that the surface of this coat was formed by the epidermis; and some respectable authors have sup- posed that they had seen cases in which portions of the epidermis of the bladder had separated and been discharged ; but these appearances are very equivo- cal, and it is by no means certain that an epidermis exists there.* The fasciculi of fibres of the muscular coat occasion this coat to appear very irregular, but these irregula- rities correspond exactly with the arrangement of the fibres of the muscular coat. When the internal coat is separated by dissection from the muscular, its surface is very smooth and uniform. In the recent subject, when no disease has previously existed, it is always spread over with mucous of a light colour, but nearly transparent, which can be easily scraped off. This mucous is spread upon the surface so uniformly, that it must be derived from sources which are situated upon every part of the surface ; but these sources are not very obvious. On the membrane of the nose the orifices of many mucous ducts are very visible, but such orifices are not to be seen on this surface. — • Haller mentions that he has seen mucous glands near the neck of the bladder; and it is stated by the pupils of Desault, that, in one of his courses, he pointed out a number of these glands, in a subject who had been afflicted with a catarrhal affection of the bladder. * In the fauces and the follicles of the tonsils an effusion of coagulable matter, in consequence of inflammation, often forms crusts, that may be mistaken for sloughs of the integuments, although those integuments re- main entire. 166 Mucous of the Bladder. Notwithstanding that the sources of this mucous are obscure, the quantity of it is sometimes immense. In some cases, where the secretion is increased by the irritation of a calculus in the bladder, the urine is rendered somewhat viscid and white coloured by the mucous mixed with it ; which, after the urine has been allowed to remain for some time, subsides in such quantities as demonstrates that many ounces must be secreted in the course of the twenty-four hours. The same circumstances occur, without the irritation of calculus, in the disease called catarrhus vesica}.* It is probable that, in healthy persons, a great deal of it passes ofF unperceived, being dissolved or dif- fused in the urine. From the quantity and the regu- lar diffusion of this mucous on the surface of the bladder, there is the greatest reason for believing that is is effused from every part of the surface; and it is a question that has not been decided whether it is discharged from glandular ducts too small to be perceived, or from the exhalent extremities of the blood vessels. It is probable that the use of it is to defend the internal coat of the bladder from the acri- mony of the urine. The symptoms of a stone in the bladder, as well as of several other diseases, evince that this coat is en- dued with a great degree of sensibility. It is evident that the essential parts in the general structure of the bladder are the muscular coat and the internal coat last described : but in addition to the account of them, there are some other important circumstances to be noted in the description of this organ, It has been already stated, that the form of the bladder was an irregular oval, although it was * In some cases this mucous soon becomes putrid, and during the putre- factive process deposits a substance which appears to be calcareous. 167 Orifice of the Urethra. somewhat varied in different persons. The oval form is not much altered at the part called the neck of the bladder, where the urethra passes off from it. The orifice of the urethra is situated anteriority at the low- ermost part of the bladder. On the lower surface of the urethra, at its commencement, and on the bottom of the bladder, immediately connected with the urethra, is situated the Prostate Gland, (to be here- after described with the organs of generation,) which is a firm body, that adheres strongly both to the bladder and urethra. This circumstance gives par- ticular firmness and solidity to that part of the blad- der. It has also been observed, that the bladder is attached firmly to the ossa pubis, at its neck, about the origin of the urethra. Each of these circum- stances have an effect upon the orifice of the urethra ; and when the bladder is opened, and this orifice is examined from within, it appears to be kept open by the connexion of the bladder with the prostate, and has been very justly compared to the opening of the neck of a bottle into the great cavity of that vessel.* The orifices of the two ureters are at equal dis- tances from the orifice of the urethra, and form with it the angles of a triangle. That part of the internal surface of the bladder which is within this triangu- lar space, is more smooth than the remainder of the same surface, probably in consequence of the adhe- sion of the bladder to the prostate, and to other parts exterior to it. * The late Mr. Lieutaud, and after him the French anatomists of the present day, have described a small tubercle at the lower and posterior part of the orifice of the urethra, which resembles the uvula in form. It has not been noticed here; and M. Boyer states, that it is often scarcely perceptible. He, however, makes a remark which is very worthy of at- tention, viz. that it is very subject to enlargement in old people, forming a tumour which impedes the discharge of urine. Sabatier has also made the same observation. 168 Ligaments and Vessels of the Bladder. That part of the bottom of the bladder which is immediately behind the triangular space, is rather lower than this space; and but a small portion of cel- lular membrane exists between it and the rectum in males, and the vagina in females. The upper part of the bladder is connected with the umbilicus by means of a ligament, which passes between the peritoneum and the abdominal muscles. This ligament consists of three cords. One of these, which is in the middle, arises from the coats of the bladder, and was, in the foetus, the duct called ura- chus; the other two, which are connected to the bladder principally by cellular membrane, were ori- ginally the umbilical arteries.* The middle cord is of a light colour and fibrous structure ; it is thickest at the bladder, and gradually diminishes as it ap- proaches the umbilicus. In a few instances it has been found to be hollow. In its progress to the umbi- licus it becomes more or less blended with the linea alba or the tendons of the abdominal muscles. The other cords are generally solid. After passing from the umbilicus to the bladder, they continue on the sides of that viscus, and finally terminate at the hy- pogastric or internal iliac artery. In the very young subject these cords are invested by distinct processes of the peritoneum, but their position is exterior to the peritoneum. As the bladder is situated very near most of the large ramifications of the hypogastric artery in the pelvis, it receives branches from several of them; viz. from the umbilical arteries before they termi- nate ; from the pubic ; from the obturators, &c. These branches ramify in the cellular membrane exterior to the muscular coat, and also in the cellular sub- stance between the muscular and internal coats. It * See the accounts of these parts in the description of the Abdomen of the Foetus. General Obsei'vations. 169 has been conjectured, that their terminations in ex- halents on the surface of the bladder are remarkably numerous. The veins correspond with the arteries, but they are very numerous on the lower and lateral parts of the bladder, and by uniting with the veins of the rectum form a remarkable plexus. The Lymphatic Vessels of this organ do not appear more numerous than those of other parts. They pass on each side the bladder in the course of its blood vessels, and unite with the larger lymphatics, and the glands which lie upon the great blood vessels on the sides of the pelvis. The JYerves of the bladder are derived both from the intercostal nerve and from the nerves of the me- dulla spinalis, which pass off through the sacrum ; and therefore the bladder is more affected than the viscera of the abdomen, by injuries of the medulla spinalis. The action of the muscular fibres of the bladder in ex- pelling urine, and the effect of those fibres which are situated near the orifice of the urethra in retaining it, can be considered with more advantage after the struc- ture of the urethra and the muscles connected with that canal have been described. It has been stated, that the internal coat of the blad- der is very sensible ; but it may be added, that in consequence of disease about the neck of the bladder, the natural sensibility appears most inordinately in- creased. When the intensity of pain which accom- panies these complaints, the frequent recurrence of paroxysms, and their duration, are taken into view, there seems reason to believe that none of the painful affections of the human race exceed those which arise from certain diseases of the bladder Happily these diseases are not very common. The functions of the kidneys is to secret urine, :nd that of the bladder to retain it, until the proper time for evacuation. Voi.. xr, 22 170 General Observations. The urine may be regarded as an excrementitious fluid, which contains many substances in solution that are constantly found in it, and many others that are occasionally in it, which are taken as aliment or me- dicine, and pass to the bladder with little, if any, change. The odour of the rose leaf, the colour of rhubarb, &c. are occasionally perceived in urine. The substances constantly found in urine are nume- rous. THe chemical account of the subject is so long, that it cannot be detailed here ; but the student ought to make himself acquainted with it, and he will read with great advantage Johnson’s History of Animal Chemistry, vol. 2d, page 363 ; and also Thompson’s Elements of Chemistry, page 333. CHAPTER V. OF THE MALE ORGANS OF GENERATION. These organs consist, 1st, of the Testicles, and their appendages. 2d. Of certain parts denominated the Vesiculse Seminales and the Prostate Gland , which are situ- ated near the commencement of the urethra, and are subservient to the purposes of generation. 3d. Of the Penis. SECTION I. Of the Testicles and their Appendages „ The Testicles are two bodies of a flattened oval form. Each of them has a protuberance on its upper and posterior part called Epididymis , and is connect- ed to parts within the cavity of the abdomen by a thick cord, which proceeds to it through the abdo- minal ring. Each testicle also appears to be contain- ed in a sac, which is suspended by this cord and co- vered by the common integuments. That portion of the common integuments which forms the external covering of the testicles, is deno- minated The Scrotum . The skin of the scrotum, although it is very often in a state of corrugation, has the same structure with that on other parts of the body, except that it is 3 *2 Structure of the Scrotum . lather thiner and more delieate. The superior deli- cacy of this portion of the skin is evinced by the great irritation produced by the application of stimulating substances, and the desquamation of the cuticle, which seems to be the effect of irritation. There are many sebaceous follicles in this portion of skin ; and after puberty there are often a few long hairs growing out of it, the bulbs of which are often very conspicuous. There is a small raised line in the middle of this skin, which commences at the root of the penis, and proceeds backwards, dividing it into two equal parts: this line is denominated Raphe. The corrugation which so- often takes place in the skin of the scrotum, appears to be occasioned by the contraction of certain fibres, which are in the cellu- lar substance immediately within it. This cellular substance appears to be attached in a particular way to the skin ; and it also invests each testicle in such a manner, that when they are withdrawn a cavity is left in it. It has long been observed, that no adipose matter is found in this cellular substance ; but it is often distended with water in hydropic diseases. As the contraction and corrugation of the scrotum has been imputed to this substance, it has been ex- amined with particular attention by anatomists, and very different sentiments have been entertained re- specting it. While some dissectors have asserted that muscular fibres could be seen in it, which they have denominated the Dartos Muscle ,* others have said that this substance was simply cellular, and without any muscular fibres. This difference of sen- timent may possibly have arisen from the different conditions of this part in different subjects ; for in some cases there are appearances which seem to jus- tify the assertion that muscular fibres exist in this structure. After the testicles are removed, so as to leave the 173 Action of the Dartos Muscle . cellular substance connected with the skin, if the scro- tum be inverted, and this substance examined in a strong light, many fibres will appear superadded to the common cellular structure ; and sometimes their colour can be distinguished to be red. It is not as- serted that this will be uniformly the case ; but cer- tainly it has often been observed in this way. The existence of an organ which possesses the power of contraction, within the skin of the scrotum and connected to it, is evinced by the corrugation which takes place when the scrotum is suddenly ex- posed to cold, after having been very warm. This corrugation occurs in a very sudden and rapid man- ner, in some cases, in which the wounded scrotum is thus exposed for the purpose of dressing : for ex- ample, upon removing an emollient poultice from this part some days after the operation for the cure of hydrops testis, by incision, if the air of the cham- ber be cool, a motion of the scrotum will take place, almost equal to the peristaltic movements of the in- testines. The Arteries of the scrotum are derived from two sources. One or two small arteries, which arise from the femoral artery, between Poupart’s ligament and the origin of the profunda, are spent upon it. These are called the external pudic arteries. It also re- ceives some small branches from the internal pudic artery. The Nerves of the scrotum are principally derived from the lumbar nerves. The Spermatic Cord. The cord which proceeds to the testicle through the abdominal ring, appears at first view like a bun- dle of muscular fibres ; but it consists of an artery and veins, with many lymphatic vessels and nerves, and also the excretory duct of the testicle, connected 174 Cremaster Muscle. —Blood vessels of the Testicle . to each other by cellular substance, and covered by an expansion of muscular fibres which are derived from the lower edge of the internal oblique muscle of the abdomen, and continue from it to the upper part of the testicle. These fibres constitute the Cremas- ter Muscle. The artery above mentioned is called the Sperma- tic. It commonly arises from the front of the aorta, very near its fellow, at a small distance below the emulgents : and is not much larger than a crow’s quill. It proceeds downwards behind the perito- neum and before the psoas muscle and ureter. — While it is in contact with the psoas muscle, it joins the ramifications of the vein. It afterwards meets the vas deferens, and proceeds through the abdomi- nal ring to the back part of the testis. Before it ar- rives at the testis it divides into several branches, two of which generally go to the epididymis, and the others penetrate the tunica albuginea on the upper and back of the testicle, and ramify very minutely on the fine membranous partitions which exist in that body. In addition to the spermatic artery, there is a small twig from the umbilical branch of the hypogastric* which passes to the spermatic cord along the vas de- ferens. The branches of the spermatic vein are much larg- er than those of the artery: several of them proceed from the testicle so as to correspond with the arterial branches ; and in addition to these there are many smaller, which also arise from the testicle and epidi- dymis. In their course up the cord they ramify, and again unite, so as to form a considerable plexus, which is called the Corpus Pampiniforme , and con- stitutes a considerable part of the volume of the sper- matic cord. As they proceed upwards they unite into a few Lymphatics and Nerves of the Spermatic Cord. 175 larger veins ; and finally, on the psoas muscle, they generally form one trunk, which continues upwards so as to unite with the vena cava on the right side, and the emulgent vein on the left. Sometimes, but not often, there are several sper- matic veins on each side. The Lymphatic Vessels of the testicle are very nu- merous, considering the size of the organ. Six or eight, and sometimes more, large trunks have been injected, running upon the cord, and continuing to the glands on the back part of the abdomen. The JYerves of the testicle are derived from those which supply the viscera of the abdomen, and are to be found in the cord although they can scarcely be traced to the testicle. A small plexus, called the spermatic, is formed by fibres from the renal plexus and from the sympathetic nerve. These fibres ac- company the spermatic vessels, and in all probability enter the body of the testis and the epididymis. The spermatic cord and cremaster muscle receive filaments from the second lumbar nerve. In addition to these vessels, the Vas Deferens, which is much firmer than either of them, is always to be distinguished in the back part of the cord. They are all covered in front and on the sides by the cremaster muscle, which passes with them from the lower margin of the internal oblique, through the abdominal ring, and continues to the upper part of the external coat of the testicle, which is a sac appa- rently containing that organ, and upon this sac it is spread out and terminates. The Tunica Vaginalis . The External Coat of the testicle, which is com- monly called the Tunica Vaginalis , is a complete sac which encloses the testicle as the pericaridum encloses the heart. It covers the body of the testicle 176 Coals of the Testicle . and epididymis, and adheres closely to them. It is then reflected from them so as to form a loose sac, which appears to contain them. The cavity of the tunica vaginalis commonly extends above the body of the testis up the cord, and is oval or pyriform. — This sac is so reflected from the body of the testicle that there is a place on the upper and back part of that body, at which the blood vessels enter it, without penetrating the sac. It resembles the peritoneum and other serous mem- branes in texture, and is therefore thin and delicate. It always contains a quantity of moisture, sufficient to lubricate the surface which it forms. When the tunica vaginalis is lain open, the testicle appears as if it were contained in the posterior part of its cavity. The testicles, as has been already stated, are of a flattened oval form. Their position is somewhat oblique, so that their upper extremities look upwards and forwards, their lower extremities downwards and backwards, and their edges present forwards and backwards. The body of the testicle is very firm, in conse- J quence of its enclosure in a very firm coat called Tu- nica Albuginea. Upon the upper and posterior part of it is the protuberant substance, called Epididymis , which is less firm, being exterior to the tunica albu- ginea. The blood vessels of the testicle pass into it on the posterior edge, at some distance below the upper end. The Tunica Albuginea , In which the body of the testicle is completely enclosed, is firm and dense ; and upon this coat its particular form depends. It is of a whitish colour, and has a smooth external surface. It is thick as well as strong. The epididymis is exterior to it. Form and Connexions of the Epididymis . 177 It is only perforated by the blood vessels, lympha- tics and nerves, and by the vasa efferentia, which carry out the secretion of the testis. One portion of the tunica vaginalis adheres very closely to it, and the other appears to contain it. The portion which adheres to it is with difficulty separated, but it is a distinct membrane. The Epididymis. Differs in colour from the testicle, being more or less reddish. It commences at the upper and ante- rior extremity of the testicle, and passes down the posterior edge to the lower end. At the commencement the epididymis is somewhat rounded in form, and its upper part, or head, has been called the globus major : as it descends it lessens, and about the middle of the testicle it is flatfish. It is firmly attached to the body of the testicle, at the upper end, where the vasa efferentia pass to it ; and it is also attached to it below ; but at the middle it appears nearly detached from it. It has therefore been compared to an arch resting with its two extre- mities on the back of the testis ; it is, however, in contact with it at its middle ; but about the middle it only adheres by one of its edges to the body of the testis, and generally by its internal edge. It has a coat which is less firm than the tunica albuginea of the testicle, described on the last page. The tunica vaginalis of the testicle is so reflected as to cover a great part of the epididymis which is not in contact with the testicle, and also those surfaces of the epi- didymis and testis which are in contact with each other and do not adhere. The Body of the Testicle. When the tunica albuginea is cut through, and the substance of the testicle examined, it appears Vol, iio 23 178 Tubuli, Septa and Bloodvessels of the Testicle » to consist of a soft pulpy substance of convoluted threads, of a yellowish brown colour, which is divi- ded into separate portions by very delicate septa, at- tached to the internal surface of the tunica albuginea at the posterior part of the testicle. After macera- tion, by using a fine needle to detach them from the cellular substance, these threads may be drawn out to a great length. In some animals they are larger than in the human species ; in them, it is said, they are evidently hollow, and that very small blood vessels appear in their Goats. When mercury is injected into the vas deferens, or excretory duct of the testis, in a retrograde course, it can be perceived in these ducts in the human subject. These delicate septa, or partitions, are united to the internal surface of the tunica albuginea at the posterior part of the testicle, at which place there is a body called Corpus Highmorianum, which has been regarded very differently by different anato- mists. It is a long whitish substance* which extends lengthwise on the posterior part of the testis ; and was supposed by Haller to resemble one of the sali- vary ducts. It is now, however, generally agreed to be of a cellular structure, and to contain and support the ducts which pass from the substance of the testi- cle to the epididymis. The blood vessels pass into the body of the testi- cle upon these septa, and are continued from them to the filaments or tubes of which the body of the testi- cle consists. As in some animals blood vessels are distinguished on these tubes, there is the greatest reason to believe that a direct communication sub- sists between them, without the intervention of any other structure, no other structure having been dis- covered : but at the same time it ought to be obser- ved, that these tubes have not yet been injected from the blood vessels. Some ingenious anatomists have Blood vessels and Structure of the Testicle, fyc. 179 injected the artery going to the testicle so success- fully, that the injection has passed from it into the veins coming out of the testicle ; but it is not now said by any of them, that they have filled the tubes in this manner. Mercury will pass into these vessels from the ex- cretory duct of the testicle ; and by means of an in- jection in that way, the structure of the testiele can be unravelled. 'this structure is as follows : The cavity formed by the tunica albuginea is divided into a number of apartments by the very thin septa, or partitions, above mentioned. From the filamentary or tubular matter which fills each of these chambers, proceeds a number of small tubes or vessels, which observe a straight course ; they are therefore called Vasa Recta . These vasa recta unite with each other and form a net-work on the back of the testis, within the tunica albuginea, which is called Rete Testis. From this net-work other vessels, from twelve to eighteen in number, denominated Vasa Ejferentia , proceed through the albuginea to the epididymis. These vessels are convoluted in such a manner as to form bundles of a conical form, which are called Coni Vasculosi. The number of these corresponds with the number of the vasa efferentia, and they com- pose about one third of the epididymis, viz. all the upper part of it. The single tubes which form each of these cones, successively unite into one duct, which is convoluted so as to form all the remainder of the epididymis. The lower part of the epididymis is turned upwards on the back of the testicle, the tube gradually enlarges and is less convoluted, and finally becomes straight : it then takes the name of Vas De- ferens , and continues on the back of the testicle and ISO Course of the Vas Deferens. at the inner side of the epididymis to the spermatic cord.*f A small solitary vessel or duct, has been observed by Haller, Monro, and several other anatomists, to proceed from the upper part of the epididymis : sometimes it Unites to the epididymis below, and sometimes it proceeds upwards. The nature of- this vessel has not been ascertained with certainty. The Vas Deferens Is a very firm tube about one line in diameter, which is not perfectly cylindrical exteriorly, although the cavity formed by it is so. This cavity is so small in diameter, that it will only admit a fine bristle. The coats of the duct have, of course, a considera- ble thickness. The internal coat forms a soft sur- face, analogous to that of the mucous membranes : the external is firm, and its texture resembles that of cartilage. Owing to the small size of the cavity, the internal coat has not been separated from the ex- ternal. * De Graff appears to have been the first anatomist who made much progress in the successful investigation of the structure of the testicle ; and Haller ought to be mentioned next to him, on account of the plate ex- hibiting this structure, and the explanation of it, which he published in the Philosophical transactions of London, for 1749. This plate has been re- published by the second Monro, in the Literary and Physical Essays of Edinburgh, and also in his Inaugural Thesis. Haller has likewise repub- lished it in his Opera Minora. It represents not only the vasa efferentia and the cones formed by their convolutions, but also the rete testis and the vasa recta. Haller could inject no further than this ; but Monro and Hunter soon after succeeded so as to fill a considerable portion of the body of the testicle with mercury, injected by the vas deferens. f In Mr. Charles Bell’s Anatomical collection in London, there is a pre- paration by his assistant, Mr. Shaw, in which the tubuli testis are com- pletely injected with quicksilver and unravelled. I saw also in Leyden, one nearly as successfully executed by Professor Sandifort.' — E d. 181 Course of the Vas Deferens. It passes upwards in the posterior part of the spermatic cord, and continues with it through the abdominal ring, under and exterior to the perito- neum ; soon after this it leaves the cord and dips down into the cavity of the pelvis, forming a curve on the side of the bladder, and proceeding back- wards, downwards and inwards. In this course it crosses the ureter, and passes between it and the bladder. On the lower part of the bladder the two vasa deferentia approach each other so gradually, that they appear to be nearly parallel. They pro- ceed forward between the vesiculse seminales, which are two bodies irregularly convoluted, that are placed in a converging position with respect to each other, and communicate with the vasa deferentia. The vasa deferentia finally terminate almost in contact with each other in the posterior part of the prostate gland, where they perforate the urethra. At the distance of about two inches and a half from their termination they enlarge in diameter, and become somewhat convoluted. At the posterior margin of the prostate they, come in contact with the anterior extremities of the vesiculse seminales, and unite with them. After this union they diminish in size, and become conical ; and passing a short distance through the substance of the prostate, during which they ap- proach each other more rapidly, they penetrate the urethra, so as. to open in it on each side of a small tubercle, called the Caput Gallinaginis } seon to be described. SECTION II, . Of the Vesiculse Seminales and the Prostate Glands The Vesiculse Seminales are two bodies of a whitish colour, and irregular form, being broad and flat at their posterior extremities, and terminating in 182 Structure of the Vesiculse Siminaies , a point at the other. Their surfaces are so convo- luted, that they have been compared to those of the brain. They are situated between the rectum and bladder, and are connected to each by cellular mem- brane. When the vesiculse seminales are laid open by an incision, they appear to consist of cells of a consider- able size, irregularly arranged ; but when they are carefully examined exteriorly, and the cellular mem- brane about them is detached and divided, they ap- pear to be formed by a tube of rather more than two lines diameter, and several inches in length, which terminates, like the caecum, in a closed extremity.? — * From this tube proceed from ten to fifteen short branches, which are closed in the same manner.-— All these tubes are convoluted so as to assume the form of the visiculm seminales above described ; and they are fixed in this convoluted state by cellular membrane, which firmly connects their different parts to each other. It is obvious, that tubes thus convo- luted, when cut into, will exhibit the appearance of cells, as in the present instance. This convoluted tube composing the vesiculse se- 3 minales, terminates in a very short duct, which is nearly of the same diameter with the vas deferens, and this duct joins the vas deferens so as to form an' acute angle. From the union of the vesicylss seminales with the vas deferens on each side, a canal, which seems to be the continuation of the vas deferens, proceeds i through part of the prostate to the urethra, which it perforates. These canals are from eight to twelve lines in length ; they are conical in form, their 3 largest extremity being equal to the vas deferens at ' 1 that part. If air or any other fluid be injected through the vas deferens into the urethra, it will pass at the same Function of the Vesieulse Seminales . 183 time into the vesieulse seminales, and distend them. It has been observed, that a fluid passes in this man- ner much more readily from the vasa deferentia into the vesieulae seminales, than it does from these last mentioned organs into the duct. These organs were generally regarded as reservoirs of semen, and analogous to the gall bladder in their functions, until the late Mr. J. Hunter published his opinion that they were not intended to contain semen, but to secrete a peculiar mucous subservient to the purposes of generation. He states the following facts in support of his opin- ion. A fluid, very different from semen, is found after death in the vesieulse seminales. In persons who have lost one testicle, a consider- able time before death, the vesieulse seminales on each side are equally distended with this peculiar fluid. In the case of a person who had a deficiency of the epididymis on one side and of the vas deferens on the other, the vesieulse were filled with their pe- culiar fluid S The sensation arising from redundance of the se- cretion of the testes, is referred to the testes, and not to the vesieulse seminales. In some animals there is no connexion between the vasa deferentia and the vesieulae seminales. See Observations on certain parts of the Animal Economy, by John Hunter*. The Prostate Gland. Is situated on the under and posterior part of the neck of the bladder, so as to surround the urethra. Its form has some resemblance to that of the ches- nut, but it has a notch on the basis like that of the figure of the heart on playing cards, and it is much larger than the chesnufc of this part of America,, 184 Prostate Gland.* — The Penis. The basis of this body is posterior, and its apex an- terior; its position is oblique, between the rectum and the symphisis pubis. Below there is in some cases a small furrow, which, in addition to the notch above, gives to the gland an appearance of being divided into two lobes. By turning away the vesicul® seminales and vasa deferentia from the under surface of the bladder we bring into view a small tubercle at the upper part of the base of the prostate, called by Sir Everard Home the third lobe. When diseased it projects into the cavity of the bladder. It adheres to the urethra and neck of the bladder. Its consistence is very firm and dense, resembling the induration of scirrhus rather more than the ordinary, texture of glands. This gland receives small branches from the neigh- bouring blood vessels, and has no artery of consider- able size exclusively appropriated to it. As it lies in close contact with the urethra, the ducts which pass between it and the urethra are not to be seen separate from these bodies ; but ducts can be seen in the substance of the gland, which perfo- rate the urethra, and open on the sides of the caput gallinaginis to the number of five or six on each side. By pressure a small quantity of a whitish fluid can be forced from these orifices, which is rather viscid, and coagulable in alcohol. The particular use of this fluid is not known. SECTION III. Of the Penis . The penis, when detached from the bladder, and the bones, to which it is connected, and divested of the skin which covers it, is an oblong body, which is rounded at one extremity and bifurcated at the other. Perns. 1&5 It is composed of three parts, viz two oblong bo- dies, called Corpora Cavernosa , which, at their com- mencement, form the bifurcated portions, and then unite to compose the body of the organ ; and a third part, of a spongy texture, which is connected to these bodies where they unite to each other, on the under side, and continues attached to them during the whole extent of their union, terminating in an expanded head which covers the anterior extremities of the corpora cavernosa. The urethra passes from the neck of the bladder, on the under side of the penis, to its anterior extremity, invested by this third body, which is therefore called Corpus Spongiosum Ure- thrae . The two bifurcated extremities are attached each of them to one of the crura of the pubis and ischium ; and they unite to form the body of the penis imme- diately anterior to the symphisis pubis, to which the lower part of it is also attached ; so that the penis is firmly connected to the middle of the anterior part of the pelvis. The urethra proceeds from the neck of the bladder, between the crura of the ischium and pubis and the crura of the penis, to join the body of the penis at its commencement, and near this place its connexion with the corpus spongiosum begins; so that there is a small portion of the urethra between the neck of the bladder and the commencement of the corpus spongiosum, which is not covered by the cor- pus spongiosum. This is called the membranous part of the urethra. The penis, therefore, consists of two oblong bo- dies of a cellular structure, which originate separate- ly, but unite together to form it ; and of the urethra, which joins these bodies immediately after their union, an$ is invested by a spongy covering, which by its expansion forms the anterior extremity not . only of the urethra but of the whole penis. These Vol, ii. 24 186 Corpora , Cavernosa Penis . three bodies, thus arranged and connected, are cover* ed by cellular membrane and skin in a manner to be hereafter described. The Corpora Cavernosa , Which compose the body of the penis, are two ir- regular cylinders, that are formed by a thick dense elastic membrane, of a whitish ligamentous appear- ance and great firmness. They are filled with a sub- stance of a cellular structure, which is occasionally distended with blood. The crura of these cylindri- cal bodies, which are attached to the crura of the ischium and pubis, are small and pointed at the com- mencement, and are united to the periosteum of the bones. In their progress upwards they enlarge, and at the symphisis of the pubis they unite so as to form an oblong body, which retains the appearance of a union of two cylinders applied to each other length- ways ; for above there is a superficial groove passing" in that direction, which is occupied by a large vein : and below there is a much deeper groove, in which the urethra is placed. Between these grooves is a septum which divides one side of the penis from the other. It appears to proceed from the strong mem- brane which forms the penis, and is composed of bundles of fibres, which pass from one groove of the penis to the other, with many intervals between them, through which blood or injection passes very freely. Sometimes these bundles of fibres, with their inter- vals, are so regularly arranged, that they have been compared to the teeth of a comb. This septum ex- tends from the union of the two crura to their termi- ;i nation. Each of these cylinders is penetrated by the main branch of the pudic artery, which is about equal in size to a crow’s quill. These arteries enter the cor- pora cavernosa near their union, and continue 181 Cells of the Corpora Cavernosa „ through their whole extent, sending off branches in their course ; the turgeseence and erection of the penis is unquestionably produced by the blood which flows through these vessels into the penis. The interior structure of the penis, when examined in the recent subject, is of a soft spongy nature, and seems stained with blood. If any fluid be injected through the arteries this substance appears cellular, and may be completely distended by it. When air is injected, and the structure becomes dry, the penis may be laid open ; the cellular structure then appears as if formed by a number of lamina and of filaments, which proceed from one part of the internal surface of the penis to another, and form irregular cells. It has been compared to thq lattice-work in the interior of bones j "and it is suggested by M. Roux, that the fibres of which the structure consists resemble those of the strong elastic coat of the penis.* If these cells are filled with coloured wax, injected by the artery, and the animal substance is then destroyed by placing the preparation in a corroding liquor, the wax which remains shows that the membranes forming the cells are very thin. These cells communicate freely with tfeaeh other ; and therefore, if a pipe be passed through the strong coat of the penis, the whole of them can be filled from it by the ordinary process of injection. * Mr. John Hunter says on this subject, “ That the cells of the corpora cavernosa are muscular, although no such appearance is to be observed in men ; for the penis in erection is not at all times equally distended. Tke penis, in a cold day, is not so large in erection as in a warm one : which probably arises from a kind of spasm, that could not act if it were not mus= eular. In the horse, the parts composing the cells of the penis appear evident- ly muscular to the eye, and in a horse just killed, they contract upon being stimulated.”— E d. 188 Corpus Spongiosum.— Bulb, The Urethra. Is a membranous canal, which extends from the neck of the bladder to the orifice at the extremity of the penis ; and for a very great part of its length is invested by a spongy structure, called the corpus spongiosum urethrae. It proceeds from the neck of the bladder along the upper part of the prostate; from the prostate it continues between the crura of the penis until their junction: it then occupies the great groove formed by the corpora cavernosa on the lower side of the penis, and continues to the orifice above mentioned. At a small distance from the pros- tate gland the spongy substance which invests it com- mences, and continues to its termination. After this spongy substance has arrived at the termination of the corpora cavernosa, it expands and forms a body of a particular figure which covers the extremities of the corpora cavernosa, and is denominated the Gians Penis. The Corpus Spongiosum begins at the distance of eight or ten lines from the anterior part of the pros- tate.* It is much larger at its commencement than at any other part except the glans, and this enlarged part is called the Bulb. It surrounds the whole of the urethra, and with the exception Of the bulb and the glans penis, is of a cylindrical figure. It is form- ed by a membrane which has some resemblance to the coat of the penis, but is much thinner, and by a pe- culiar spongy substance, which occupies the space be- tween the internal surface of this membrane and the external surface of the canal of the urethra. The membrane and the spongy substance, form a coat to the urethra, which, with the exception of the enlarge- ment before mentioned, is about one line thick. After this spongy substance has arrived at the termination,, its coat adheres firmiy to the coat of the penis. Structure of the Corpus Spongiosum. 189 The Bulb, or first enlargement of the corpus spon- giosum, is oblong, and rather oval in form ; it is mark- ed by a longitudinal depression in the middle, which is very superficial. It consists entirely of the spongy substance above mentioned. The Gians Penis is also composed of the same spongy substance, but the coat which covers it is more thin and delicate than that of the other parts of the urethra. The lower surface of the glans is fitted to' the extremities of the corpora cavernosa, but it is broader than the corpora cavernosa, and therefore projects over them on the upper and lateral parts of the surface of the penis. The edge of the promi- nent part is regularly rounded, and is denominated the Corona Glandis. Several small arteries pass to this spongy structure. The pudic artery, as it passes on each side to the cor- pora cavernosa, sends a branch to the bulb of the urethra. The same vessel in the substance of the penis, also sends branches to the urethra : and the artery on the back of the penis terminates in small branches, which penetrate the substance of the glans. By these vessels blood is carried to the spongy substance of the urethra, which is occasionally dis- tended in the same manner that the cavernous bodies of the penis are distended during the erection of that organ. But the cellular structure, of this organ is not so unequivocal as that of the corpora cavernosa ; for if it be injected with coloured wax, and corroded in the usual manner, the injected matter will exhibit an appearance which has the strongest resemblance to a convoluted vessel, like the vas deferens in the epididymis.* * Mr. Hunter says, “ that the corpus spongiosum urethrse and glans penis are not spongy or cellular, but made up of a plexus of veins. This structure is discernible in the human subject ; but is much more distinctly sees, in many animals, as the horse,” &e — - Ep. 190 Internal Surface of the Urethra . The Canal of the Urethra , Which conveys the urine from the bladder, is a very important part of the urinary organs. It consists of a vascular membrane with a smooth surface, which is perforated by the qrifices of many mucous follicles, some of which are of considerable size. It is ex- tremely sensible, and has so much power of contrac- tion, that some persons have supposed muscular fibres to exist in its structure. It is dilferently circumstanced in different parts of its course. While surrounded with the prostate it ad- heres firmly to that body, seeming to be supported by it ; and here its diameter is larger than it is farther forward. On the lower or posterior side of this por- tion of the urethra, is an oblong eminence, called Verumontanum, or Caput Cfallinaginis , which com- mences at the orifice of the urethra, and continues throughout the whole portion that is surrounded by the prostate gland, terminating at the point of that body. The posterior extremity of this tubercle begins abrupt^, and soon becomes thick and large ; ante- riorly it gradually diminishes to a line, which is some- times perceptible for a considerable distance in the urethra, in a straight forward direction. In the upper edge or top of this body is a groove, which ‘is produced by a mucous follicle ; on the lateral surfaces, anterior to the middle, are the orifices of the common ducts of the vesiculse seminales and vasa deferentia, (see page 182,) which are sufficiently large to receive a thick bristle. N ear these, on each side, are five or six smaller orifices of the excretory ducts of the prostate gland. At the distance of an inch before the extremity of the bulb of the urethra, in the lining membrane, are the openings of two ducts, one on each side, that lead to small glandular bodies called Cowper’s glands, which are situated on each side of the urethra below the bulb, but are covered bv the accelerator es urinse muscles. v <• 191 Mucous Ducts of the Urethra. The diameter of the urethra lessens after it leaves the prostate. That portion of the canal which is be- tween this gland and the bulb, without investment, and therefore called the membranous part, is the smallest in diameter. After it is invested with the spongy substance, it has a small enlargement, and then continues nearly of one size, until it arrives near the glans penis, when it again enlarges and alters its form, being no longer cylindrical but flattened. Its broad surfaces have now a lateral aspect. From the bulb of the corpus spongiosum to this last enlargement, the appearance of the inner surface of the urethra is uniform. The membrane is thin and delicate, and in a healthy subject, who has been free from disease of these parts, is of a whitish co- lour ; but blood vessels are very perceptible in it. When it is relaxed, it appears to be thrown into longitudinal wrinkles; but it admits of considerable extension, being somewhat elastic : when extended, its surface, appears smooth, as if it were covered with an .epithelium. Mr. Shaw, of London, has described a set of vessels immediately below the in- ternal membrane of the urethra, which, when empty, are very similar in appearance to muscular fibres.— He says he has discovered that these vessels form an internal spongy body, which passes down to the membranous .part of the urethra, and forms even a small bulb there. His preparation with a quicksilver injection of the part is certainly a very satisfactory proof of its existence.* Throughout the whole ex- tent of this part of the' urethra, are the orifices of a great many mucous ducts or sinuses, which pass obliquely backwards. Many of these are so small that they cannot be penetrated by a bristle, or probe * Sea Sled, Clururg—Tfansactions of Londca— vol 10th. 192 Mucous Dads of the Urethra a of that size; but some are larger. It has not been observed that any glandular body immediately sur- rounds them, although they secrete the mucous with which the urethra is lubricated. — On the lower side of the urethra, near the commencement of the glans penis, there is one or more of them, so large that their orifices sometimes admit the point of a small bougie.* These organs, when inflamed, secrete the puriform discharge which takes place in gonorrhoea. — In a na- tural state they produce 'the mucous which is con* stantly spread over the surface of the urethra, to de- fend it from the acrimony of the urine, and which passes away with that fluid unperceived. The surface of the urethra is endued with great sensibility, and is therefore liable to great irritation from contact with any rough body or any acrid sub- stance. Irritation, thus excited, induces a state of contraction, which is particularly remarkable, as no muscular fibres are to be seen in its structure. — When a bougie has been passed into the urethra for a consi- derable distance, if it cannot proceed the whole way, it sometimes happens that the instrument will be dis- charged by a steady uniform motion, which seems to proceed from a progressive contraction of the urethra, beginning very low down. At particular times, after the urethra has been much irritated, it will not receive a bougie, although at other times a bougie of equal size may be passed to the bladder without opposition. This cannot depend upon that elasticity which was noticed before. f * They were discovered by Plazzoni, of Padua, in 1621. Their number, according to Loder, amounts to about 65 — See his plates. Ed. f Sir Everard Home, whose professional opinions are of' great weight, has lately described in the Transactions of the Royal Society, the appear- ance of the lining membrane of the Urethra, when viewed through a Contractile Power of the Vrethra . 19J Upon the two crura of the penis, or the beginning of the corpora cavernosa, are fixed the muscles call- ed Ereetores Penis, which are described in the first volume.* These muscles cover the crura of the penis from their origin to their junction, and not only compress them, but also influence the motion oi the penis when it is distended. The bulb of the urethra is covered by a muscular coat, called the Accelerator Urinse * which has the effect of driving forwards any fluid contained in the cavity of the urethra, and also of giving the same direction to the blood in that part of the corpus spongiosum. There is also the Transversus Perinei on each side, that passes transversely from the tube- rosity of the ischium to the bulb of the urethra.— Finally the lower part of the sphincter ani muscle, which is nearly elliptical in form is inserted by its anterior point into the muscular covering of the bulb of the urethra. Upon removing the integuments, these muscles are in view ; and the course of the urethra from the bladder is concealed, particularly by the anterior point of the sphincter ani. When the sphincter ani is dissected away from its anterior connexions, and the cellular and adipose substance, which is sometimes very abundant, is also removed, the lower surface of the membranous part of the urethra may be brought into view, as it proceeds from the prostate gland to the balb of the corpus spongiosum.f microscope of great powers. From this paper it seems that he is full/ convinced of its muscular structure. — E d. * See description of “ Muscles about the Male Organs of Generation,' Vol. I. Part II. Chap. II. t The natural situation of the membranous part of the urethra, and of the prostate gland, as well as their relative position with respect to the sphincter ani, rectum, &c. can be best stadied by a lateral view of the Tol, ii, 25 194 Contractile Power of the Urethra . When the accelerator u rinse is removed from the bulb of the urethra, there will appear two bodies, which have some resemblance to flattened peas. They lie one on each side of the urethra, in contact or nearly so with its bulb, and from each gland pro- ceeds an excretory duct of an inch and a quarter in length, between the corpus spongiosum and the lining membrane of the canal of the urethra, and opens into the latter. Its orifice is found with some difficulty, but is large enough to admit a bristle. These are Cowper’s glands.^ The penis is connected to the symphisis pubis by a ligamentous substance, which proceeds from the back or upper surface of the organ to the anterior part of the symphisis, and connects these parts firm- ly to each other. Thus constructed, of the corpora cavernosa and the urethra with its corpus spongiosum, and attach- ed to the pelvis as above mentioned, the penis is in- vested with its integuments in the following man- ner, f contents of the pelvis ; which is to be obtained by removing carefully one of the ossa innominata, and dissecting the parts which were enclosed by it. * These glands were discovered by Mery, in 1684, and described by Cowper, in 1699. A third gland, smaller than the preceding, connected with the curve of the urethra under the symphisis pubis, was discovered by Cowper, and Morgagni speaks of having observed a fourth. — Ed. t There are several fasciae and ligaments about the perineum which should be connected with the account of its viscera. Immediately beneath the skin of the perineum is the Perineal Fascia, a thin but strong mem- brane, which extends from bone to bone, occupying the space between the anus and the posterior part of the scrotum. It is rather better seen in lean subjects than in fat ones, for in the latter it is converted in part into adipose membrane. When a rupture occurs in the posterior part of the urethra, this fascia prevents the urine from showing itself immedi- ately in the perineum, and drives it into the cellular structure of the scrotum. Immediately beneath the perineal faseia are placed the muscles ; when they are removed the bulb of the urethra may be seen very advantage- ously; extending in the middle of the perineum almost to the anus. It is Integuments of the Penis . 195 Integuments of the Penis . The glans penis, the structure of which has been already described, is covered by a continuation of the skin, which appears altered in its texture so as to resemble in some respects the skin of the lips, and in like manner is covered by a delicate production of cuticle. Around the corona of the glans, especially on its upper part, there are whitish tubercles, which are of different sizes in different persons, but always very small. The skin adheres firmly to the whole extent of the corona of the glans, and is very delicate in its structure as it continues from the glans upon the body of the penis ; but it gradually changes so as to assume the appearance and structure of a common skin, and continues in this state over the penis. The adhesion of the skin to the ligamentous coat of the corpora cavernosa also becomes more loose, owing to the quantity and texture of the cellular substance which connects them. The skin thus connected to the penis, has commonly more length than that or- gan, even in its extended state. In consequence of this greater length, and of its adhering firmly around the corona glandis, it necessarily forms a circular not loose and pendulous, but is attached by its pelvic surface to the trian- gular ligament of the urethra. This ligament is a septum between the perineum and the pelvis, and connects itself to the pelvic or internal edges of the rami of the pubis and ischii as far down as the organs of the crura penis. It extends from the arch of the pubis to the line mentioned, and fills up all the space between the bones of the opposite sides. It consists of two lamina, and Cowper’s glands are placed between them. About an inch below the symphisis pubis a perforation is made in this lig- ament for the passage of the membranous part of the urethra. Just below the symphisis pubis, between the two lamina of the triangu- lar ligament, is placed a much stronger ligament called the pubic, which is about half an inch broad, its lower edge is thick and rounded. For further detail on the subject of the:fasciae of the pelvis see Lessons in Practical Anatomy by the present editor.— Ep. A 96 Integuments of the Penis, fold or plait, which varies in size according to the length of the skin. This fold is generally situated at the commencement of the firm attachment of the skin to the body of the penis, or around the glans; but it may be formed any where upon the body of the penis by artificial management. This duplicature, or fold of the skin, when it takes place so as to cover the glans, is called the Prepuce ; and the skin which is very tender and delicate for some distance from the glans, form that surface of the prepuce which is in contact with the glans when it covers that body. There is also a small fold of the skin, which is longitudinal in its direction, that commences at the orifice of the urethra, and extends backwards, on the lower surface of the penis. It is unvarying in its position, and is called the Frenum. It is a general observation, that adeps is not found in the cellular substance which connects the skin to the body of the penis ; but this cellular substance is distended with water in some hydropic cases. From the skin immediately below the glans, and from small follicles on each side of the frenum, is secreted an unctuous fluid, which, when allowed to continue, becomes inspissated, and acquires a caseous consistence and colour, as well as a peculiar odour. It sometimes also acquires an acrimony which produ- ces inflammation on the surface with which it is in contact, as well as the copious seeretion of puriform fluid. The distribution of the pudic artery in the penis, has already been mentioned; and a further account of its origin and progress to its destination, will be found in the general account of the arteries. Some- times small branches of the external pudic arteries, which originate from the femoral, are extended to the penis; and it has been asserted, that branches of Blood Vessels and Absorbents of the Penis . 19/ the middle hsemorrhoidal artery have also been found there, but this does not often occur. The Veins of the penis are of two kinds : those which originate in the corpora cavernosa, accompany the corresponding branches of the pudic artery, but communicate more or less with the plexus of veins on the lower and lateral part of the bladder. There is also a great vein, which occupies the groove on the back of the penis, between the corpora caver- nosa, that appears particularly appropriated to the corpus spongiosum urethrae ; for it originates in the glans penis, and receives branches from the urethra as it proceeds backwards. There are often two of these veins, one in the groove, and the other more superficial : they generally unite near the root of the penis. The common trunk then passes between the body of the penis and the symphisis pubis, and ter- minates in a plexus of veins at the neck of the blad- der, which is connected to the plexus above men- tioned on the lower and lateral parts of the same viscus. The Absorbent Vessels of the penis take two differ- ent directions on each side. Those which arise from the integuments generally, unite so as to form a few trunks on the back of the penis, which divide near the root of the organ, and proceed to the glands of the groin. Those which originate from the interior parts of the penis, accompany the blood vessels, and terminate in the plexus of lymphatics in the pelvis. It ought to be noted, that the superficial lympha- tics generally enter the upper inguinal glands. The JYerves of the penis are principally derived from the lower secral nerves, which unite in the plexus that forms the great ischiatic. From these nerves a branch on each side originates, which passes like the pudic artery, between the sacro-sciatic liga- ments. In this course it divides into two branches, one 198 JVerves. -^General Observations. of which passes below to the museles of the penis and urethra, and to the contiguous parts ; and some of its branches seem finally to terminate in the dartos : the other branch proceeds along the crura of the pu- bis and ischium, and passing between the symphisis pubis and the body of the penis, arrives at the upper surface or dorsum of the penis, along which it conti- nues on the outside of the veins to the glans, in which it terminates. In this course it sends off several branches, some of which terminate in the integuments of the penis. After an examination of the relative situation of the mus- cles and blood vessels of the male organs of genaration, there appears reason to doubt, whether the erection of the penis can be referred to pressure upon the veins which return from that organ. Albinus has written on this subject. See Academicarum Annotationum, lib. ii. caput xviii. Haller has also considered it, and stated the opinions of several anatomists, in bis Ele- menta Physiologiae, tom. vii. page 555. The manner in which the urine is confined in the blad- der does not appear to be clearly understood. The connexion of the neck of the bladder with the prostate, and the appearance of the contiguous parts of the blad- der, do not render it probable that these parts act like a sphincter. The late J. Hunter, who paid great at- tention to the functions of these organs, was very de- cided in his opinion that the contraction of the urethra I iroduced the effect of a sphincter of the bladder. He las published some very ingenious observations respect- ing the manner in which urine is discharged from the bladder, in his Treatise on the Venereal Disease, part III. chapter IX. Mr. Hunter also long since asserted, that the vascular convoluted appearance of the corpus spongiosum ure- thrae was more distinct in the horse than the man. In the fifth volume of the Lemons d’Anatomie Comparee of Cuvier, the very learned and ingenious author con- firms the declaration of Hunter, respecting the vascu- lar convolutions of the corpus spongiosum of the horse. He states, that the corpora cavernosa of the penis of the elephant appear to be filled in a great degree with General Observations . 199 the ramifications of veins, which communicate with each other by such large and frequent anastomoses, that they have a cellular appearance. A similar structure exists in the horse, camel, bullock, deer, &c. and in them all these communicating branches can be distinguished from those which extend the whole length of the penis. The corpus spongiosum urethrae, according to M. Cu- vier, is constructed in a similar manner. From these facts he is induced to believe that this structure per- vades the whole class of mammalia. CHAPTER VL OF THE FEMALE ORGANS OF GENERATION, The female organs of generation consist of the Uterus and Ovaries , with their appendages ; and of the Vagina , with the structure which surrounds its external orifice. The uterus is situated in the pelvis, between the bladder and rectum ; and the ovaries are on each side of it. The vagina is a very large membranous canal, which passes from the uterus downwards and forwards, also between the bladder and rectum, and opens externally. Connected with the orifice of the vagina are seve- ral bodies, which are called the external parts of generation , in order to distinguish them from the uterus and ovaries, and their appendages, and also from the canal of the vagina ; which are called the internal parts. The bladder of urine lies above and in contact with the vagina : the urethra is also intimately connected with it. The description of the bladder and urethra is therefore placed at the end of this chapter. SECTION I. Of the External Parts of Generation. The adipose membrane, immediately anterior to the symphisis pubis, and on each side of it, forms a considerable prominence in females, which, at the age of puberty, is covered with hair, as in males. This prominence is denominated the Mons Veneris. The exterior orifice commences immediately be- low this. On. each side of this orifice is a nronfib Labia Externa. — Clitoris. 2gi uence continued from the mons veneris, which is largest above, and gradually diminishes as it descends. These prominences have some hair upon them. They are called the Labia Externa. Their junction be low is denominated the Fourchette. The space be- tween the place of their junction and the anus is ra- ther more than an inch in extent, and is denominated the Perineum. As the skin which forms the labia is continued in- ternally, it becomes more thin and soft, and is cover ed by a more delicate cuticle. It is also more or less florid, and secretes a peculiar mucous. In the upper angle, formed by the labia externa, is the upper extremity and glans of the clitoris. The Clitoris is a body which has a very strong resemblance to the penis, but there is no urethra at taehed to it. It has two crura of considerable length, which originate, like those of the penis, from the crura of the pubis and ischium, and unite at the symphisis of the pubis so as to form a body, which is not much more than an inch in length, and is broad in proportion. The extremity of this organ, called the Gians of the Clitoris , forms a small tu- bercle, which is covered above and on the sides by a small plait or fold of the skin, denominated the Pre- puce. These parts are lubricated by a secretion similar to that which is observed round the glans penis. The crura of the clitoris have muscles similar to the erectores penis. The interior structure of the Clitoris is very similar to that of the corpora caver- nosa of the penis, or the corpus spongiosum of the urethra. It appears constructed for a similar disten- tion, and is endued with the same sensibility as the penis. The two lateral parts are also separated from each other by a septum, resembling that of the penis. It is united to the symphisis pubis by a ligament Vol.'h, " 26 ^02 JVyniphse .* — Hymen , The Prepuce of the clitoris lias a semicircular form : below its extremities two folds or plaits com mence, one on each side, which are situated obliquely with respect to each other, so as to form an angle. These folds are denominated the Nymphsc. The Nymphse extend from the clitoris downwards nearly as far as the middle of the orifice of the vagina. They are situated within the external labia, and are formed by the skin after it has become more delicate in its texture. Their surface however.is often some- what corrugated. There are many blood vessels in their internal structure, and it is supposed they are occasionally somewhat tumid. They are flat, and their exterior edge is convex; so that they are nar- row at their extremities, and broad in the middle. Their breadth is very variable, and in some instances is great. In a majority of cases, it is equal to one- fourth of their length. Their colour in young sub- jects is of a bright red; in women advanced in years and who have had many children, they are of a brown red, and sometimes of a dark colour. The use of these parts is not very evident. They have been supposed to regulate the course of the urine as it flows from the urethra, but their effect in tliis respect is not great. They have also been sup- posed to favour the necessary enlargement of the parts in parturition. The orifice of the urethra is situated about an inch and one quarter further inward than the clitoris. It A is often rathe® less than the diameter of the urethra, and is somewhat protuberant. The orifices of mu- cous ducts arq to be perceived around it. The orifice of the urethra is at the commencement of the Canal of the vagina. Immediately within this orifice is situated the membrane denominated f Hymen . The Hymen is an incomplete septum, made by a * 203 The Vagina. fold op duplicature of the membrane which forms the surface contiguous to it. Sometimes it is circu- lar, with an aperture in the centre. Sometimes it has a resemblance to the crescent, the aperture be- ing at the upper part of it. The hymen has fre- quently been found without a perforation, and has therefore prevented the discharge of the menstrual evacuation. It is generally ruptured in the first in- tercourse of the sexes ; and some small tubercles, which are found on the surface of the vagina near the spot where it was situated, are supposed to be the remains of it. These tubercles are called Car unci din Mi/rtiformes. ' SECTION 11 . Of the Vagina. The canal of the vagina, commencing at the hy- men and the orifice of the urethra, is rather more narrow at its beginning than it is further inward.— From this place it extends backwards and upwards, and partakes in a small degree of the curve of the rectum : while the bladder, which is above it, and rests upon it, increases the curvature of the anterior part. It is much larger in women who have had children than in those who have not. The membrane which lines the vagina resembles to a certain degree, the membranes which secrete mucous in different parts of the body. Its surface appears to consist of very small papillae ; and at the anterior extremity of the vagina it forms a great number of rugEe, which are arranged in a transverse direction, both on the part of the vagina connected to the bladder, and on that part which is connected to the rectum, while the lateral parts of the vagina are 204 Corpora Cavernosa . — Sphincter Vaginx. smooth. These rugae are most prominent in the mid- tile ; so that a raised line appears to pass through them at right angles. This line extends from with- out inwards. The rugae on the part next to the blad- der are the strongest. This arrangement of the surface of the vagina does not extend beyond the external half of the canal : on the internal half part, or that nearest the uterus, the surface is smooth. The rugae are considerably diminished in women who have had children. Throughout this surface are to be seen, in some cases with the naked eye, the orifices of mucous fol- licles or ducts, which occasionally discharge conside- rable quantities of mucous. Exterior to this lining membrane of the vagina is a dense cellular structure, which has not yet been completely investigated : it is of a lightish colour, and has some resemblance to the texture of the body of the uterus. It is very vascular, and appears to be of a fibrous structure. It may be very much distended, and seems to have a contractile power. At the anterior extremity of the vagina, on each side of it, there is superadded to this, a cellular, or vascular substance, from eight lines to an inch in breadth ; which, when cut into, resembles the cor- pora cavernosa, or the corpus spongiosum of the pe- nis. These bodies commence near the body of the clitoris, and extend downwards on each side of the vagina. They have been called Plexus Eeteformis, and Corpora Cavernosa Vaginx , and are supposed to be occasionally distended with blood, like the cli- toris and penis. These corpora eavernosa are covered by muscular fibres, which pass over them on each side from the sphincter ani to the body of the clitoris ; to each of which organs they are attached. These fibres con- Unimpregnated Uterus , 205 stitute the sphincter vagina muscle, and contract the diameter of the vagina at the place where they are situated. The transversus perinei muscles also exist in the female. They pass from the tuberosities of the ischia, and are* inserted into a dense whitish substance in the perineum, to which the anterior extremity of the sphincter ani is likewise attached. The vagina is in contact with the rectum behind 5 the bladder lies upon it and anterior to it. A small portion of peritoneum, to be reflected to the rectum, is continued from the uterus upon the posterior part • of it. TJie lateral portions of it are invested with eellular substance. The anterior extremity of the uterus, which is called the Os Tinea}, projects into it from above. * SECTION III. Of the Uterus , the Ovaries and their Appendages, The Uterus Has been compared to a pear with a long neck.— There is of course a considerable difference between the body and neck ; the first being twice as broad as the last. Each of these parts is somewhat flattened,* In subjects of mature age, who have never been pregnant, the whole of the uterus is about two inches and a half in length, and more than one inch and a half in breadth at the broadest part of the body : it is also near an inch in thickness. .It is generally larger than this in women who have lately had children. The uterus is situated in the pelvis between the bladder and rectum, and is enclosed in a duplicature or fold of the peritoneum, which forms a loose septum that extends from one side of the pelvis to the other, ^ and divides it into an anterior and posterior cham 2(>o Ligaments of the Uterus. her. The posterior surface of this septum is oppo- sed to the rectum, and the anterior to the bladder. The two portions of this septum, which are between the uterus and the lateral parts of the pelvis, are called the Broad Ligaments. On the. posterior surface, the Ovaries are, situated on each side of the uterus, being enclosed by a pro- ct&&of the ligament or septum. Above them, in the upper edge of the septum, are the Fallopian Tubes , which are ducts that commence at the upper part of the uterus on each side, and proceed, in a lateral di~ : *iion for some distance, when they form an angle and incline downwards to the ovaries. These ducts are enclosed between the two lamina of the septum for the greater part of their length. The peritoneum, which forms the septum, is re- flected from it, posteriorly, to the rectum and the posterior surface of the pelvis, and anteriorly, to the bladder. In its progress, in each direction, it forms small plaits or folds ; two of which extend from the uterus to the rectum posteriorly, and two more to the bladder anteriorly : these are called the Anterior and Posterior Ligaments of the Uterus. The other ligaments, which proceed more imme- diately from the uterus, are called the Bound Liga- ments. These arise from each side of the uterus, at a small distance before and below the origins of the fallopian tubes, and proceed in an oblique course to the abdominal rings. These ligaments are also in- vested by the peritoneum. They pass through the rings and soon terminate. In the body of the uterus is a cavity, which ap- proaches to the triangular form ; and from which a canal proceeds through its neck. This cavity is so small that its sides are almost in contact, and the ca- nal is in proportion ; so that this organ is very thick in proportion to its bulk. Structure and Cavity of the Uterus. 20 7 The substance of which the uterus consists is very firm and dense : it is of a whitish colour, with a slight tinge of red. There are many blood vessels, with nerves and absorbent vessels, in its texture. The nature and structure of this substance has not yet been precisely ascertained. It appears very different indeed from muscle ; but the uterus occasionally con- tracts, with great force, during labour. It is not ren- dered thin by its enlargement during pregnancy, and the blood vessels in its texture are greatly enlarged at that time. Exteriorly, the uterus is covered by the peritone- um, as ha;? already been mentioned. Internally it is lined with a delicate membrane that has. some resem- blance to those which secrete mucous, and is gener- ally of a whitish colour, abounding with small orifi- ces that can be seen with a magnifying glass. This membrane is so intimately connected to the substance of the uterus that some anatomists have supposed it was merely the internal surface of that substance, but this opinion is now generally abandoned. It is sup- posed that the colour of this membrane is more florid about the period of menstruation. The cavity of the uterus, as has been observed be- fore, is triangular in form. When the organ is in its natural position, the upper side of this triangle is transverse with respect to the body, and the other * sides pass downwards and inwards. In each of the upper angles are the orifices of the fallopian tubes, which are of such size as to admit a hog’s bristle. The two lower lines of the triangle are slightly curved outwards at their upper extremities ; so that the upper angles of the triangle project outwards,, and the orifices of the fallopian tubes are nearer to the external surface than they otherwise would be. ’ The lower angle of the cavity of the uterus is oc-' 208 Structure and Cavity of the Uterus . cupied by the orifice of the canal, which passes through the neck of the organ ; this orifice is from three to four lines in diameter. The canal is about an inch, in length, and is rather wider in the middle than at either end. On the anterior and posterior portions of its surface are many small ridges which have an arborescent arrangement, one large ridge passing internally from the commencement of the canal, from which a number of other ridges go off in a transverse direction. These ridges extend nearly the whole length of the canal. In the grooves, be- tween the ridges, are the orifices of many mucous ducts. There are also on this surface a tnumber of transparent bodies of a round form, equal in bulk to a middle sized grain of sand the nature and use of which is unknown. They have been called Qvida JYabothi, after a physiologist, who published some speculations respecting their use, about the com- mencement of the last century. The canal of the neck of the uterus is very dif- ferent from other ducts, for it seems to he a part of the cavity to which it leads, and when the cavity of the uterus becomes enlarged in the progress of preg- nancy, this canal is gradually converted into a part of that cavity. The lower extremity of the neck of the uterus is irreglarly convex and tumid. The orifice of the canal in it is oval, and so situated that it divides the convex surface of the neck into two portions, which are called the Lips. The anterior or upper portion is thicker than the other. This extremity of the uterus protrudes into the va- gina, and is commonly called Os Tincse . As the an- terior portion or lip is larger and more tumid than ‘the posterior, the vagina extends further beyond the os tincse on the posterior part than on the anterior., Fallopian Tubes. —Round Ligaments 209 The Fallopian Tubes Are two canals, from four to five inches in length? which proceed between the lamina of the broad liga- ments, from the upper angles of the uterus, in a transverse direction, to some distance from the ute- rus, when they form an angle, and take a direction downwards towards the ovaries. They are formed, for a considerable part of their extent, by a substance which resembles that of which the uterus consists, and are lined by a membrane continued from the internal membrane of the uterus* Their extremities appear to be composed of mem- brane, which is rendered florid by the blood vessels in its texture* At the commencement their diameters are extremely small ; but they enlarge in their pro- gress. This enlargement is gradual for the first half, and afterwards sudden ; the enlarged part is more membranous than the small part, and has a bright red colour. The large extremity is loose in the ca- vity of the pelvis, and is not invested by the lamina of the broad ligaments. Near the termination the diameter is often contracted ; after which the mem- brane which forms the tube expands into an open mouth, the margin of which consists of fringed pro- cesses : this margin is also oblique, as respects the axis of the tube ; and the different fringed processes are not all of the same! length ; but the longest are in the middle, and the others regularly diminish on each side of them : these processes constitute the Fimbriae of the fallopian tubes. The internal surface of the large extremities of these tubes is extremely vascular ; and there are some longitudinal fibres of a red colour to be seen on it. The Round Ligaments , Which have already been mentioned, are cords of Vjpffc. n, 27 21 U * Structure of the Ovaries . a fibrous structure, with many blood vessels in them. They arise from the uterus below the origin of the fallopian tubes, and proceed under the anterior la- mina of the broad ligaments to the abdominal rings, through which they pass; and then the fibres and vessels are expanded upon the contiguous cellular I substance. The Ovaries Are two bodies of a flattened oval form ; one of which is situated on each side of the uterus on the posterior surface of the broad ligament, and invested completely by a process of the posterior fa men, which forms a coat, and also a ligament for it. The size of this organ varies in different subjects, but in a majority of those who are about the age of matu- rity it is between ten and twelve lines in length. It is connected to the uterus by a small ligament, or bundle of fibres of the same structure with the round ligaments, which is not more than two lines in dia- meter, and is included between the lamina of the broad ligament. The process of the broad - ligament forms an exter- nal coat to the ovary ; within this is the proper coat of the organ, which is a firm membrane. This mem- brane is so firmly connected to the substance of the ovary which it encloses, that it cannot be easily se- parated from it. The ovary is of a whitish colour and soft texture, and has many blood vessels. Ia virgins of mature age it contains from fen to twenty vesicles, formed of a delicate membrane, filled with a transparent coagulable fluid. Some of these vesi- cles are situated so near to the surface of the ovary that they are prominent on its surface; others are near the centre. They are very different in size ; the largest being between two and three lines ir Arteries of the Uterus „ 21 1 diameter, and others not more than one-third of that size. In women who have had children, or in whom conception has taken place, some of these vesicles are removed ; and in their place a cicatrix is found. It has been ascertained, that during the sexual intercourse with males, one of these vesicles, which was protuberant on the surface, is often ruptured, and a cavity is found. A cicatrix -is soon formed, where the membrane was ruptured ; and in the place occupied by the vesicle there is a yellow substance denominated Coi'pus Luteum. This corpus luteum generally continues until the middle of pregnancy: it often- remains during that state, and for some time after delivery, but it gradually vanishes. The cica- trization continues during life. In many cases these cicatrices correspond with the number of conceptions which have taken place ; but they often, exceed the number of conceptions, and they have been found in cases where conception has not been known to have taken place. In very old subjects, where conception has never taken place, the vesicles are either entirely removed, or small dense tubercles only remain in their place. The Arteries Of the uterus are derived from two very different sources ; viz. from the spermatic and from the hy- pogastric arteries. The spermatic arteries, instead of passing directly down to the abdominal ring, proceed between the lamina of the broad ligament, and send branches to the ovaries, which may sometimes be traced to the vesicles. They also send branches to the fallopian tubes and to the uterus. Those which are on the opposite sides of the uterus anastomose with each other, and also with the branches of the hypogastric 3X2 .Veins of the Uterus . arteries. There are also branches of these arteries in the round ligaments which accompany them to their termination outside of the abdominal ring. The principal arteries of the uterus are those de- rived from the hypogastric, which sends to each side of it a considerable branch, called the Uterine. This vessel leaves the hypogastric very near the origin of the internal pudic, and proceeds to the cervix of the uterus : it passes between the lamina of the broad li- gaments, and sends branches to the edge of the ute- rus, which penetrate its texture. The branches which are in the texture of the uterus, are very small in- deed. in young subjects. In women who have had children they are considerably larger; but during pregnancy they gradually enlarge with the growth of the uterus, and become very considerable. These arteries observe a serpentine and peculiarly tortu- ous course. Those on the opposite sides anastomose with each other. The Veins Of the uterus, like the arteries, form spermatic and uterine trunks. The Spermatic Vein is much larger than the artery. It ramifies as in males, and forms a very large plexus, w hich constitutes the cor- pus pampiniforme. Many of the veins which form this body, originate near the ovary: a considerable jjumber also come from the fallopian tubes and the uterus. The spermatic vein and its branches are greatly enlarged indeed during pregnancy ; and it is said that they are enlarged the same way during the menstrual discharge. The most imporant veins of the uterus are the branches of the Uterine Veins. They are extremely numerous, and form a plexus on the side of the ute- rus ; from which two or more uterine veins proceed in the course of the artery, and join the hypogastric Lymphatics. —Bladder and Urethra. 213 These veins also are greatly enlarged during preg- nancy. The Lymphatic Vessels Of the uterus, and its appendages, are very nu- merous. In the unimpregnated state they are small ; but, during pregnancy, they increase greatly. They proceed from the uterus in very different directions. Some that accompany the round ligaments go to the lymphatic glands of the groin. Others which take the course of the uterine blood vessels pass to glands in the pelvis, and a third set follows the spermatic arteries and veins to the glands on the loins. The JYerves Of the ovaries are derived from the renal plexus, and those of the uterus and vagina from the hypo- gastric plexus, or the lower portions of the sympa- thetic, and the third and fourth sacral nerves, SECTION IV. Of the Bladder and Urethra. The situation of the Bladder , as respects the sym phisis pubis, is nearly alike in both sexes ; but that part of it which is immediately behind the insertion of the ureters is rather lower in males than in fe- males. The bottom of the bladder rests upon the upper part of the vagina, a thin stratum of cellular substance only intervening: when that viscus is dis- tended it forms a tumour, which compresses the va- gina. The ureters are inserted, and the urethra commen- ces in the same part of the bladder, in both sexes. The length of the Urethra is between one and two inches. When the body is in a direct position, it is nearly horizontal; but it is slightly curved? with 314 General Change in (he Gravid Uterus. its convexity downwards. It is immediately above the vagina, and it passes below the body of the cli- toris. The external orifice of it is rather more than an inch within the gians or head of the clitoris. This orifice is somewhat prominent in the vagina. In the internal or lining membrane of the urethra there are many orifices of mucous follicles, and also longitudinal wrinkles, as in the urethra of males. The diameter of the female urethra and its orifice in the bladder are greater than they are in the male. For this reason it has been supposed, that women are less liable to calculus of the bladder than men.* • The urethra is intimately connected with the ex- ternal coat of the vagina, and between them there is a spongy cellular substance which makes the fough surface of the vagina prominent ; so that the urethra has been supposed, although erroneously, to be in- vested with the prostate. It is capable of great ar- tificial dilatation. - Of the Changes induced in the Uterus in the pro- gress of Pregnancy . The alteration which takes place in the size of the uterus during pregnancy is truly great. About the conclusion of that period, instead of the small body above described, which is almost solid, the uterus forms an immense sac, which extends from the ter- mination of the vagina in the pelvis, into the epigas- tric region ; and from one jside of the abdomen to the other ; preserving, however an ovoid figure. This change is so gradual at first, that the uterus does not extend beyond the cavity of the pelvis be- fore the third month, although at the end of the se- venth month it is very near the epigastric region. For the first six months the body of the uterus * It has however been asserted that they are also less liable to calculi m the kidneys. General Observations, 215 appears principally concerned in the enlargement : after this the cervix begins to change, and is gradu- ally altered so as to compose a portion of the sac, rather of less thicknes than the rest of the uterus ; the mouth being ultimately an aperture in a part which is much thinner than the other portions of the organ. The change which takes place in the texture of some of the appendages of the uterus is very import- ant. The Broad Ligaments , which seem particularly calculated to favour the extension of the uterus, are necessarily altered by the change in the ’size of that organ, but not entirely done away. The portion of peritoneum of which they are formed must be very much enlarged with the growth of the uterus, as it continues to cover it. The Round Ligaments are much elongated ; and they observe a more straight course to the abdominal ring. The Fallopian Tubes are enlarged ; and instead of passing off laterally from the uterus, they now proceed downwards by the side of it. The Ovaries appear rather larger and more spongy : their relative situation is necessa- rily lower. The change in the Uterus itself is particularly in- teresting. The great increase of its size is not at- tended with any considerable diminution of thick- ness in its substance ; nor are the arteries much less convoluted than before pregnancy , as might have been expected. They are greatly enlarged in diam eter, and the orifices of the exhalent vessels on the internal surface of the uterus are much more percep- tible. The veins are much more enlarged than the arte- ries, and in some places appear more than half an inch in diameter. They are not regularly cylindri- 216 Peculiarity of the Female Hottentots. cal, but rather flat. They anastomose so as to form an irregular net work. The uterus appears much more fibrous and mus- cular in the gravid than in the unimpregnated state. The contractile power of the gravid uterus is not only proved by the expulsion of its contents, but also by very vigorous contractions, which are occasionally observed by accoucheurs. Although the general effects which result from the par- ticular conditions of the uterus in pregnancy, men- struation, &c. evince that the influence of this organ upon the whole system is very great, yet it seems pro- bable'that the sexual peculiarities of females are espe- cially dependent upon the ovaria. This sentiment is confirmed by an account of a wo- man in whom the ovaria were deficient, which is pub- lished in the London Philosophical Transactions for 1805, by Mr. C. Pears. The subject lived to the age of twenty-nine years. She ceased to grow after the age of ten years, and therefore was hot more than four feet six inches in height : her breadth across the hips was but nine inches, although the breadth of tne shoulders was fourteen. Her breasts and nipples ne- ver enlarged more than they are in the male subject. There was no hair on the pubes, nor were there any indications of puberty in mind or body. She never menstruated. At the age of twenty -nine she died of a complaint in the breast, attended with convulsions. — The uterus and os tine® were found not increased beyond their usual size during infancy. The cavity of the uterus was of the common shape, but its coats were membranous. The Fallopian Tubes were per- vious. “ The Ovaria were, so indistinct that they ra- ther showed the rudiments which ought to have form- ed them , than any part of the natural structure .V Another case, which confirms the aforesaid sentiment, is related in one of the French periodical publications.. It has been long known that a race of savages near the Cape of Good Hope were distinguished trom the generality of their Species by a* peculiarity about the pudendum. An account of this structure has been Peculiarities in the Abdomen of the Fcetus 217" given with some precision by Messrs. Peron and Le- sueur, in a paper which was read to the National Insti- tute of France. It is a flap or apron, four inches in length, which is united to the external labia near their upper angle, and hangs down before the clitoris and the external orifice of the parts of generation. It is divided below into two lobes, which covers the orifice. It is formed by a soft distensible skin, free from hair, which is occasionally corrugated like the scrotum, and is rather more florid than the ordinary cutis. s The Abdomen of the Fcetus. The difference between the foetus and the adult, in the cavity of the abdomen, is very conspicuous at the first view. The Liver in the foetus is so large that it occupies a very considerable part of the abdomen. Its left lobe, which is larger in proportion than the right, extends far into the left hypochondriac region. The Bladder of urine , when filled, extends from the cavity of the pelvis a considerable distance to- wards the umbilicus : so that the greatest part of it is in the cavity of the abdomen. A ligament of a conical figure extends from the centre of the upper part of the bladder to the umbilicus ; with an artery on each side of it, which is soon to be described. This ligament, which is in the situation of the ura- chus of the fcetus of quadrupeds, is hollow, and thus frequently forms a canal, which has a very small diameter, that communicates with the bladder by an aperture still smaller, and continues a short distance from the bladder towards the umbilicus. In a few rare instances this canal has extended to the umbili- cus, so that urine has been discharged through it, but the ligament is commonly solid there. * This paper has not yet been published by the Institute, but it is refer- red to by M. Cuvier in his Lemons d’Anatomie Comparee, voL v. page 124.— Messrs. Peron and Lesueur were naturalists who accompanied captain Baudin in his voyage of discover} - ; the latter has been for some years re- sident in Philadelphia, VOL-v IT, 18 218 Descent of the Testicle in the Foetus . The Stomach appears to be more curved in the foetus than in the adult. The Great Intestine does not extend sufficiently far, beyond the insertion of the ileon, to form the cfecum completely. The Glandalse JRenales are much larger in pro- portion in the foetus than in the adult. The colour of the fluid they contain is more florid. The Kidneys are lobulated. The Testicles in the foetus are found above the pelvis, in the lumbar region, behind the peritoneum, until two months before birth. Thus situated, their blood vessels and nerves proceed from sources which are near them ; but the vas deferens, being connected to the vesiculac seminales by one extremity, is neces- sarily in a very different situation from what it is in the adult: it proceeds from the testicle downwards to the neck of the bladder. — While each testicle is in this situation, it is connected with a substance or liga- ment, called Gubernaculwn , of a conical or pyrami- dical form, attached to its lower end, and extends from it to the abdominal ring. This substance is vascular, and of a fibrous texture: its large extremity adheres to the testicle, its lower and small extremity passes through the abdominal ring, and appears to terminate in the cellular substance exterior to that opening, like the round ligament in females. The Gubernaculum, as well as the testicle, is behind the peritoneum ; and the peritoneum adheres to each of them more firmly than it does to any of the surround- ing parts. It seems that, by the contraction of the Gubernaculum , the testicle is moved down from its original situation to the abdominal ring, and through the abdominal ring into the scrotum. The perito- neum, which adheres firmly to the gubernaculum and testicle, and is loosely connected to the other parts, yields to this operation ; and \yhen the testicle has Descent of the Testicle in the Fcetus . 219 arrived near the abdominal ring, a portion of the pe- ritoneum is protruded a little way before it into the scrotum ; forming a cavity like the finger of a glove. The testicle passes down behind this process of the peritoneum, and is covered by it as it was in the ab- domen. Although it appears protruded into the ca- vity, it is exterior to it, and behind it ; and the ves- sels, &c. which belong to the testicle are also ex- terior to it. The cavity formed in the scrotum, by this process of the peritoneum, necessarily communicates with the cavity of the abdomen at its formation ; but very soon after the testicle has descended into the scrotum, the upper part of this cavity is closed up, while the low- er part of the process continues unchanged, and con- stitutes the Tunica Vaginalis Testis. In some in- stances the upper part of this process does not close up, and the communication with the cavity of the abdomen continues. The descent of the intestine into the cavity thus circumstanced, constitutes that species of Hernia which is denominated Congenital.* The most important peculiarities in the abdomen of the foetus are those connected with the circulation of the blood. The internal iliac or hypogastric arteries are larger than the external iliacs. Their main trunks are * These interesting circumstances respecting the original situation of the testicle, and its descent into the scrotum, were discovered and elu- cidated by Haller, Hunter, Pott, Camper, and several other very respect- able anatomists and surgeons. There is, however, a difference of opin- ion, between some of them, as to the time when the testicle leaves the abdomen. Haller thought the testicles were seldom in the scrotum at birth. Hunter and Camper found them so generally. It has been suggested that there are some national peculiarities in this re- spect ; that amongst the Hungarians, for example, the testicles often re- main above the abdominal ring until near the age of puberty. The student will find an interesting description of the situation of the testis, and its descent, in the foetus, in the “ Observations o.n certain parts of the Animal Economy, 51 by John Hunter, Umbilical Vessels. ;i2U continued on each side of the bladder to its fundus t and proceed from it, with the ligament, to the umbi- licus ; when they pass out of the abdomen to go along the umbilical cord to the placenta. These arteries are now denominated the Umbilical, and are very considerable in size. After birth, as there is no cir- culation in them, they soon begin to change : the ca- vity of them is gradually obliterated, and they are converted into ligaments. They are exterior to the peritoneum, and contained in a duplicature of it. A vein also called the Umbilical, which is much larger in diameter than both of the arteries, returns from the placenta along the cord, and enters the ca- vity of the abdomen at the umbilicus. It proceeds thence, exterior to the peritoneum, but in a duplica- ture of it called the Falciform Ligament , to the liver, and enters that viscus at the great fissure ; along which it passes to the left branch of the sinus of the vena portarum, into which it opens and dis- charges the blood which flows through it from the placenta. It opens on the anterior side of the branch of the vena portarum, and from the posterior side of the branch, opposite to this opening, proceeds a duct or canal, which opens into the left hepatic vein near its junction with the vena cava. This communicating vessel is called the Ductus, or Canalis Venosus ; to distinguish it from the duct which passes from the pulmonary artery to the aorta, and is called Ductus , or Canalis Arteriosus. This venous duct carries some of the blood of the umbilical vein directly to the vena cava ; but it is much smaller than the um- bilical vein, and of course a considerable quantity of the blood which passes through the umbilical vein must pass through the liver, by the vena portarum, before it can enter the cava. In some foetal subjects, if a probe of sufficient Object of the Circulation in the Placenta. 221 length be introduced within the umbilical vein and pushed forwards, it will pass to the heart without much difficulty or opposition, as if it proceeded along one continued tube, although it really passes from the umbilical vein across the branch of the vena por- tarum, and then through the ductus venosus, and through a portion of the left hepatic vein, into the inferior vena cava. If the umbilical vein be injected with a composi- tion, which will be firm when cool, it appears to terminate in a rounded end, which is situated in the transverse fissure of the liver : the sinus of the vena portarum, into which this vein enters, appears like two branches going off, one from each side of it, and the ductus venosus like a branch continuing in the direction of the main trunk of the umbilical vein. The umbilical vein, in its progress through the fissure of the liver, before it arrives at the sinus of the vena portarum, sends off a considerable number of branches to each of the lobes of that organ, but more to the left than to the right lobe. After birth, when blood ceases to flow through the umbilical vein, it is gradually converted into a liga- ment ; and the venous duet is also converted into a ligament in the same manner. The vena portarum, which before appeared very small, when compared with the umbilical vein, now brings all the blood which fills its great sinus, and increases considerably in size. It has been ascertained by anatomical investiga- tion, that the umbilical arteries above mentioned, after ramifying minutely in the placenta, communi- cate with the minute branches of the umbilical vein ; and it is probable that the whole blood carried to the placenta by these arteries, returns by the umbilical vein to the foetus. It is dearly proved by the effects of pressure on 222 Object of the Circulation in the Placenta. the umbilical cord, in cases of delivery by the feet, as well as by other similar circumstances* that this circulation cannot 'be suspended for any length of time without destroying the life of the foetus.' From these circumstances, and from the florid colour which the blood acquires by circulating in the pla- centa, it seems probable that the object of the circu- lation through that organ is somewhat analogous to the object of the pulmonary circulation through the lungs of adults.* * * During the first four months of pregnancy a very small vesicle, which does not exceed the size of apea, is found between the chorion and the amnios, near the insertion of the umbilical cord into the placenta. It is connected to the fetus by an artery and a vein, which pass from the abdomen through the umbilicus, and proceeding along the cord to the placenta, continue from it to the vesicle. The artery arises from the mesenteric, and the veii^ is united to the mesenteric branch of the vena portarum. It is proba- ble that these vessels commonly exist no longer than the vesicle, viz. about four months ; but they have been seen by Haller fujd-Chaussier at the termination of pregnancy. They are called Omphat&iiHesenteric -ves- sels. The vesicle is denominated the Umbilical Vesicle. This inexplicable structure is delineated in Hunter’s Anatomy of the Gravid Uterus, plate xxxiii. figures v. and vi. ; in the Academical Annota- tions of Albums, first book, plate i. figure xii.; and also in the leones Em- bryonum Humanortun of SoemmeriDg, figure & SYSTEM OF ANATOMY. PART IX. OF THE BLOOD VESSELS. The blood vessels are flexible tubes, of a peculiar texture, through which blood passes from the heart to the different parts of the body, and returns again from these parts to the heart. They are to be found, in varying proportions, in almost every part of the body, and seem to enter into its texture. The tubes which carry blood from the heart, are more substantial and more elastic than those through which it returns to the heart. They are generally found empty after death ; and, therefore, were called Arteries by the ancient anatomists, who supposed that they carried air, and not blood. The tubes which return the blood to the heart are denominated Veins . They are less substantial and less elastic than arteries, and are generally fall of blood in the dead subject. There are two great arteries, from which all the Other arterial vessels of the body are derived. They are very justly compared to the trunks of trees, and the smaller vessels to their branches. One of these great arteries, called the Aorta, carries blood to eve- ry part of the body. The other great vessel, called the Pulmonary Artery , carries blood exclusively to the lungs. S24 The Blood vessels in general . The veins which correspond to the branches of the Aorta, unite to each other, so as to form two great trunks that proceed to the heart. One of these trunks, coming from the superior parts of the body, is called the Superior, or Descending Vena Cava. The other, which comes from the lower parts of the body, is called the Inferior, or Ascending Vena Cava. The veins which correspond with the branches of the Pulmonary Artery, and return to the heart the blood of the lungs, are four in number : two of them proceeding from each lung. They are called Pul- monary Veins. In many of the veins there are valves, which pre- vent the blood they contain from moving towards the surface and extremities of the body, but allow it to pass towards the heart without impediment. From the construction of the cavities of the heart, and the position of the valves which are in them ; as well as the situation of the valves at the commence- ment of the great arteries, and the above mentioned valves of the veins, it is evident, that when the blood circulates, it must move from the heart, through the aorta and its branches, to the different parts of the body, and return from these parts through the venae cavae, to the heart ; that, when deposited in the heart by the venae cavae, it must proceed through the pul- monary artery to the lungs, and return from the lungs through the pulmonary veins to the heart, in order to pass again from that organ into the aorta. It is also certain, that the blood is forced from the heart into the arteries, by the contraction of the mus- cular fibres of which the heart is composed ; and that the blood vessels likewise perform a part in the cir- culation, they propelling the blood which is thus thrown into them : but their action appears to depend upon causes of a complex nature. CHAPTER I. OF THE GENERAL STRUCTURE AND ARRANGEMENT OF THE BLOOD VESSELS, SECTION r. Of the Arteries. The arteries are so much concerned in the im- portant function of the circulation of the blood, that every circumstance connected with them is very in- teresting. They are composed of coats or tunics, which are very elastic and strong, and which are also very thick. In consequence of the firmness of their coats, they continue open, after their contents are discharged, like hard tubes. They submit to great dilatation, and elongation, when fluids are forced into them, and return to their former dimensions when the distend- ing cause is withdrawn. This elasticity is particu- larly subservient to the circulation of the blood. ' It admits the artery to distend readily, and receive the biood which is thrown into it by the contraction of the heart. It also produces the contraction of the artery; which takes place as soon as the action of the heart ceases ; and this contraction of the artery neces- sarily forces the blood forward, as the valves at its orifice prevent it from returning to the heart. The motion of the artery, which is so easily per- ceived by the touch, and in many instances also by the eye, is completely explained by the discharge of blood into the arterv from the heart, and by the VOL.il. 29 ' 226 Structure of the Arteries. elasticity of the vessel, by which it re-acts upon the blood. In some cases it is not simply the diameter of the artery which is enlarged, but a portion of the vessel is elongated; and this elongation, by pro- ducing a curvature of it, renders its motion more visible. In the aorta, and probably in its large branches, Elasticity seems to be the principal cause of the con- tinuance of the motion which is originally given to the blood by the heart. But there are many circum- stances connected with the smaller vessels, which evinee that they exert a power which is very differ- ent indeed from elasticity. Thus the application of local stimulants or rubefacients, and of heat, is fol- lowed by an increase of motion in the arteries of the parts to which they are applied. Neither of these causes could produce their effect by the influence of elasticity : but the effect of these and other similar causes is uniformly produced ; and a power of inde- pendent motion, or Irritability , is thus proved to exist in these vessels, and seems essentially necessa- ry to the circulation of the blood. The Structure of the Arteries Is, therefore, a subject of importance; and has re- ceived a considerable degree of attention from anato- mists. They are composed of a dense elastic substance, of a whitish colour. Their external surface is rough, and intimately connected with the cellular membrane, which every where surrounds it in varying quanti- ties. Internally, they are lined with a thin mem- brane, which is very smooth and flexible, and is also very elastic. The substance which composes the artery, and is situated between the cellular invest- ment and the internal membrane, consists of fibres, which are nearly, though not completely, circular, 227 / Structure of the Arteries . but so arranged as to constitute a cylinder. These fibres may be separated from each other so. as to form lamina, which have been considered as differ- ent coats of the arteries ,• but there is no arrange- ment of them which composes regular distinct strata. The coats of arteries may, therefore, be separated into a greater or smaller number of lamina, accord- ing to the thickness of these lamina. The fibres which compose these lamina appear to be united to each other in a way which readily al- lows of their separation, at the same time that they form a firm texture. Although arteries thus appear essentially different from muscles in their hardness and their elasticity, as well as in their general tex- ture, they are considered, by a great majority of anatomists, as partaking more or less of a muscular structure. In the human subject their structure is very diffi- cult of demonstration, and great differences exist in the accouuts which are given of it, even by anato- mists who agree in the general sentiment that the arteries are muscular. Thus Haller believed that muscular fibres were most abundant in the large arteries, while J. Hunter thought the reverse. Hunter appears to have investigated this subject with great attention, and supposed the muscular substance, in the composition of arteries, to be inte- rior, and the elastic matter exterior ; that in large arteries this muscular substance is very small in quantity, and gradually increases in proportion as the artery diminishes in size. He however observes, that he never could discover the direction of the muscular fibres* When the great talents of Mr. Hunter as an ' Treatise on the Blond, &c, Voi I, p. ITS. Bradford’s edition 228 Question? expecting the muscularity of Arteries . anatomist, are considered, this circumstance cannot fail to excite a belief that the existence of these ' fibres is not certain : and if to this be added the fact, that even the red coloured substance of the arteries is elastic, and in that respect different from muscu- lar substance, the reasons for doubting must be in- creased. Bichat appears to have entertained very strong doubts on the subject ; but he stands almost alone ; for a large number both of the preceding and cotem- porary anatomists, seem to have adopted the senti- ment, that the arteries have a muscular structure. The student of anatomy can very easily examine this subject himself, by separating the coats of arte- ries into different lamina ; and by viewing the edges of the transverse and longitudinal sections of those vessels. While thus engaged with this question, he will read with great advantage what has been writ- ten upon it by Mr. Hunter, in his Treatise on the Blood, &c. See chapter second, section 3. Bichat ought also to be read upon this subject, which he has discussed in his Anatomie Generate — -Systeme Vasculaire a Sang Rogue , article Troisieme, &c. and also in his Traite des Membranes , article Sixieine. The belief of the irritability of arteries does not however, rest upon the appearance of their fibres. 1. It is asserted by very respectable authors,* that they have been made to contract by the appli- cation of mechanical and of chemical irritation, and also of the electric and galvanic power. 2. A partial or local action of arteries is often produced by the local application of heat and rube- facients, as has been already observed. * See Soemmering on the structure of the Human Body, Vol. IV. Ger- man edition. Dr. Jones, on the Process employed by nature for suppressing Hemorrhage, &c. Proofs of the Irritability of the Jlrteries, 3. Arterial action is often suspended in a particu- lar part by the application of cold. It has also been observed that the arteries have for a short time ceas- ed to pulsate in cases of extreme contusion and lace- ration of the limbs.* 4. When arteries are divided transversely in living animals, they often contract so as to close completely the orifice made by the. division. 5. In a horse bled to death, it was ascertained by Mr. Hunter, that the transverse diameter of the ar- teries was diminished to a degree that could not be explained by their elasticity. He also found that, after death, the arteries, especially those of the smaller size, are generally in a state of contraction, which is greater than can be explained by their elas- ticity: for if they are distended mechanically, they do not contract again to their former size, but conti- nue of a larger diameter than they were before the distention; although their elasticity may act so as to restore a very considerable degree of the contraction observed at death. The contraction, which is thus done away by dis- tention, Mr. Hunter supposed to have been produced by muscular fibres : for, if it had been dependent on elasticity, it must have re-appeared when the distend- ing power was withdrawn. It therefore seems certain, that the arteries have a power of contraction 'different from that which de- pends upon elasticity ; but whether this depends upon muscular fibres superadded to them, or upon an irritable quality in the ordinary elastic fibres of blood vessels, is a question which is not perhaps completely decided. * This local suspension of arterial motion by cold, &c. applied locally, is very difficult to explain; as the' action of the heart and the elasticity of the arteries appear sufficient to account for the pulsation of the large ar- teries. 230 General Observations on ■ the Arteries. The motion of the blood in the arteries appears to depend, 1st, Upon the impulse given to it by the action of the heart. 2dly, Upon the elasticity of the arteries, in conse- quence of which they first give way to the blood impelled into them, and then re-act upon it; and 3dly, Upon the power of contraction in the arte- ries, or their irritability. In the larger arteries the blood seems to move as it would through an inanimate elastic tube, in conse- quence of the impulse given by the heart, and kept up by the arteries themselves. In the smaller ves- sels it seems probable that the motion of the blood depends in a considerable degree upon the contrac- tion which arises from their irritability. The obvious effect of the elasticity of the arteries is to resist distention and elongation, and to contract the artery to its natural state, when the distending or elongating cause ceases to act. But it must also resist the contraction induced by the muscular fibres, and restore the artery to its natural size when the muscular fibres cease to act after contracting it, as has been observed by Mr. Hunter. It seems probable that all the fibres of which the artery consists are nearly but not completely circular, for it is not certain that there are any longitudinal fibres in the structure of an artery. The internal coat of these vessels is very smooth, but extremely dense and firm; and seems to be ren- dered moist and flexible by an exudation on its sur- face. It adheres very closely to the contiguous fibres of the coat exterior to it, but may be very readily peeled off from them. It is of a whitish colour, and, like the fibrous structure of the artery, is very elastic. Like that substance also it is easily torn or broken, and when ligatures have been applied to General Observations on the Arteries . 231 arteries, it has often been observed that the fibrous structure and the internal coat have been separated while this external cellular coat has remained en- tire. The arteries are supplied with their proper blood vessels and lymphatics. It is to be observed, that the blood vessels are not derived from the artery on which they run, but from the contiguous vessels. These vessels have nerves also, which are rather small in size, when compared with those which go to other parts. Arteries appear to have a cylindrical form, for no .diminution of diameter is observable in those portions of them which send off* no ramifications. When an artery ramifies the area of the differ- ent branches exceeds considerably that of the main trunk. Upon this principle the aorta and its branches have been compared to a cone, the basis of which is formed by the branches, and the apex by the trunk.* The transverse section of an artery is circular. There are no valves in the arteries, except those of the orifices of the aorta and the pulmonary artery, at the heart. The valves of the pulmonary artery have been described in the 51st page of this volume, and those of the aorta have an exact resemblance to them, but are rather larger. The course of the arteries throughout the body is obviously calculated to prevent their exposure to pressure, or to great extension from the flexure of the articulations by which they pass. With this view they sometimes proceed in a winding direction ; and when they pass over parts which are subject to great distention or enlargement, as the cheeks, they often * According to Brassiere the relation of the branches of the aorta to its trunk is as 25 to 16; Helvetius reckons the orifice of the aorta in compa^ visori ■with its branches as 64 to 71. Lassus.— Es. I 232 General Observations on the Arteries, meander; and, therefore, their length may be in- creased by straightening, without stretching them. Their course appears sometimes to have been calculated to lessen the force of the blood, as is the case with the Internal Carotid and the Vertebral ar- teries. In the trunk of the body the branches of arteries generally form obtuse angles with the trunks from which they proceed. In the limbs these angles are acute. The communication of arteries with each other is termed Anastomosis. In some instances, two branches which proceed in a course nearly similar, unite, with an acute angle, and form one common trunk. — Sometimes, a transverse branch runs from one to the other, so as to form a right angle with each. In other cases, the two anastomosing branches form an arch, or portion of a circle, from which many branches go off. By successive ramifications, arteries gradually di- minish in size, until they are finally extremely small. The smallest arteries do not carry red blood, their diameters being smaller than those of the red parti- cles of that fluid, the serous or aqueous part of the blood can, therefore, only pass through them. Many of the arteries which carry red blood, and of the last mentioned serous arteries terminate in veins, which are in some respects, a continuation of the tube reflected backwards.# * Malpighi and Leeuwenhoek declare, that by the aid of a microscope, they have seen arteries terminating in the veins. Haller advances for- mally his own experience in support of his assertion. Other anatomists have seen, that in blowing into an artery, the air passed into the corres- ponding veins. Nevertheless, Duverney and some others say, that a particular substance is interposed between the extremities of these ves- sels. Ruyseh in his Thesauras Anatomic'us, VI. No. 73, says, in repletione arteriarum, replentur et plurimum quosque vense, et vice versa, ita ut im- possible videatur precise dicere quomodo res .se habeat. Discours sur i’Anat — E p. ~ ' • General Observations on the Ferns. 233 They likewise terminate in exhalent vessels, which open upon the external surface, and upon the various internal surfaces of the body. The secretory vessels of glands are likewise the termination of many arte- ries. SECTION II. Of the Feins. These tubes, which return to the heart the blood carried from it by the arteries, are more numerous than the arteries, and often are larger in diameter. They generally accompany the arteries, and very often two veins are found with one artery. In addition to these last mentioned veins, which may be called deep-seated , there are many subcuta- neous veins which appear on almost every part of the surface of the body. The capacity of all the veins is therefore much greater than that of all the arteries. Those subcutaneous veins, which are of consider- able size, communicate very freely with each other, and also with the deep-seated veins. The trunks of the veins, in those places where no branches go off, are generally cylindrical. There are, however, some exceptions, in which these ves- sels are irregularly dilated, as sometimes happens in the case of the internal jugular vein. It is how- ever, not easy to determine from the appearance of veins injected after death, respecting their situation during life, as their coats are very yielding ; and it is very probable that they are, therefore, preterna- turally dilated by the injection. Veins , directly or indirectly, originate from the termination of arteries : but they do not pulsate as the arteries do, because the impulse given to the Vgl. EE. 30 234 Coats of Veins. blood by the heart is very much diminished in con- sequence of the great diminution of the size of the vessels through which the blood has passed. In some cases, however, when blood flows from an opened vein ; the extent of its projection is alter- nately increased and diminished, in quick succes- sion, as if it were influenced by the pulsation of the heart. The Coats of Veins differ considerably from those of Arteries, — for they are thinner , and so much less firm, that veins, unlike arteries, collapse when they are empty. They Consist of a dense elastic substance, the fibres of which are much less distinct than those of arteries, but some of them are to be seen in a longi- tudinal direction. These fibres can be made to con- tract by local irritation ; for if a vein be laid bare in a living animal, and then punctured, it will often contract so as to diminish its diameter very consi- derably, although no blood shall have escaped from the puuctures. Next to the elastic substance is the internal coat, which is smooth and polished. It is separated from the substance exterior to it with difficulty, although it may be taken from it very easily in the vena cava. This internal coat is more ostensible than the in- ternal coat of arteries, and is not, like the latter, dis- posed to ossification. It is frequently so arranged as to form valves, which are plaits or folds, of a se- milunar form, that project from the surfaces into the cavities of these vessels.* Two of these valves are generally placed opposite The valves of the veins were first described by Charles Etienne of Pa- ris in 1546. In 1547, Amatus, a Portuguese, saw at Ferrara those at the mouth of the vena azygos. Sylvius of Paris announced them about the same time in the jugular, brachial and crural veins. Fabricus ab Acjua- osndente claims the discovery for himself in 1574 Lassus. — Ei?- Colour of the Blood in the / 'inns, 235 to each other ; and, when raised up, they form a sep- tum in the cylindrical cavity of the vessel. The sep- tum, thus composed, is concave towards the heart. The valves have a great effect in preventing the contents of the veins from moving in a retrograde course : they, therefore necessarily modify the effects of lateral pressure, in such a manner, that it propels the blood forward, or to the heart. These valves are generally found in the veins of the muscular parts of the body, especially in those of the extremities. They are not found in those veins which are in the cavities of the body, nor in the in- ternal jugulars. — They are placed at unequal dis- tances from each other. The coats of the veins are somewhat transparent ; and, therefore, those veins which are subcutaneous have a bluish aspect, which is derived from the co- lour of the blood they contain. The colour of the blood in the veins is different from that in the arteries, being of a darker red. The situation and arrangement of the large trunks of veins is much alike in different subjects ; but the branches, especially those which are subcutaneous, are very variable in their situations. CHAPTER II. A PARTICULAR ACCOUNT OF THE DISTRIBUTION OF THE ARTERIES. SECTION r. Of the AORTA,* Or the Great Trunk of the Arterial System „ When the heart ia in its natural position, the right ventricle is nearly anterior to the left, and, therefore, the aorta, where it originates from the left ventri- cle, is behind the pulmonary artery, and covered by it. Its first direction is so oblique towards the right side of the body, that it crosses the pulmonary artery behind, and appears on the right side of it. It has scarcely assumed this position before its course alters, for it then proceeds obliquely backwards, and to the left; so as to form a large curve or arch, which ex- tends to the left of the spine. The position of this curve or arch is so oblique, with respect to the body, that the cord or diameter of it, if it were extended anteriorly and posteriorly, would strike the cartilage of the second or third right rib about the middle of its length, and the left rib near the head. In consequence of this position of the curve, the aorta crosses over the right branch of the pulmonary artery, and the left branch of the windpipe : and assumes a situation, in front, and to the left of the third dorsal vertebra: from this situa- tion it proceeds downwards; in front, but rather on * This name was given by A ristotle.— Et>. Situation ofthe Aorta in the Thorax fy Abdomen. 237 the left side of the spine, and in contact with that column. The aorta, as well as the Pulmonary Artery , for a small distance from the heart, is invested by the pericardium ; and, when that sac is opened, ap- pears to be contained in it. After crossing the right branch of the Pulmonary Artery , a ligament is inserted into it, which proceeds from the main trunk of the Pulmonary artery at its division : this ligament was the Canalis Arteriosus in the foetus. As the aorta proceeds down the spine, it is si- tuated between ,the two lamina of the mediastinum, and in contact with the left lamen, through which it mav be seen. It descends between the crura of the diaphragm, in a vacuity which is sufficiently large to admit of its passage without pressure from the surrounding parts, and is still in contact with the anterior surface of the spine, but rather te) the left of • the middle of it. It continues this course along the spine until it arrives at the cartilaginous substance between the fourth and fifth lumbar vertebra, when it divides into two great branches of equal size, which form an acute angle with each other. These are denominated the common, or primitive iliac Arteries. From the aorta in this course are sent off the ar- teries which are distributed to all the parts of the body for their nourishment and animation. ^ From the curve proceed the great branches which supply the heart, the head, the upper extremities, and part of the thorax. Between the curve and the great primitive iliac arteries, the aorta sends off those branches which supply the viscera contained in the cavities of the thorax and abdomen,* and part * It ought to be observed here, that the viscera iu the lower part of the pelvis receive some branches from the internal iliac arteries. 238 Origin of the Subclavian and Carotid Arteries . of the trunk of the body. The great ILIAC branches of the AORTA are divided into smaller arteries, which supply the whole of the lower extremities and some of the viscera of the pelvis. SECTION II. Of the Branches which go off from the (well of the- AORTA, The proper arteries of the heart, denominated co- ronary arteries, proceed from the aorta so near to the heart that their orifices are covered by the semilunar valves, vrhen those valves are pressed against the sides of the artery. These arteries have been described in the account of the heart.— -See p. 50. The arteries of the head and of the upper extremi- ties proceed from the upper part of the curve in the following manner. A large trunk, called arteria innominata, goes off first. This is more than sixteen lines in length, when it divides into two branches : one of which supplies the right side of the head, and is denomi- nated the right carotid : the other proceeds to the right arm, and from its course under the clavicle, is called, at first, the right subclavian. Almost in contact with the first trunk, another artery goes off, which proceeds to the left side of the head, and is called the left carotid. Very near to this, arises the third artery, which proceeds to the left arm, and is denominated the left subclavian. From these great branches originate the blood vessels, which are spent upon the head and neck and the upper extremities. As these arteries arise from the curve of the aorta, they are situated obliquely with respect to each other. The arteria innominata is not only Situation of the Common Carotid Arteries. 239 to the right, bat it is also anterior to the two others : and the left subclavian is posterior, as well as to the left of the left carotid and the arteria in- NOMINATA. THE CAROTID ARTERIES. The two carotid arteries above mentioned have been denominated common carotids, to distinguish them from their first ramifications which are called internal and external carotids. THE COMMON CAROTIDS Proceed towards the head on each side of the trachea : at first they diverge, but they soon become nearly parallel to each other, and continue so until they have ascended as high as the upper edge of the thyroid cartilage, when they divide into the internal and EXTERNAL CAROTIDS. These arteries are at first very near each other, and rather in front of the trachea ; they gradually diverge and pass backwards and outwards on the sides of it, and of the oesophagus, until they have ar» rived at the larynx. In the lower part of the neck they are covered by the sterno mastoidei, the sterno hyoidei, and thyroidei, as well as by the platysma myoidei muscles. Above, their situation is more su- perficial ; and they are immediately under the pla- tysma myoides. On the inside, they are very near the trachea and larynx, and the oesophagus ; on the outside, and ra- ther anterior to them, are the internal jugular veins; and behind, on each side, are two important nerves called the intercostal and the par vagum . These blood vessels and nerves are surrounded by absor- bent vessels. £40 External Carotid Artery. The common carotid artbries send off no branches from their origin to their bifurcation ; and they appear to preserve the same diameter through- out their whole extent. In some few instances the right carotid has been found larger than the left. The external and internal branches into which they divide, are nearly equal in the adult ; but it is sup- posed that the internal is the largest during infancy. The relative position of these branches is also differ- ent at the commencement from what it is afterwards. The internal carotid forms a curve which pro- jects outwardly, so as to be exterior to the exter- nal carotid, while this last proceeds upwards, and rather backwards. THE EXTERNAL CAROTID ARTERY May be considered as extending from its commence- ment, which is on a line with the superior margin of the thyroid cartilage, to the neck of the condyle of the lower jaw, or near it. At first it is superficial ; but as it proceeds up- wards it becomes deep-seated ; and passing under the digastric and stylo-hyoidei muscles, and the ninth pair of nerves, is covered by the Parotid Gland. After this, it again becomes superficial ; for the tem- poral artery, which may be regarded as the contin- uation of the external carotid, passes over the zygo- matic process of the temporal bone. As the external carotid supplies with' blood the upper part of the neck and throat, the exterior of the head and face, and the inside of the mouth and nose ; its branches must necessarily be numerous, and must pass in very various directions. Thus, soon after its commencement, it sends off, in an anterior direction, three large branches; viz. to the upper part of the neck, to the parts within the lower jaw, and to the cheeks and lips. These are deno- External Carotid and its Branches. 241 minated, the superior thyroid, the sublingual, and the facial. It then sends off to the back of the head one which is called the occipital ; and, as it proceeds upwards near the condyle of the lower jaw, another which passes internallv, behind the jaw, to the deep-seated parts in that direction. After this, it forms the temporal artery, which supplies the fore- head and central parts of the cranium. Besides these larger branches, the external carotid sends of two which are smaller ; one from near the origin of the sublingual artery, which is spent principally upon the pharynx and fauces, and is called the inferior pha- ryngeal : and another, while it is involved with the parotid gland, which goes to the ear; and is there- fore called posterior auris. These arteries are distributed in the following manner : 1. The superior thyroid branch Comes off very near the root of the external carotid, and sometimes from the common trunk ; it runs ob- liquely downwards and forwards, in a meandering’ course, to the thyroid gland, where it is speut. Dur- ing this course it sends off one branch to the parts contiguous to the os hyoides ; another to the neigh- bourhood of the larynx : and a third branch, which may be termed laryngeal , that passes with a small nerve derived from the laryngeal branch of the par vagum, either between the os hyoides and thyroid cartilage, or the thyroid and cricoid cartilages, to the interior muscles of the larynx ; and finally re- turns again to terminate externally. While in the thyroid gland this artery anastomo- ses with the inferior thyroid, and also with its fellow •on the opposite side., VoL, IX. 31 242 Brandies of the External Carotid . t. The LINGUAL, or sublingual branch, Goes off above the last mentioned artery, and very near it ; but in a very different direction, for it runs upwards and forwards, to the tongue. In this course it crosses obliquely the os hyoides, and is commonly within the hyoglossus muscle. It gives off branches to the middle constrictors of the pharynx, and to the muscles contiguous to the tongue. It also sends off a branch which penetrates to the back of ' the tongue, which is called, from its situation, Dorsalis Linguse. ^t the anterior margin of the hyoglossus muscle it divides into two branches, one of which passes to the sublingual gland and the adjacent parts, and is thence called Sublingual ,* while the other branch, thei?am- na , passes by the side of the genio glossus muscle to the apex of the tongue. 3. The facial or external maxillary, Runs obliquely upwards and forwards under the ninth pair of nerves, the stylo hyoideus muscle and the ten- don of the digastric, across the lower jaw and cheek, towards the inner corner of the eye, in a serpentine course. Before it crosses the jaw it sends off several branches, viz. to the pharynx, the tonsils, the inferior maxillary gland and the parts contiguous to it. It also sends a branch towards the chin, which passes between the mylo-hyoideus, the anterior belly of the digastric, and the margin of the lower jaw : and some of its branches continue to the muscles of the under lip. This branch is called the Submental. This artery then passes round the basis or inferior edge of the lower jaw, very near the anterior margin of the masseter muscle, and is so superficial that its pulsations can be readily perceived. After this turn, its course is obliquely upwards and forwards. Near f he basis of the jaw it sends off a branch to the mas Branches of the External Carotid. 243 seter, which anastomoses with small branches from the temporal ; and another which passes superficially to the under lip and contiguous parts of the cheeks. This last is called the Inferior Labial. After the artery has passed as high as the teeth in the lower jaw, it divides into two branches ; which go, one to the under, and the other to the upper lip; that to the upper lip is largest. These branches are called Coronary. The Coronary Artery of the lower lip passes under the muscles called Depressor Angtili Oris, and Or- bicularis Oris, into the substance of the lip, and anastomoses with its fellow of the opposite side. The Coronary Artery of the upper lip passes un- der the zygoinaticus major and the orbicularis, and very near the margin of the upper lip internally. It also anastomoses freely with its fellow on the oppo- site side. These anastomoses are frequently so con- siderable that the arteries on one side can be well filled by injecting those of the other. The coronary branches, as well as the main trunk of the facial ar- tery, observe a serpentine or tortuous course ; in con- sequence of which they admit of the motions of the cheeks and lips, which they would greatly impede if they were straight. from the upper coronary artery a branch conti- nues in the direction of the main trunk of the fa- cial artery, by the side of the nose, which extends upwards, sending olf small branches in its course, and finally terminates about the internal angle of the eye and the forehead. 4. The INFERIOR PHARYNGEAL Is a very small artery ; it rises posteriorly from the external carotid, opposite to the origin of the sublin- gual, and passes upwards to the basis of the era- 244 Branches of the External Carotid. nium. In this course it sends several branches to the pharynx, and to the deep seated parts immediately contiguous. It also sends branches to the first ganglion of the intercostal nerve, to the par vagum, and to the lym- phatic glands of the neck ; and finally it enters the cavity of the cranium by the posterior foramen ia- cerum. In some cases it also sends a small branch through the anterior foramen lacerum. 5. The occipital artery Arises from the posterior side of the external caro- tid, nearly opposite to the facial, but sometimes higher up ; it ascends obliquely, and passes to the back part of the cranium, between the ; transverse process of the atlas and the mastoid process of the temporal bone. In this course it passes over the internal jugular vein and the eighth pair of nerves, and under the posterior part of the digastric muscle ; it lies very near to the base of the mastoid process, and under the muscles which are inserted into it. After emerging from these muscles, it runs superficially upon the occiput, dividing into branches which ex- tend to those of the temporal artery. The occipital artery sends off branches to the muscles which are contiguous to it, and to the glands ef the neck. It also gives off the following branches : one called the Meningeal , which passes through the posterior foramen lacerum to the under and back part of the dura mater : one to the exterior parts of the ear : another which passes downwards, and is spent upon the complexus, trachelo mastoideus, and other muscles of the neck : and' several smaller ar- teries. Branches of the External Carotid. 245 The artery next to be described, is sometimes sent off by the occipital artery. 6. The posterior auricular, or stylo mastoid ARTERY, When it arises from the external carotid, comes off posteriorly from the artery, where it is involved with the parotid gland, and passes backwards between the meatus auditories externus and the mastoid process. It then ascends, in a curved direction, and termin- ates behind the ear. In this course it sends off small branches to the parotid gland, and to the digastric and sterno mas toid muscles. Sometimes a distinct branch, which is particularly visible in children, passes through an aperture in the meatus auditorius externus, and is spent on its internal surface. , * It also sends off a branch which enters into the Stylo Mastoid Foramen, and supplies small vessels to the membrana tympani and the lining membrane of the cavity of the tympanum; to the mastoid eeiis; to the muscle of the stapes, and to the external semi- circular canal. One of these vessels anastomoses, in the upper and posterior part of the cavity of the tym- panum, with a small twig derived from the artery of the dura mater. When it has arrived behind the ear, the Posterim' Auricular Artery terminates upon the external ear and the parts contiguous to it. 7. The INTERNAL MAXILLARY ARTERY^' Arises from the external carotid under the parotid gland, at a little distance below the neck of the con- * The general situation of this artery, and the distribution of several of its most important branches, cannot be understood without a knowledge of the bones through which they pass. The student of surgery will therefore derive benefit from a re-examination of these bones, and of the zygomatic fossa, &c. when he studies this artery. (See Y< 1. 1. page 73.) — He ought 246 Th e Internal Maxillary Artery. • dyloid process of the lower jaw, and extends to the bottom of the zygomatic fossa : varying its direction in its course. It is rather larger than the temporal. a. It first sends off one or two small branches to the ear, and a twig which penetrates into the cavity of the tympanum by the glenoid fissure. b. It also sends off a small artery called the Les- ser Meningeal, which passes upwards, and after giving branches to the external pterygoid and the muscles of the palate, passes through the foramen ovale, and is spent upon the dura mater about the sella turcica. c. It then sends off one of its largest branches, the Great or Middle Artery of the Dura Mater , which passes in a straight direction to the foramen spinale, by which it enters into the cavity of the cranium. This artery ramifies largely on the dura mater, and makes those aboreseent impressions which are so visible in the parietal bone. It generally divides into two great branches : the anterior, which is the largest, passes over the anterior and inferior angle of the parietal bone: the posterior branch soon divides into many ramifications, which are extended lateral- ly and posteriorly. It furnishes the twig which passes to the ear by the hiatus of Fallopius, and anastomoses with the small branches of the stylo mastoid artery. It also supplies some other small vessels which pass to the cavity of the tympanum by small forami- na near the junction of the squamous and petrous portions of the temporal bone. d. The next branch sent off by the internal max- illary leaves it about an inch from its origin, and is to be well acquainted with this subject, if he should undertake the man- agement of necrosis of the jaw bones ; or of those fungous tumours, which sometimes originate in the antrum iraxillare its well as of several other complaints. The Internal Maxillary Artery, 247 called the Inferior Maxillary . It passes between the internal pterygoid muscle and the bone, and after giving small branches to the contiguous miiscles, en- ters the canal in the lower jaw, in company with the nerve. This canal has a very free communication with the cellular structure of the jaw, and the artery in its progress along it sends branches to the respective teeth and the bone. At the anterior maxillary fora- men, this artery sends off a considerable branch, which passes out and anastomoses with the vessels on the chin, while another branch passes forward and sup- plies the canine and incisor teeth and the bone con- tiguous to them. Sometimes the inferior maxillary artery divides into two branches before it has arrived at this fora- men. In this ease, one of thp arteries passes out of the foramen, while the other continues to the sym- phisis. e. Two branches pass off to the temporal muscle, which originate at a small distance from each other : one of them passes upwards on the tendon of the temporal muscle ; the other arises near the tuberosity of the upper maxillary bone: they are called the ex- terior deep , and the interior deep temporal artery . They are both spent upon the temporal muscle; . but the interior branch sends a small twig into the orbit of the eye. F. There are some small branches which pass to the Pterygoid Muscles and to the Masseter , which arise either from the internal maxillary artery, or from the anterior deep temporal. They are generally small, and often irregular, g.. An artery, particularly appropriated to the cheek, perforates the buccinator muscle from within outwards, and generally terminates on the buccinator, the zygomatieus major and the muscles of the lips. This Artery of the Cheek is very irregular in its 248 The Internal Maxillary Artery. origin, sometimes arising from the internal maxillary, sometimes from the deep temporal, and sometimes from the suborbitary, or from the alveolar artery, to be immediately described. h. The Alveolar Artery, or the Artery of the Upper Jaiv, arises generally from the internal maxil- lary, but sometimes from one of its branches. It winds round the tuberosity of the upper jaw, and sends branches to the buccinator muscle, to the bone and the gums, to the antrum highraorianum, and some of the molar teeth: and also to the teeth generally, by means of a canal which is analogous to that of the lower jaw. i. The Infra orbitar Artery arises from the inter- nal maxillary in the zygomatic fossa, and soon enters the infra orbitary canal, through which it passes to the face, and emerges below the orbit of the eye, supplying the muscles in the vicinity, and anasto- mosing with the small ramifications of the two last described arteries, and also of the facial artery and the ophthalmic. This artery in its course sends off small twigs to the periosteum, the adipose membrane, and the mus- cles in the inferior part of the orbit, and also to the great maxillary sinus or antrum highmorianum, and to the canine and incisor teeth. j. The Palato Maxillary , or Superior Palatine Artery, arises also in the zygomatic fossa, and, de- scending behind the upper maxillary bone, enters the posterior palatine canal. It generally forms two branches, the largest of which advances forward, supplying the palate and gums, and finally sends a twig through the foramen incisivum to the nose, while the posterior branch, which is much smaller, supplies the velum pendulum palati. k. The Ptery go Palatine, or Superior Pharnygeal, is a small vessel ; which sometimes arises from the 249 Temporal Artery, artery next to be mentioned. It is spent upon the upper part of the pharynx, and a branch passes through the pterygo palatine foramen, which is spent upon the arch of the palate and the contiguous parts. L. The internal maxillary at length termi- nates in the Spheno- Palatine, or Large Nasal Ar- tery, which passes through the spheno-palatine fora- men to the back part of the nose. This artery sometimes separates into two branches before it en- ters the foramen; sometimes it enters singly, and divides into two branches soon after; one of them is spread upon the septum, and the other upon the ex- ternal side of the nose ; each of these branches rami- fies very minutely upon the Schneiderian membrane and its processes in the different sinuses, and also in' the ethmoidal cells. 8. The TEMPORAL ARTERY Is considered as the continuation of the external ca- rotid, because it preserves the direction of the main trunk, although the internal maxillary is larger. After parting with the internal maxillary it pro- jects outwards ; and passing between the Meatus Auditorius Externus and the condyle of the lower jaw, continues upwards, behind the root of the zygo- matic process of the temporal bone, to the aponeuro- sis of the temporal muscle : on the outside of which, immediately under the integuments, it divides into two large branches denominated anterior and poste- rior. Before this division the temporal artery sends off several branches of very different sizes. One, which is considerable'in size, and called the Transverse Facial Branch, advances forwards across the neck of the condyle of the lower jaw, and giving small branches to the masseter, runs parallel to the parotid duct,, and below it. This branch is spent Vol, it. • -32 250 Temporal Artery.-— Internal Carotid . upon the muscles of the face, and anastomoses With the other vessels of that part. The temporal gives off small branches to the pa- rotid gland and to the articulation of the jaw. From the last mentioned branch small twigs pass to the ear, one of which enters the cavity of the tympanum by tbe glenoid fissure. ■While this artery is on a line with the zygoma, it sends off a branch called the middle temporal artery , which penetrates the aponeurosis of the temporal muscle, and ramifies under it upon the muscle in an anterior direction. The two great branches of the temporal artery are distributed in the following manner. The Anterior passes up in a serpentine direction on the anterior part of the temple, and supplies the front side of the head, and the upper part of the forehead. The Posterior extends upwards and backwards, and supplies the scalp on the lateral and middle part of the cranium, and also the bone. Ramifications from each of these branches anasto- mose on the upper part of the cranium with those of its fellow of the opposite side. The anterior branch also anastomoses on the forehead with the facial and ophthalmic artery; and the posterior branch with the occipital artery on the back part of the head. THE INTERNAL CAROTID ARTERY Is sometimes called the Artery of the Brain , as it is almost entirely appropriated to that viseus. From its origin to the commencement of its rami- cations the course of this blood vessel is peculiarly tortuous. In consequence of which the force of the blood in it is greatly diminished before it arrives at the brain. An instance of this curvature occurs immediately Cnrmtnres cf the Internal Carotid . 251 after its separation from the external carotid, when it protrudes outwards so much as to be exterior to that vessel ; after this it ascends to the carotid canal, and. in its course is in contact, or very near the par va- gum and intercostal nerves. The carotid canal in the os petrosum is by no means straight; it forms a semicircular curve, for- wards and inwards; and its upper portion, which is nearly horizontal, opens obliquely against the body - of the sphenoidal bone, at a small distance from it. Therefore, after the artery lias passed through the canal, it must turn upwards to get fairly into the cavity of the cranium ; and of course, its direction while in the canal, forms almost a right angle with its direction before it enters, and after it emerges from it. In consequence of this curvature, much of the mo- mentum of the blood must be impressed upon the cranium. After the artery has arrived at the end of the caro= tid canal, and has turned upwards to get within the cavity of the cranium, it bends forwards, and passes nearly in a horizontal direction, through the caver- nous sinus on the side of the sella turcica, to the an- terior clinoid process ; here it again forms a conside- rable curve, which is directly upwards, and then it perforates the dura mater. These curvatures must also deprive the blood of the carotid of a portion of the momentum which it has retained after leaving the bone. The object of these various flexures of the internal carotid appears to be analogous to that of the Rete Mirabile in certain quadrupeds, which is formed by the division of this artery into many small branches, that reunite again, without producing any other ef- fect than the diminution of the momentum of the blood . 252 Ophthalmic Artery. During its course from the place of bifurcation to its entrance into the carotid canal, the internal carotid artery very rarely sends off any branches. In the canal it gives off a small twig which enters the cavity of the tympanum ; and sometimes a second which unites with the Pterygoid branch of the internal maxillary. As it goes by the sella turcica, it passes through the cavernous sinuses, and gives off two branches which are called the Postet'ior and Anterior Arteries of the Cavernous Sinus or Receptacle. The posterior branch goes to that part of the dura mater which is connected with the posterior clinoid process, and the cuneiform process of the occipital bone. It likewise gives branches to several of the nerves which are contiguous, and to the pituitary gland. The anterior artery also gives branches to the con- tiguous nerves, to the dura mater, and the pituitary gland. When the internal carotid turns upwards at the anterior clinoid process, it sends off the OPHTHALMIC ARTERY Which passes under the optic nerve through the fo- ramen opticum into the orbit of the eye, and is about a line and a half in diameter. Although this artery enters the orbit under the op- tic nerve, it soon takes a position on the outside of it, but afterwards gradually proceeds to the inner side of the orbit, crossing over this nerve in an oblique di- rection, and finally passes out of the orbit near the in- ternal angle. In this spiral course it sends off nu- merous branches, viz. a. To those parts which are auxiliary to the eye. b. To the bail of the eye. Branches of the Ophthalmic Artery. 253 c. To the cavity of the nose, through small fora- mina in the ethmoid bone, and d. To the forehead and external side of the nose. These branches generally go off in the following order. 1. The Lachrymal artery arises soon after the ophthalmic arrives within the orbit, and passes above the abductor muscle to the lachrymal gland, where it terminates, sending off many small branches in its course. 2. The Central artery of the retina also leaves the ophthalmic soon after its arrival in the orbit: it is a small vessel which penetrates into the centre of the optic nerve, and passing with it into the eye is spread upon the internal surface of the retina. Here it appears to terminate in the adult ; but in the foetus it is continued through the vitreous humour to the capsule of the crystalline lens. 3. While the ophthalmic is passing over the optic nerve the branches which enter the ball of the eye leave it. Their number varies, but they form three classes, viz. The Long Ciliary , the Short Ciliary. and the Anterior Ciliary arteries. (See description of the eye, vol. i. p. 352,) the supra orbitary and muscular branches leave it also near the same places. 4. The Supra Orbitary Branch often gives off several muscular twigs : but it passes out of the orbit through the supra orbitar foramen, and generally divides into two branches, one of which is spent upon the periosteum, and the other upon the skin and muscles of the forehead. 5. There are sometimes two muscular branches, a Superior and an Inferior. The superior branch is often deficient : when it exists it supplies the levator palpebrae, the levator oculi, obliquus superior, &c. ; but these parts are often supplied by the branches above mentioned. The supra orbitar so frequently &o4 Branches of the Ophthalmic Artery* gives off branches to the muscles that it has been called the Superior Muscular Branch. The inferior muscular branch is more constant. It commonly supplies the rectus inferior, the adductor, and the inferior oblique muscles, and also the lachrymal sac, and the lower eyelid, &c. When the artery is on the inside of the nerve it sends off the two branches to the cavity of the nose, viz, The Ethmoidal Arteries ; and also, branches to the eyelids. 6. The Posterior Ethmoidal branch is first. It passes between the levator and abductor muscles, and above the obliquus superior ; and penetrates the cavity of the cranium by the posterior orbitary fora- men : after giving some twigs to the dura mater, it passes to the posterior cells of the ethmoid by the foramina of the cribriform plate of that bone, and sends a small branch to the Schneiderian membrane on the back part of the septum of the nose. 7. The Anterior Ethmoidal artery arises from the ophthalmic nearly opposite to the anterior orbitary foramen, through which it passes: and after enter- ing the cranium is distributed like the other through some of a foramina of the cribriform plate to the anterior cells of the ethmoid bone, and to the ante- rior part of the Schneiderian membrane on the sep- tum of the nose, to which it sends a considerable branch. In its course it sends twigs to the frontal sinuses, and to the dura mater and its falciform process. 8. The arteries of the Palpebrse are called Supe- rior and Inferior ; they leave the ophthalmic near the loop or pully of the superior oblique muscle. The inferior comes off first ; it sends branches to the ligaments of the tarsus, the caruncula lachrymalis, and the parts connected with the cartilage of the un- der eyelid, and unites with the lachrymal artery near Brandies of the Ophthalmic Artery. Carotid. 2 53 the external canthus, forming ail arch called the In* ferior Tarsal Arch. 9. The Superior Artery supplies the superior part of the orbicularis muscles, the ligament and carun- cula also : and it likewise unites with a twig of the lachrymal, and forms the superior tarsal arch. Soon after sending off the palpebral branches, the Ophthalmic Artery arrives at the internal canthus, and then finally divides into two branches, the nasal aiid the frontal. 10. The A r asal Branch passes above the superior part of the lachrymal sac and the ligament of the eye-lid to the nose ; after sending a twig to the fron- tal muscle and the lachrymal sac, it passes down the side of the nose and anastomoses with the facial ar- tery. 11 . The Frontal Artery is not so large as the na- sal ; it generally divides into three parts. . A super- ciliary branch which is principally spent upon the eyebrows ; a superficial branch which is spent upon the forehead ; and a branch which is distributed to the pericranium. The mTEENAL carotid, soon after parting with the ophthalmic, sends off, in a posterior direction, a branch to join one from the vetebral artery. From its distination this vessel is called the arteria com - municans , After this it sends off another branch, which is so large that it may be considered as a continuation of the main trunk : this is called the middle cadery of the brain, or the Arteria Sylviana. It runs outwards nearly in the direction of the fossa Sylvii, which se- parates the anterior from the middle lobes of the ce- rebrum. In its course it divides and subdivides into numerous branches which are spread upon the Pici Mater , and finally enter the surface of the brain, in a very minute state. 256 Right and Left Subclavian Arteries , The internal carotid then terminates in a branch which is smaller than the last mentioned, and from its situation is called the Anterior Artery of the Brain , or Arteria Callosa, This vessel first inclines towards its fellow on the opposite side, and after ap- proaching within half an inch of it, forms another curve, and runs forward to the anterior part of the brain, dividing itself gradually into branches which pass in several directions. When these anterior arteries are nearest to each other, a small transverse branch, which passes at right angles, connects them together. This branch completes the anterior part of the Circle of Willis. It crosses immediately before the sella turcica and pituitary gland, and sends off branches. Which pass to the third ventricle, to the fornix and septum luei- dum, and also to the pia mater. The Anterior Arteries of the brain also send off branches to the optic and olfactory nerves ; to the opposite surfaces of the two hemispheres on each side of the falx ; to their inferior surfaces, and to the corpus callosum. They have likewise some branches which anasto- mose with those of the middle artery of the brain, and of the vertebral artery. The SUBCLAV’IAN Arteries. The right subclavian may be considered as the continuation of the arteria innominata. This last mentioned artery, after leaving the aorta, forms a curve or arch, which extends obliquely backwards and outwards, over the first rib to the axilla, crossing the trachea in its course. At the distance of an inch and a quarter, or an inch and a half from its origin, it sends of the right carotid, and then, assuming the name of Right Subclavian , continues in the above stated direction- ' 257 The Left Subclavian Jittery. The chord of the carve of this artery, and the chord of the curve of the aorta, are not in the same direction, but form an angle with each other. The position of the left subclavian is somewhat different from that of the right. Its origin is posterior, and, therefore, the direction of the chord of its curve is more immediately lateral. The curve or arch is also smaller. The situation of the two subelavians as relative to the contiguous parts, is, therefore, somewhat different ; but each of them proceeds be- tween the anterior and the middle scaleni muscles, and when they have arrived at these muscles, their respective positions are very similar. The anterior and middle scaleni muscles arise from the transverse processes of several of the cervical vertebrae, and are inserted into the first rib, one be- fore the other, so as to leave a considerable space be- tween them. The subclavian arteries § pass through this 3pace, and before they arrive at it, but when they are very near the above mentioned muscles, they send off several very important branches in various directions, viz. to the cavity of the cranium, to the parietes of the thorax, to the thyroid gland, and to the lower part of the neck. They proceed near to the scaleni muscles before they send off any branches ; and it is to be observed, that the subclavian veins which correspond with these arteries, are anterior to them, for they pass before the scaleni muscles, and not between them. The internal mammary Jlrtery Goes downwards, from the lower and anterior part of the subclavian, along the inner side of the anterior scalenus muscle. It proceeds, exterior to the pleura, across the cartilages of the true ribs, and and near their middle ; and, continuing between the cartilages and the diaphragm, exterior to the peritoneum, term!* tTo h. Ho 3fS 258 Course and Distribution of the Inferior nates on the rectus .abdominis muscle, in branches which anastomose with those of the epigastric artery. In this course it gives branches to almost all the parts to which it is contiguous, viz. to the muscles and glands at the lower part of the neck ; to the thymus gland ; to the parts in the intercostal spaces; to the sternum ; to the mediastinum and pericar- dium ; to the diaphragm and to the muscles of the abdomen. From some of its ramifications upon the parts be- tween the ribs, small branches go off to the mamma, and thereby give a name to the artery. There is also a small vessel which is sent off by the mammary ar- tery, or by one of its upper branches, which accom- panies the phrenic nerve to the diaphragm. The inferior thyroid Artery Arises from the upper side of the subclavian nearly opposite to tKe origin of the internal mammary. It passes upwards and inwards, between the carotid artery and the spine, to the thyroid gland : then it anastomoses with the branches of the superior thyroid on the same side, and with those of its fellow on the opposite side. This vessel sometimes sends off large branches to the muscles at the lower part of the neck. The vertebral Artery Arises from the upper and posterior part of the sub- clavian. It goes upwards and backwards between the muscles which lie on the front of the spine, and passing under the transverse process of the sixth or seventh cervical vertebra, enters into the canal form- ed in the transverse processes of the vertebrae. In this course, as it proceeds from the «third to the se- cond cervical vertebra, it inclines outwards laterally , and, in its passage from the traosvere process of Thyroid and the Vertebral Arteries, 259 the second to that of the first vertebra, it forms a considerable curve, the convexity of which has a lateral and external aspect. After passing the trans- verse process of the Atlas , it is turned suddenly backwards, in a groove, and finally passes through the great occipital foramen into the cavity of the cra- nium. It then proceeds upon the cuneiform process, of the occipital bone, under the Medulla Oblongata , and joins its fellow so as to form an acute angle with it near the union of the medulla oblongata with the pons Varojii. From each of the vertebral arteries, before their union, there generally goes off a small branch called the Posterior Meningeal , which is spent upon the posterior part of the dura mater. The trunk formed by the union of the vertebral arteries is called The BASILAR Artery . It extends forward near to the anterior part of the pons Varolii, where it bifurcates ; but previously sends off several branches on each side. The first pair go off in a lateral direction, soon after its com- mencement, near the back part of the pons Varolii, and are spent upon the medulla oblongata, the pons Varolii, and the other contiguous parts, and also upon the fourth ventricle and the Plexus Choroides of that cavity. They are called the Posterior or Inferior Arteries of the Cerebellum. Two other lateral branches, which are called the Superior Arteries of the Cerebellum , go off from the Basilar artery, near its anterior extremity. These are principally spent upon the crura of the cerebel- lum and cerebrum ; upon the cerebellum itself, and the contiguous parts. Soon after sending off the last mentioned arteries, the Basilar artery divides into two branches, which also take a lateral direction, and are of considerable 260 Arteries of the lower part of the JYech. size. In their course outward, these branches are curved with their convexity forward. About ten of twelve lines from its commencement, each of them sends off a branch called the Arteria C omnium cans? - which passes directly forward, and communicates with the internal carotid, thus forming the arrange- ment which is called the Circle of Willis .* After sending off these arteries, they continue their lateral direction, and are distributed principally to the pos- terior parts of the cerebrum. These terminating branches of the Basilar Artery , are called the Pos- terior Arteries of the Cerebrum . The superior intercostal Artery Arises from the upper part of the Subclavian , after the Vertebral and Thyroid arteries, and very near them. It descends by the side of the spine across the first and second ribs, near their heads, and exte- rior to the great intercostal nerve. It generally forms two branches, which are appropriated to the mus- cles, &c. in the first and second intercostal spaces, and sometimes a small branch is continued to the third intercostal space. From each of these branches a small vessel proceeds backwards, and is spent upon the contiguous muscles, &c. on the back of the tho» rax. The Intercostal Artery also sends a branch upwards to the deep-seated parts of the neck. In addition to the arteries above mentioned, there are several others of considerable size which origi- nate either directly or indirectly from the subcla- * The arteria communicans is also considered as a branch of the Inter- nal Carotid. The arrangement here alluded to is very remarkable. As the branches which pass off laterally from the single trunk of the Basilar Jittery unite to the Internal Carotids, and the Internal Carotids are united to each other, there is an uninterrupted continuation of artery. Which en- closes a portioh of space of a determined form ; but this form resembles an oblong square more than a circle. By this connexion blood will pas? from any one of the four arteries of t’ne brein ceed differently in different subjects : the right pass- ing behind, and the left before, the subclavian artery of its respective side. Each of them, however, be- comes contiguous to its corresponding artery. When it has arrived at the place in the transverse processes, where the artery enters the vertebral canal, it sends off an external branch, which passes up, before and nearly in contact with, those processes, and gives ramifications to the contiguous muscles, and also to 304 Vertebral Veins. the cavity of the spine. These last mentioned, rami- fications enter by the lateral apertures between the transverse processes, and anastomose with the veins and sinuses of the cavity. The branch often finally terminates in the lateral sinus of the dura mater, by passing through the foramen near the mastoid process of the temporal bone. The Main Trunk of the ver- tebral vein generally sends off another external branch to the muscles near the basis of the neck, and after- wards enters the canal with the vertebral artery. While in this canal it generally sends off two branches through each of the lateral apertures between the ver- tebrae. One of these branches passes backwards to the muscles of the neck, and the other proceeds into the great spinal cavity, and communicates with the venous sinuses. When it has arrived at the atlas, the Vertebral vein sends branches to the contiguous muscles of the neck. It also frequently sends a branch through the posterior condyloid foramen of the occipital bone to the lateral sinus. It is evident, from these circumstances that the vertebral vein carries a portion of blood from the sinuses of the brain and of the spinal marrow, as well as from the muscles of the neck, into the sub- clavian veins. The veins of the head are frequently very differ- ent in different subjects. The INTERNAL JUGULAR , Already mentioned, is often almost exclusively ap- propriated to the cavity of the cranium ; and all the exterior veins of the head are ramifications of one or more smaller vessels, which pass up superficially on the neck, and are denominated External Jugulars . In some instances almost all the exterior veins of the head are united to the internal jugular at the upper The internal Jugular , 305 part of the neck, and it of course conveys the blood of the exterior as well as of the interior parts of the head. Frequently these veins are divided between the internal and external jugulars, but they are di- vided very differently in different subjects. The Internal Jugular, however, almost always passes in the same direction from the inside of the origin of the sterno-mastoid muscle to the posterior foramen lacerum of the cranium. It is deeply seat- ed on the external side of the common carotid artery, and under the sterno-mastoid muscle. Between the upper margin of the thyroid cartilage and the angle of the lower jaw, it often sends off branches which are very different in different subjects, but commonly pass to the anterior parts of the neck and face : above these it generally sends another to communi- cate with the external jugular. One of the branches which often go off from the internal jugular is that which corresponds with the superior thyroid or la- ryngeal artery. This vein, which has sometimes been called the Guttural, sends many ramifications to the thyroid gland. The Ranular veins, which are so conspicuous under the tongue, are also derived from it ; and it likewise sends branches to the larynx and pharynx. Before the internal jugular enters the foramen la- cerum, it suffers a partial dilatation, which is gener- rally larger on one side than the other. * This dila- tation occupies the fossa at the foramen lacerum. After passing through the aforesaid foramen, the in- ternal jugular terminates in the lateral sinuses of the * When the veins of the neck are injected, it very often appears that a considerable portion of the internal jugular is much larger on one side than the other, as if it were affected with varicose distention. It also often appears that the general arrangement of the exterior vein is different on the two sides of the head and neck, VOL, II. 39 306 The External Jugular . dura mater.* These and the other sinuses within the cavity of the cranium are important portions of the venous system, which are interposed between the smaller branches spread upon the pia mater and the great trunks of the neck. They are described in the account of the brain, (Yol. I. page 316.) Into these sinuses the very numerous veins of the pia mater open, proceeding to the sinuses in a direction the reverse of that in which the blood flows in those channels. These veins are divided very minutely on the pia mater before they enter the substance of the brain. Into one of these sinuses, denominated the Ca- vernous, the ophthalmic vein discharges its contents. This vein proceeds from the anterior part of the sinus into the orbit of the eye through the sphenoid fissure. t Its ramifications correspond generally with those of the ophthalmic artery^ and some of them pass out of the orbit to anastomose with the branches of the facial vein. The superficial veins of the neck are variously arranged in different persons. There is often one considerable vein, The EXTERNAL JUGULAR, Which is sent off by the subclavian, very near its union with the internal jugular ; but sometimes it goes off from that vein much nearer the shoulder. There are sometimes two external jugulars, an ante- * It is asserted that the internal coat, or lining membrane of the inter, nal jugulars, is continued into the lateral sinuses, and extends throughout all the sinuses of the dura mater ; so that the blood, during its passage through the sinuses, does not come in contact with any membrane different from that of the veins. f See the account of this fissure in vol. i. p. 63. t The Vasa Vorticasa of the choioides aie one of the exceptions to this. See vol. i. p- 356. General Account of the Great Fein of the Arm. 307 rior and a posterior, nearly of equal size. More fre- quently one of them is much smaller than the other. In a majority of cases, the principal external jugular goes off near the junction of the internal jugular and subclavian, as above stated, and proceeds upwards towards the angle of the lower jaw, passing between the platysma myoides and the stern o- mastoid muscle. It often sends off, at the basis of the neck, one or more branches to the contiguous muscles, and then proceeds upwards. Near the angle of the jaw, it often com- municate&with the internal jugular : itthen continues upwards, covered with the parotid gland, near the temporal artery, and finally divides into superficial and deep-seated temporal branches. The External Jugular, near the angle of the jaw, often sends off the facial vein, which crosses the basis of the lower jaw, near the facial artery, and distri- butes branches to the side of the face and to the fore- head. It also very often sends off, near this place, the internal maxillary vein, which generally ramifies in such a manner that its branches correspond with those of the internal maxillary artery. Veins which correspond to some of the other branches of the exter- nal carotid artery, the lingual, occipital, &c. are of- ten sent off near this place by the external jugular. They take the names of the arteries to which they correspond, and commonly accompany them. The SUBCLAVIAN Vein, Although it originates differently on the two sides of the neck, is situated alike on each of them. After parting with the internal jugular, it proceeds over the first rib, under the clavicle, and does not pass between the scaleni muscles, as is the case with the arteries, but before the anterior muscle. It soon joins the great artery of the arm, and proceeds be- fore or below it to the axilla. In this situation it gives 308 General Account of the Great Vein of the Arm. off branches to the contiguous parts, which correspond with those given off by the artery. In this course it also often gives off a large branch, called the CEPHALIC, Which soon becomes superficial, and proceeds down- wards between the margins of the deltoid and pecto- ral muscles : it continues superficial on the external side of the biceps muscle, sending off many subcuta- neous branches. Near the external condyle of the os humeri, it generally sends off a branch towards the middle of the anterior part of the fore-arm, which is called the Median Cephalic, and also some other su- perficial branches. It then continues over the radius, and inclining to the back of the fore-arm, until it ar- rives at the back of the hand, where it divides into branches, some of which go to the thumb. In the axilla, the great vein, there called The AXILLARY Vein, Generally divides into two or three branches. One, which is commonly the largest, and appears like the continuation of the main trunk, is called The basilic Vein . This vessel passes down, deeply seated, to the bend of the elbow. It becomes superficial near the inter- nal condyle, and divides into several branches. — One of these generally proceeds to join the median branch of the cephalic, and from the union of the two branches is formed the median vein, which passes down near the middle of the anterior part of the fore- arm. This vein generally sends off a branch which proceeds internally, and anastomoses with the deep- seated veins of the fore- arm. There are frequently two other branches of the basilic vein. One, which is small, passes down on Situation of the Inferior Vena Cava. 309 the ulnar side of the anterior part of the fore-arm, but does not extend to the wrist. The other passes down on the ulna, and gradually proceeds to the back of the hand, when it divides into several branches, one of which is generally appropriated to the little finger. The axillary vein, after the Basilic leaves it, sometimes divides into two branches, and sometimes continues undivided. In either case it accompanies the humeral artery, and takes the name of humeral Vein or Veins. It sends off branches which corres- pond to those of the artery, and continues to the bend of the elbow ; here it is so divided, that two of its ramifications accompany each of the three arteries of the fore-arm. These ramifications sometimes com- municate with each other by anastomosing branches near the elbow, and they communicate also with the superficial veins. The superficial veins of the arm are so different in dif- ferent subjects, that a general description will rarely apply accurately to an individual case. It may, how- ever, be observed, that a Cephalic vein will generally be found, which very frequently arises from the sub- clavian instead of the axillary, and commonly conti- nues to the hand on the radial side of the arm. The superficial veins, on the ulnar side of the fore-arm, * very frequently are branches of a large vein which ac- companies the humeral artery to the elbow, viz. the basilic ; but the median vein, formed by branches of the cephalic and basilic veins, is very often not to be found. SECTION II. Of the INFERIOR VENA CAVA, and the Veins which are connected with it. This great vessel exceeds the Superior Cava in diameter. It proceeds from the lower part of the 310 Situation of the hiferior Vena Cava. right auricle, and very soon perforates the dia- phragm, at a small distance in front of the spine, and rather to the right of the centre. As the peri- cardium adheres to the diaphragm at this place, the vessel appears to leave it abruptly. Immediately after leaving the diaphragm, it proceeds along a groove in the posterior edge of the liver, formed by the great lobe and the lobulus Spigelii.* After leaving the liver, it continues downwards, inclining backward and to the left, and is soon in contact with the aorta, which is on the left of it. It accompanies the aorta to its great bifurcation, and divides in the same manner. It sends off, during this course, branches to the Diaphragm, Liver, Right Renal Gland, the Kidneys, and the testicles ; and also the Lumbar and Middle Sacral veins. The Inferior Phrenic Veins Are thus denominated to distinguish them from other veins, which are derived from the internal mammary, &c. They generally accompany the phrenic arteries, and are distributed in the same manner. The hepatic Veins Pass off from the vena cava, nearly at right angles, into the substance of the liver, while it is in the groove of that viscus, and before it has proceeded more than eight or ten lines from the heart. They arise from the anterior part of the vena cava, and are generally three in number. Some- times there are two only, but then one of them di- vides immediately after it enters the substance of the gland. The distribution of these vessels in the liver has 4 Sometimes it is completely surrounded by the live/. The Vena Portarum.**— Splenic Vein. 311 been detailed in the account of that organ, and there- fore need not be stated here ; but the veins which unite to form the vena portarum, and the trunk of that great vein also, before it is connected with the liver, may be regarded as a portion of the regular venous system, and ought now to be considered. The VENA PORTARUM Passes downwards from the great sinus of the liver behind the pancreas, and inclining to the left. In this course it sends branches to the gall bladder, the sto- mach and pylorus, and the duodenum. At the up- per and posterior edge of the pancreas, it sends off a very large branch to the spleen, which often pass- es, with slight meanders, along a groove in the pan- creas. The SPLENIC Vein Often sends off the inferior mesenteric vein, which proceeds downwards between the aorta and the left portion of the colon. It also sends off some of the coronary veins and the left gastro epiploic vein to the stomach ; many small branches to the pan- creas; and, finally, either from the main trunk or its branches before they enter the spleen, the venae breves, which pass to the great extremity of the sto- mach. Before it enters the spleen, it forms several ramifications, which accompany the branches of the splenic artery. After sending off the splenic, the Vena Portarum takes the name of The SUPERIOR MESENTERIC Vein; Which is larger than the splenic, and passes from behind the pancreas > before the transverse portion of 312 Superior Mesenteric Vein. —Emulgen t Veins, the duodenum , into the mesentery ; where it accom- panies the superior mesenteric artery. It is evident that the above, described portion of the vena portarum simply performs the functions of a great vein ; but when it takes on the arrangements for entering the liver, it no longer acts like a vein, but an artery. The lower portion of the trunk of this vein and its ramifications is denominated Vena Portse Ventralis. The part which ramifies in the liver, Vena Portx Hepatica. The Capsular Veins Are small vessels, one on each side. That on the right passes from the vena cava to the right glandula renalis. That on the left arises from the left emul- gent vein. The EMULGENT, or RENAL Veins, Are very large vessels; and, like the arteries, go off nearly at right angles, one to each kidney. The right emulgent vein is not so long as the left, and it is rather anterior to its corresponding artery. The left emulgent, in its course to the kidney, cross- es the aorta, and is anterior to it. These veins pass to the sinus of each kidney, and ramify before they enter it. The ramifications fol- lows those of the arteries. The Spermatic Veins Arise one on each side : the right from the vena cava and the left from the emulgent vein. They proceed downwards behind the peritoneum, and on the psoas muscle generally divide into many branches which communicate with each other as they progress down- wards, and form a plexus denominated Corpus Pampiniforme, These branches proceed in the Internal Iliac and its Ramifications. 313 spermatic cord to the back of the testis. The prin- cipal part enters the body of that gland; but some of the branches go to the epididymis. In females the spermatic vein, like the artery, passes to the ovary, the uterus and its appendages, &c. The Lumbar Veins Correspond to the arteries of the same name. They arise from the posterior and lateral parts of the in- ferior cava, and those on the left side pass under the aorta. The Middle Sacral Vein Resembles the artery of the same name in its origin and distribution. The INFERIOR VENA CAVA accompanies the aorta to the space between the fourth and fifth lumbar vertebrae, and there it also divides into the two PRIMITIVE ILIAC VEINS. • The left vein crosses behind the artery of the right side, and rather behind the left primitive iliac artery, which it accompanies until they are opposite to the junction of the sacrum and ilium, when it divides again, like the artery, into the internal and exter- nal iliac veins. The INTERNAL ILIAC , or HYPOGASTRIC Vein ' Descends into the pelvis behind the artery, which i£ accompanies. Its ramifications correspond in gene- ral with those of the artery, and, therefore, need not be particularly described. The VENiE VESICALES Have such peculiarities that their ramifications re- quire particular attention. They arise from the hy- Vol. ii« 40 314 JRamiJications of the External Iliac . pogastric, very near the origin of the obturator, and are large as well as numerous. They are somewhat different in the two sexes. In men they form a remarkable plexus on the lateral and inferior portions of the bladder, and on the ve- siculse seminales. This plexus extends more or less to the prostate : from it a number of veins proceed to the symphisis of the os pubis, which communicate in their course with the pudic vein. From thence arises the great vein of the penis, which proceeds in the groove between the corpora cavernosa, and ter- minates in the glans penis. This vein often divides, near the root of the penis, into two : one of which is in the groove, and the other more superficial.* In females, the venae vesicales form a considerable plexus on each side of the bladder and vagina. — • Many veins pass from these to the upper portions of the bladder and the contiguous parts, and form plexuses. The clitoris has a dorsal vein like the penis, and it originates in a manner analogous to the dorsal vein of the male. The EXTERNAL ILIAC Vein . The great trunk of the veins of the lower extremity proceeds on the inside of the artery, under the crural arch or Poupart’s ligament. Before it passes from under the arch, it sends off two branches which an- swer to the circumflex artery of the ilium and to the epigastric artery. The Circumflex Vein Arises from the external side of the iliac vein, and passes towards the anterior end of the spine of the * The pudic veins accompany the arteries of that name. They com- wmnicate with the plexus, as above mentioned, and continue into the penis 315 Veins of the Leg . ilium. It divides into branches which accompany those of the artery of the same name. The Epigastric Vein Arises from the external iliac, and accompanies the epigastric artery.— After passing a small distance in=? ward and downward, it turns up on the inside of the abdominal muscles. In the first part of its course it sends off some small branches to the spermatic cord. After passing beyond Poupart’s ligament the name of the great vessel is changed from external iliac to FEMORAL VEIN. It proceeds downwards at first on the inside of the femoral artery, but gradually changes its relative situation, so that in the thigh and in the ham it is behind or on the outside of that vessel. At a short distance below Poupart’s ligament, af- ter giving off some small branches to the external or- gans of generation, and to the glands of the groin, it sends off on the internal side of the thigh a very large vein which is called the SAPHENA MAJOR. This vein immediately becomes superficial, and passes down on the internal side of the thigh, some- what anteriorly; giving off some small branches to the contiguous parts, soon after it originates; and many superficial veins afterwards. It continues along the inside of the knee and leg to the internal ankle, the anterior part of which it passes over. It then proceeds along the internal part of the upper surface of the foot to the middle, when it curves towards the external edge, and joins the lesser saphena. On the GIG Instances of peculiar arrangement of the Veins . leg and foot it also sends off many branches, which anastomose with each other, and with those of the aforesaid vein. The femoral vein, after parting with the saphena, soon sends off the vena profunda, and the circum- flex® also, when they do not arise from the profun- da. These veins are generally larger than the arte- ries to which they correspond, and their branches are more numerous ; but they observe the same course. The great vein accompanies the artery down the thigh and through the perforation in the biceps ; but it changes its relative position, so that it is placed behind or on the exterior side of the artery at the lower part of the thigh. It is very often be- hind it in the ham, where, like the artery, it takes the name of popliteal. In the ham it sends off an- other superficial vein, which seems very analogous to the basilar vein of the arm. This is called The Lesser or External Saphena. It proceeds from the ham over the external head of the gastrocnemius, and down the outside of the leg, sending off many branches in its course. It passes behind the external ankle and near the exterior edge of the upper surface of the foot, about the middle of which it inclines towards the great saphena, and forms with it the anastomosis already mentioned. The popliteal vein, after passing across the arti- culation, ramifies like the artery, but sends two veins, which accompany each of the three arteries of the leg. In a few instances some of the larger veins have been found to be arranged in a manner very differ- ent from that which is commonly observed. Pulmonary Vessels, 317 One case of this kind has already been mentioned in the account of the liver,* where the Vena Porta- rum terminated in the Vena Cava , below the liver, without entering into it. Another very remarkable instance of peculiar ar- rangement is to be seen in a preparation now in the University of Pennsylvania, in which the Inferior Cava, instead of opening into the lower part of the right auricle, passes behind it, in the tract of the Vena Azygos, and opens into the Superior Cava, in the place where the Vena Azygos usually communi- cates with that vessel, receiving the Intercostal Veins in its course. In this preparation, the HepaticVeins communicate directly with the right auricle, at its lower part; the middle and left hepatic veins forming one trunk before they enter, and the right vein passing in singly.f Of the PULMONARY Arteries and Veins, Those portions of the Pulmonary artery and veins which are distinct from the lungs may be described very briefly. It has been already observed , % that the pulmonary artery arises from the left and most anterior part of the basis of the right ventricle, and proceeds thence * See note to p. 132, of this volume, f The foregoing preparation was made by the present editor in 1814, since which two other anomalous cases have occurred to him. 1819, Case 1st. The ascending cava passed into the thorax on the left side of the spine, and getting as far as its upper part, was joined there by the trunk of the internal jugular and subclavian of the left side. It there passed across the vessels of the arch of the aorta and joined with the de» scending cava. The vessels of th'e liver entered the heart at the usual place, in the lower part of the right auricle. 1820, Case 2d. The trunk formed by the junction of the internal jugu- lar and subclavian of the left side instead of taking its usual course, passed down vertically, before the left branch of the pulmonary artery and be fore the left auricle, then making a slight curve between this auricle and the diaphragm joined with the ascending cava.-— E d. t See page 59 oi this volume 3i 8 Pulmonary Vessels. obliquely backwards, inclining gradually to the left side for about eighteen or twenty lines when it divides into two branches, which pass to the two lungs. This course places it under the curve of the aorta : for that great vessel passes over the right branch of the pul- monary artery, and the right side of the main trunk of it, in such a manner that it proceeds downwards between the two branches and behind the angle form- ed by their bifurcation. From this place of bifurcation a short ligament proceeds to the lower part of the curve of the aorta, which is almost in contact with it. This ligament was originally the canal that formed the communication between the pulmonary artery and the aorta of the foetus. Each of the great branches of the pulmonary artery takes a direction backwards, and to its respective side. It soon joins the corres- ponding branch of the trachea and the two pulmona- ry veins, being anterior to the branch of the trachea, and above the pulmonary veins. It is also invested, in common with them, by that portion of the pleura which forms the mediastinum, and thus enters into the composition of the root of the lungs. The Pulmonary Veins are four in number — two on each side. In conformity to the mode of descrip- tion which we have adopted, it may be said that they arise from the sides of the Left Auricle , and proceed nearly in a transverse direction, two of them to each lung ; where they accompany the branches of the artery and of the trachea, being invested by the mediastinum in common with these branches. It has been observed, that they differ from veins in general, by preserving a diameter nearly similar to that of the arteries which they accompany. SYSTEM OF ANATOMY. PART X. OF THE NERVES. The nerves are those whitish cords which pass from the brain and spinal marrow to the various parts of the body. A general account of their origin is contained in the description of the basis of the brain and of the spinal marrow,* which may be considered as intro- ductory to the present subject. The nerves, in general, appear to be bundles or fasciculi of small cords, each of which is composed of a series of fibres that are still smaller. These fibres consist of medullary matter, which is derived from the brain and spinal marrow, and is enclosed in a membranous sheath that appears to arise from the pia mater. The smaller the fibre the more delicate is the membrane which invests it. As the nerves proceed from the brain and spinal marrow, through the foramina of the cranium and the spine, they are enclosed in a sheath formed by the dura mater; but when they arrive at the exterior extremities of the foramina in those bones, this coat, derived from the dura mater, appears to separate into two lamina. The exterior lamen combines with the periosteum, and the interior continues to in- * See volume I. p^ge 33? 320 General Structure of the Nerve. vest the nerve, but seems to change immediately into cellular substance ; so that the exterior coat of the nerves may be regarded as composed of cellular membrane, which is continued from the sheath de- rived from the dura mater. It has been supposed that the membrane which forms the sheaths for the medullary fibrils, of which the nerves are composed, is of a peculiar nature ; but it appears to be derived from the pia mater, in- vesting the brain and the spinal marrow. It is very vascular.* The ramification of a nerve is simply the separa- tion of some fibres from the general fasciculus. The branch commonly forms an acute angle with the main trunk. The course of these branches from their origin to their termination, is generally as straight as possible. When the nervous cords are examined in an ani- mal recently dead, there is an appearance of white lines arranged in a transverse or spiral direction. The cause of this appearance is not well understood. In various parts of the body net works are formed by the combination of different nerves, or the branches of nerves. In those instances the branches of one nerve, unite with those of another, and form new branches. These new branches again divide, and their ramifications unite with other new ramifications to form other new trunks. These new trunks divide again, and form new combinations in the same way. The trunks last formed proceed to the different * Several authors have written professedly on the structure of the nerves, viz. Monro, in his “ Observations on the Structure and Functions of the nervous System.” — Bichat, “ Anatomie Generale.” — Fontana, “ Treatise on the Poison of the Viper.” — Reil, “ Exercitationes Anato- mies.” — Scarpa, “ Annotations Academicse.” — Prochaska, “ De Struc- tura Nervorum.” I regret, that it has not been in my power to procure Reil, Prcchask:!, or Scarpa. 321 Plexuses . — - Ganglions. parts of the body, as other nerves do which arise im- mediately from the brain. These combinations are denominated Plexuses. There are several of them in the cavities of the abdo- men and thorax, formed by the ramifications of the par vagum and the sympathetic nerves. The four lower cervical and the first dorsal nerve form a very remarkable plexus of this kind, which extends from the side of the neck to the axilla, and forms the nerves of the arm. The lumbar nerves form a similar plexus, although not so complex, from which the cru- ral nerve arises. The anterior nerves of the sacrum also unite for the formation of the great sciatic nerve. It appears to be clearly ascertained, that the great object of this peculiar arrangement is the combina- tion of nervous fibres from many different sources, in each of the nerves, which are distributed to any organ. Thus, the smaller nerves of the arm that are distributed to the different parts, are not to be re- garded simply as branches of any one of the five nerves which are appropriated to the upper extre- mity, but as composed of fibres which are derived from each of them. Many of the nerves are enlarged in particular places so as to form small circumscribed tumours, which are denominated Ganglions. These Ganglions are generally of a reddish co- lour. By very dextrous management, they can be shown to consist of a texture of fibres. The larger cords, which compose the nerve, seem suddenly to be resolved into the small fibres, of which they con- sist. These small fibres, after proceeding separate- ly a greater or lesser distance, according to the size of the ganglion, and changing their relative situa- tion, are again combined in cords whieh recompose the nerve. These fibres appear to be surrounded by a fine Vol, ii. 41 322 Structure of Ganglions. cellular substance, which is vascular, moist and soft. It is asserted that, in fat subjects, an oily substance, resembling fat ; and in hydropic subjects, a serous fluid has been found in this texture. Ganglions are often connected with but one nerve, which seems to enter at one extremity and go out at the other. But they frequently receive additional branches from other nerves, and send off additional branches to parts different from those to which their principal nerves are directed. When connected with but one nerve, they have been called simple gang- lions: when they receive and give off additional branches, they are denominated compound gang- lions.— It does not appear that there is any impor- tant difference in their structure in these cases. The simple ganglions occur in the nerves of the spinal marrow— the posterior fasciculus of the nerves having always formed a ganglion before it is joined by the anterior fasciculus. The sympathetic nerve, throughout its whole extent, forms compound gang- lions. The use of this particular structure does not ap- pear to be perfectly known. It seems, however, certain, that the different fibres— (of which the nerves forming ganglions are composed)— are blend- ed together and arranged in a manner different from that in which they were arranged before the nerve entered the ganglion. It ought to be observed, that the combination of nervous fibrillse, so as to bring together those fibrils which originally belonged to different cords, seems to have been kept in view throughout the whole ar- rangement of the nervous system. It is not only in the plexus and the ganglion that this appears, but also in some of the larger nerves ; for in them, the fibres which form the cords that compose the nerve, instead of running parallel to each other, along the Reproduction of Nerves. 323 whole extent of the nerve, form a species of plexus in their course ; separating from the fibres with which they were originally combined, and uniting with the fibres of other cords ; as in other cases of plexus.* There have been doubts respecting the possibility of a reproduction of the substance of the nerves when it has been destroyed ; but it appears to have been clearly proved by the experiments of Mr. Haighton, that a reproduction does really take place.f Nine pair of nerves proceed from the brain through the foramina of the cranium. They are called Nerves of the Brain , or Cerebral Nerves. One pair passes off between the cranium and the spine, which is call- ed Sub-Occipital. Twenty-nine or thirty pair pass through the foramina of the spine : they are deno- minated Cervical, Dorsal, Lumbar , and Sacral , from the bones with which they are respectively connected. There are seven pair of Cervical nerves, twelve Dorsal, five Lumbar, and five or six Sacral — - amounting, with the nerves of the brain, to thirty- nine or forty pair. NERVES OF THE BRAIN The nerves which go off from the brain and me- dulla oblongata are named numerically, according to the order in which they occur ; beginning with the anterior. They also have other names, which generally are expressive of the functions of the dif- ferent parts to which they are distributed. Those which go to the nose are anterior to all the others, and are therefore denominated THE FIRST FAIR, OR THE OLFACTORY NERVES. They arise by three delicate white fibres from the * See Monro’s Observations on the Structure and Functions of the Ner- vous System. Plate xviii. t See London Philosophical Transactions, for 1795, Part I. 324 Olfactory Nerves . under and posterior part of the anterior lobes of the brain, being derived from the Corpora Striata. They proceed forward to the depression on the cribriform plate of the ethmoid bone, on each side of the crista galli. The upper surface occupies a small sulcus formed by the convolutions of the lower surface of the brain, and, therefore, has a longitudinal ridge on it. The lower surface is flat. Their texture is like that of the medullary part of the brain. On each side of the crista galli each of them forms a pulpy enlargement of a brownish colour, which is called the bulb , and has been considered as a gang- lion. From this bulb many fine and delicate cords go off, which proceed through the dura mater and the fora= mina of the cribriform plate to the Schneiderian membrane. — These ramifications of the olfactory nerve seem to receive a coat from the dura mater, as they are much more firm after they have passed through it. They appear to be arranged in two rows as they proceed from the ethmoid bone — one running near to the septum, and the other to the opposite surface of the ethmoid bone."*' THE SECOND PAIR, OR THE OPTIC NERNES, Originate from the Thalami Nervorum Opticorum , and appear on the external and lower surface of the brain, on each side of the sella turcica. Each of them seems like a cord of medullary mat- ter, enclosed in a coat derived from the pia mater, and has not the fasciculated appearance of the other nerves. The medullary matter, however, appears to be divided by processes that pass through it, which are derived from the coat of the nerve. See VoJ- n. p a. 325 Second and Third Pair of JYerves, They proceed obliquely forward, and inward, on each side of the sella turcica, in contact with the brownish cineritious substance, in which the infundi- bulum and the corpora albican tia of Willis are situa- ted.* Anterior to this substance they come in con- tact with each other, and again separate, in such a way, that it is an undecided question whether they decussate each other, or whether each forms an an- gle, and is in contact with the other at the angle. From this place of contact, each nerve proceeds to its respective foramen opticum, where it receives a coat from the dura mater, which extends with it to the eye, as has been described in the account of that organ. THE THIRD PAIR OF NERVES Are sometimes called Motor es Oculorum , in conse- quence of their distribution to several muscles of the eye. They arise at the inside of the crura cerebri, and make their appearance on the basis of the brain^ at the anterior part of the pons Varolii. They originate by numerous threads, which soon unite so as to form a cord, which passes through the dura mater, on each side of the posterior clinoid pro- cess, and continues through the cavernous sinus, and the foramen lacerum, to the orbit of the eye. Before this nerve enters the orbit it generally di- vides into two branches, which are situated one above the other. The Uppermost Branch is spent princi- pally upon the rectus superior muscle of the eye, but sends a twig to the levator palpebrae. The Inferior Branch is distributed to two of the recti muscles, viz. the internus and the inferior, and also to the inferior oblique. It likewise sends a twig to a small ganglion in the orbit, called the Lenticular or Oph - * See VoL I. page 323 ^526 Fourth and Fifth Fair of Nerves. thalmic Ganglion,* from which proceed the fine nervous fibres that perforate the sclerotica coat.f THE FOURTH FAIR OF NERVES Are called the Pathetic, in consequence of the ex- pression of the countenance produced by the action of the muscle on which they are spent. They arise from the side of the valve of the brain, below and be- hind the Tubercula Quadrigeminal and are so small that they appear like sewing thread. They proceed round the crura of the cerebrum, and appear on the surface between the pons Varolii arid the middle lobes of the brain. They proceed along the edge of the tentorium which they perforate, and passing through the upper part of the cavernous sinus, enter the orbit by the foramina lacera. They are exclu- sively appropriated to the Superior Oblique or Troch- learis muscle. THE FIFTH PAIR OF NERVES Are called Trigemina, because each nerve divides into three great branches. These nerves arise from the crura of the cerebel- lum where they unite to the pons Varolii, by distinct fibres, which are connected so as to form a cord or nerve, that is larger than any other nerve of the brain. In many subjects this cord seems partially divided into two portions, the anterior of which is much smaller than the posterior, and appears softer at its origin. It passes into a short canal formed by the dura mater, near the anterior extremity of the petrous por- * This ganglion, which is considered as the smallest in the body lies on the outside of the optic nerve, near its entrance into the orbit, and is ge- nerally surrounded by soft adipose matter, t See Vol I. page 356. S2 7 First Branch of the Fifth Fair . Sion of the temporal bone, at a small distance below the edge of the tentorium. It is perfectly loose and free from adhesion to the surface of this canal ; but it soon passes out of it under the dura mater, and then adheres to that membrane. After leaving the canal it expands like a fan, but still consists of fine fibres which have some firmness. It is said that there are seventy or eighty of these fibres in the expansion, but they appear to be more numerous. Round the cir- cumference of the expansion is a substance of a brown- ish colour, into which the fibres enter. This is the Semilunar Ganglion, or the Ganglion of Gasser , and from it the three nerves go off. These nerves pass off from the convex side of the Ganglion, and are denominated the Ophthalmic, the Superior Maxillary, and the Inferior Maxillary. The Ophthalmic Nerve Passes into the orbit of the eye through the foramen lacerum : it there divides into several branches, which are called, from their distribution, the Frontal or Su- pra Orbitar, the Nasal, and the Lachrymal. The Frontal or Supra Orbitar branch proceeds forward in the upper part of the orbit, exterior to the membrane which lines it, and divides into two ramifications. One of these is small, and passes out of the orbit near the pulley of the superior oblique, to be spent upon the orbicularis muscle and the con- tiguous parts. The other ramification passes through the Supra Orbitary Foramen, or through the notch, which is in the place of that foramen, and divides into a number of twigs, some of which pass transversely towards the side of the head, and communicate with twigs from the portio dura. Most of the others extend up- wards on the head. Some are distributed to the an- 328 Second Branch of the Fifth Pair. terior part of the oceipito-frontalis muscle, and the integuments of the forehead ; others are spent upon the upper portion of the scalp. Some of the extreme parts of these ramifications also communicate with the portio dura. The Nasal Branch proceeds obliquely forward towards the inner side of the orbit, and sends a twig in its course to the lenticular ganglion. It also sends off some small twigs, to join the ciliary nerves which go from the ganglion. On the inside of the orbit a branch leaves it, which proceeds through the Fora- men Orhitare Internum Anterius to the cavity of the cranium, and passes a small distance upon the cribriform plate of the ethmoid bone, under the dura mater, to a fissure in the said plate near the crista galli, through which it proceeds into the cavity of the nose. Here it divides into twigs, some of which pass on the septum near its anterior edge, and ter- minate on the integuments at the end of the nose, while others pass down on the inferior turbinated bone. After parting with the ramification to the nose, the remainder of the nasal branch continues to the in- ternal canthus of the eye, and sends twigs to the lachrymal sac, the caruncula lachrymalis, the eye- lids, and the exterior surface of the upper part of the nose. The Lachrymal Branch proceeds obliquely for- ward and outwards, towards the lachrymal gland. In its course it sends off a twig which passes through the spheno maxillary fissure, and communicates with a twig of the upper maxillary nerve, and one or more twigs that pass to foramina in the malar bone. The main branch passes to the lachrymal gland, and some twigs continue beyond it to the contiguous parts. 829 Second Branch of the Fifth Pair. The Superior Maxillary Nerve. The second branch of the fifth pair is examined with great difficulty on account of its peculiar situa- tion. It proceeds from the semilunar ganglion, and passes through the foramen rotundum of the sphe- noid bone into the upper part of the zygomatic fossa. In this situation it sends a twig to the orbit by the spheno maxillary fissure, and a branch, called the Infra Orbitar, which appears like the main nerve, as it preserves a similar direction, to the infra orbitar eanal. At the same place it sends downwards two branches which* unite together almost immediately af- ter their origin, and, as soon as they have united, en- large into a ganglion.* This ganglion is called the Spheno- Palatine. It is rather of a triangular figure, and lies very near the spheno-palatine foramen. It gives off a posterior braneh, which passes through the pterygoid foramen to the cavity of the cranium : some branches which proceed through the spheno-palatine foramen to the nose, and are called the Spheno- Pala- tine or Lateral JYasal Nerves: and an inferior branch that proceeds through the posterior palatine canal, and is called the Palatine Nerve. The small braneh, which was first mentioned, as going to the orbit by the spheno maxillary fissure, divides into two ramifications. One of them unites with a twig of the lachrymal branch above mention- ed, and passes out of the orbit, through a foramen in the malar bone, to the face ; where it is distributed. The other passes also through a foramen of the ma- lar bone, into the temporal fossa, and, after uniting with twigs from the Inferior Maxillary Nerve , pro- ceeds backwards and perforates the aponeurosis of * Sometimes a single branch passes downwards instead of two ; but it forms a ganglion in the same place. Vol. n. 42 330 Superior Maxillary Nerve. the temporal muscle, to terminate on the integuments of the temporal region. Before the Infra Orbitar branch enters the canal of that name, it sends off two twigs, called Posteriw Dental Nerves , which pass downwards on the tube- rosity of the upper maxillary bone, and enter into small canals in that bone, that are situated behind the Antrum Maxillare. They subdivide into fine twigs that proceed forward to the alveoli of three or four of the last molar teeth ; and penetrate each of the roots by a cavity at its extremity. Twigs also proceed from thege nerves to the posterior part of the gums and buccinator muscle. After the posterior dental nerves have left it, the Infra Orbitar nerve proceeds forwards in the canal of that name ; and near the extremity of it, gives off the anterior dental nerve, which accompanies it for some distance, and then proceeds downwards in a canal in the bone anterior to the antrum maxillare. In its course this nerve divides into many fibres, which pass to the roots of the incisor, canine, and small molar teeth, each in its proper eanal. These dental branches sometimes pass in the antrum maxillare between the lining membrane and the bones. The Infra Orbitar nerve passes out of the foramen upon the cheek, and divides into several branches of considerable size, which are distributed on the face from the side of the nose to the back of the cheek, and also upon the un- der eye-lid and the upper lip. The Pterygoid Nerve , or posterior branch, passes backwards, from the ganglion to a eanal in the base of the pterygoid process of the Os Sphenoides, and proceeds through it. After leaving this canal, it passes through a substance almost as firm as carti- lage, which closes the anterior foramen lacerum, at the basis of the cranium; and divides into two branchy The smallest of them, called the Vidian Superior Moccdlldry Nerve , 331 Nerve , proceeds with a small artery to the small fo- ramen, or Hiatus Fallopii, on the anterior side of the petrous portion of the temporal bone, and continues, through a small canal, to join the Portio Dura of the seventh pair in the larger canal, called the Aqueduct of Fallopius , at the first tarn in that canal.* The other branch of the pterygoid nerve proceeds to the Foramen Carotieum, and passes through it, with a twig of the sixth pair, to join the first cervical gan- glion of the Intercostal Nerve. The Spheno- Palatine, or Lateral Nasal Nerves, consist of several branches which pass from the spheno-palatine ganglion through the Spheno-pala- tine foramen into the nose. Some of them are dis- tributed to that part of the pituitary membrane, which is above the upper meatus, and others to the part which is immediately below it. Some of the branches which thus enter th£ nose are spread upon the septum ; one among them extends upon it, downwards and for- wards to the anterior part of the palatine process of the upper maxillary bone, where it enters into the foramen incisivum, and terminates in a papilla in the roof of the month. f The Palatine Branch proceeds through the canal formed by the upper maxillary and palate bones, to the roof of the mouth and the soft palate. Soon af- ter its origin, it sends off a twig which proceeds down a small canal that is behind it. It also sends off, as it proceeds downwards, several twigs to that part of the membrane of the nose which covers the * The late Mr. John Hunter believed that this nerve parts from the portio dura at the lower end of the aqueduct, and is the chorda tym= pani. f The curious distribution of this nerve appears to have been known to the late John Hunter, and also to Cotunnius; but it is minutely described by Scarpa, and is delineated by Soemmering in his plate of the nose — See “ Observations on certain parts of the Animal Economy,” by J. Hunter, page 219, and also Scarpa “ De Organo Olfactus.” In this last are some interesting observations relative to the ducts of Steno. 332 Third Branch of the Fifth Pair . inferior turbinated bone. When it arrives at the roof of the mouth, it divides into several branches which run forwards, and are distributed to the mem- brane which lines the roof of the mouth. Some of its branches pass to the soft palate, the uvula, and the tonsils; small filaments pass into the back part of the upper jaw. The inferior Maxillary Nerve , or the Third Branch of the Fifth Pair , Passes through the foramen ovale into the zygomatic fossa, and divides into two branches , one cf which sends ramifications to many of the contiguous mus- cles, as the Temporal, the Masseter, the Buccina- tor, the Pterygoid ; and also to the anterior part of the ear and the side of the head. The other branch passes between the pterygoid muscles, and divides into two ramifications, one of which proceeds to the tongue, and is called the Lingual or Gustatory , while the other passes into the canal of the lower jaw. The Lingual Nerve proceeds between the ptery- goid muscles, and in its course is joined by the chorda tympani. It continues forward between the maxillary gland and the lining membrane of the mouth ; and passes near the excretory duct of that gland, above the mylo-hyoideus and the sublingual gland, to the under side of the tongue, near the point : it then divides into a number of branches which enter into that body between the genio-hyoi- deus and lingualis muscles. This nerve has been supposed to be particularly concerned in the function of taste, because many of its branches continue to the upper surface of the tongue, especially near the point. In its course it has a communication with the ninth pair of nerves, and it, sends twigs to the The Sixth Pair of Nerves. 333 membrane of the mouth and gums, and the contiguous parts. After parting with the lingual nerve, the inferior maxillary continues to the upper and posterior orifice of the canal in the lower jaw. Before it enters this canal it sends a branch to the sub-maxillary gland, and to the muscles under the jaw. It then enters the canal, attended by blood vessels, and proceeds along it to the anterior maxillary foramen, on the side of the .chin, through which it passes out. In this course it sends twigs to the sockets of the teeth, and generally supplies all the large and one of the small grinders. Before it leaves the jaw it sends a branch forwards, which supplies the remaining teeth on the side to which it belongs. After passing out, through the an- terior foramen, it is spent upon the muscles and inte- guments of the front of the cheek, the chin, and the under lip. THE SIXTH PAIR OF NERVES Are called Motores Externi. They arise from the commencement of the medulla oblongata, and pro- ceed forward under the pons Yarolii. They proceed through the dura mater on the inside of the fifth pair, and appear to pass through the cavernous sinuses, but are enclosed in sheaths of cellular membrane while they are in those sinuses. When in this situa- tion they are near the carotid arteries, and each nerve sends off one or more very fine twigs, which being joined by a twig from the pterygoid branch of the fifth pair, accompany the carotid artery through the carotid canal, and then unite themselves to the upper extremity of the upper cervical ganglion of the in- tercostal nerve. The sixth pair afterwards pass into the orbit of the eye, each through the foramen lacerum of its respective 334 Composition of the Seventh Pair. side, and is spent upon the Rectus Externus or Ab- ductor muscle of the eye. THE SEVENTH PAIR OF NERVES Comprises two distinct cords which have very differ- ent destinations; and have, therefore, been considered as different nerves, by several anatomists. One of these cords is appropriated to the interior of the ear, and is the proper Auditory Nerve. The other is principally spent upon the face, and, therefore, has been called the Facial. They have, however, more frequently been denominated the Seventh Pair , and distinguished from each other, in consequence of a great difference in their texture, by the appellations of Portio Dura and Portio Mollis. These two cords pass off nearly in contact with each other, from the side of the upper part of the Medulla Oblongata , where it is in contact with the pons Varolii ; but the Portio Mollis can be traced to the fourth ventricle, while the Portio Dura is seen to rise from the union of the pons Varolii with the me- dulla oblongata and the crura Cerebelli. The Portio Dura , at its*origin, is on the inside of the Portio Mollis. Between these cords are one or more small fibres, called Portio Media , which seem to originate very near them, and finally unite with the Portio Dura. Each of the seventh pair of nerves, thus compo- sed, proceeds from its origin, to the Meatus Audito- rius Internus of the temporal bone ; and the Portio Mollis divides into fasciculi, which proceed to the different parts of the organ of hearing, in the man- ner described in the account of the ear.* The Portio Dura enters an orifice at the upper and anterior part of the end or bottom of the Meatus See Vol. I. p. SS6-. Chorda Tymptmi^Pes Ansennus. 33 5 Audiiorius Interims. This orifice is the commence- ment of a canal, which has been called the Aqueduct of Fallopius, and proceeds from the Meatus Audit o- rius Intemus to the external foramen, between the mastoid and styloid processes at the basis of the era- nium. This canal first carves backwards and out- wards, near to the upper surface of the petrous bone,, then forms an acute angle, and proceeds, (back- wards and downwards,) to the stylo-mastoid foramen, passing very near the cavity of the tympanum in its course. The Portio Dura, as it passes into the canal from the meatus internus, seems to receive an investment from the dura mater. It fills up the canal, but does not appear to be compressed.* Near the angle it is joined by the twig of the vidian nerve, whieh proceeds from the pterygoid branch of the fifth pair, and enters the petrous bone by the small foramen innommatum on its anterior surface. In its course through the canal it sends off some very small twigs to the muscles and appurtenances of the small bones of the ear, and to the mastoid cells ; and, when it has arrived almost at the end of the canal, it sends off, in a retrograde direction, a small branch which proceeds into the ca- vity of the tympanum, (entering it by a foramen near the base of the pyramid,) and crosses the upper part of it, near the membrana tympani, between the long processes of the Malleus and Incus. This twig is th eCorda Tympani ; it proceeds from the cavity, by a fissure on the outside of the Eustachian tube, to join the lingual branch of the fifth pair, as has been already mentioned.* The Portio Dura , after passing out of the Fora- * The late John Hunter believed that the chorda tympani is merely a continuation of the twig of the pterygoid branch which joins the portio dura above— See Observations on certain par's of the Animal Econcmv, 220 336 Composition of the Eighth Pair of Nerves* men Stylo -Mastoideum , is situated behind and with- in the parotid gland. Here it gives small twigs to the back of the ear and head, and to the digastric and stylo-hyoideus muscles. It perforates the gland after sending filaments to it, and then divides into branches which are arranged in such a manner that they constitute what has been called the Pes Jlnse- rinus. To describe the various branches in this expan- sion would be more laborious than useful. Some of them are spread upon the temple and the upper part of the side of the head, and unite with the supra-or= bitar branches of the ophthalmic nerve. Some pass above and below the eye, and are distributed to the orbicularis muscle, and communicate with nervous twigs that pass through foramina in the malar bone, &c. Some large branches pass transversely. They cross the masseter muscle, and divide into ramifica- tions which are spent upon the cheek and the side of the nose and lips, and communicate with the small branches of the superior maxillary nerve. A large number of branches pass downwards. Many of them incline forwards, and are spent on the soft parts about the under jaw ; while others pro- ceed below the jaw to the superficial muscles and integuments of the upper part of the neck, communi- cating with the branches of the contiguous nerves.*' THE EIGHTH PAIR OF NERVES Are very frequently denominated the Par Vagum 9 on account of their very extensive distribution. * A most minute and laboured description of the nerves of the face was published by the celebrated Meckel, in the seventh volume of Memoirs of the Royal Academy of Sciences of Berlin, for the year 1751, accompa- nied with a plate, exhibiting the'side of the head, of three times the natu- ral size. This is republished in the Collection Academiaue : Partie Etrang6re.—' Tom. viii. Composition of the Eighth Pair of Nerves. 337 They arise from those portions of the medulla ob- longata which are denominated the Corpora Olivaria . Each nerve consists of a cord, which is anterior, and called the Glosso Pharyngeal ; and of a considerable number of small filaments, which arise separately, but unite and form another cord, the proper Par Fa- gum. Associated with these is a third cord, called the Spinal , or Accessory Nerve of Willis, which ori- ginates in the great canal of the spine, and, passing up into the cavity of the cranium, goes out of it with these nerves through the foramen lacerum. The two first mentioned nerves proceed from their origin to the posterior foramen lacerum, and pass through it with the Internal Jugular vein, — being separated from the vein by a small process of bone. They are also separated from each other by a small process of the dura mater. In the foramen they are very close to each . other ; but soon after they have passed through it, they separate and proceed towards their different destinations. The Glosso -Pharyngeal proceeds towards the tongue, between the stylo-pharyngeus and the stylo- glossus muscles, following the course of the last men- tioned muscle to the posterior part of the tongue. At the commencement of its course it receives a twig from the Portio Dura and one also from the Par Vagum. It soon gives off a branch which passes down on the inside of the common carotid to the lower part of the neck, where it joins some twigs of the intercostal to form the cardiac nerves. Afterwards it sends off several twigs to the muscles of the pharynx and its internal membrane, and also some twigs which unite with others from the upper cervical ganglion of the Sympathetic , and form a network that lies over the anterior branches of the external carotid. The Glosso- pharyngeal finally enters the tongue, at the termina- Vol. ii.' 43 338 Giosso- Pharyngeal Nerve,*— Par Vagum . tion of the hyo-glossns muscle ; and after sending t ranches to the lingualis, and the various muscles in- serted into the tongue, terminates in small ramifica- tions that are spent upon the sides and middle of the root of the tongue, and upon the large papillae. THE PAR VAGUM Are slightly enlarged after passing through the fora- men lacerum. As they descend, they adhere to the superior ganglion of the intercostal, and also to the ninth pair. They proceed behind and on the outside of the carotid, and are contained in the same sheath of cellular membrane which encloses that artery and the internal jugular vein. Each of these nerves, soon after it leaves the cranium, gives a twig to the glosso- pharyngeal ; that soon after it sends off a branch called the Pharyngeal, which unites to one from the acces- sory nerve, and to one or more from the glosso-pha- ryngeal, and proceeds to the middle constrictor of the pharynx, when it expands into ramifications that form a plexus from which proceed a number of small twigs that go to the larynx, and some that pass down on the common carotid artery. It then sends off, downward and forward, the Su- perior Laryngeal nerve, which continues in that di- rection behind the carotid artery, and divides into an external and internal branch. The Internal Branch , which is the largest, pro- ceeds between the os hyoides and the thyroid carti- lage ; and divides into numerous ramifications which are distributed to the arytenoid muscles and to the membrane which lines the larynx and covers the epi- glottis. It is said, that fine twigs can be traced into the foramina, which are to be seen in the cartilage of the epiglottis ; — some ramifications can be traced to the pharynx ; — others communicate with the branches of the recurrent nerve. Superior Laryngeal Branch of the Par Vagum . 339 The External Branch sends twigs to the pharynx, to the lower and inner part of the larynx, and to the thyroid gland. In its course downwards, the great nerve some- times sends off a twig, which unites with one from the ninth pair that passes to the sterno-hyoidei and sterno-thyroidei muscles. It uniformly sends off one or more twigs, which pass into the thorax and combine with small branches from the sympathetic or intercostal nerve, to form the Cardiac plexus , which sends nerves to the heart. After entering the thorax, the right trunk of the Par Vagum passes before the subclavian artery ; and the left trunk before the arch of the aorta ; and im- mediately after passing these arteries, each of the nerves divides into an interior and posterior branch. The anterior is the continuation of the Par Vagum ; the posterior is a nerve of the Larynx; which, from its retrograde course, is called the Recurrent Nerve. On the left side the Recurrent Nerve winds back- wards round the aorta, and on the right side round the subclavian artery, and proceeds upwards, deeply seated, on the side of the trachea, to the Latnjnx. Soon after its origin it sends filaments to a ganglion of the sympathetic, to the cardiac plexus, and to a pulmonary plexus soon to be mentioned. In its course upwards it sends twigs to the trachea and the oesophagus. It proceeds behind the thyroid gland, and sends twigs to that organ. At the lower part of the larynx it sends off a branch which communicates with branches of the superior laryngeal nerve. It also divides into branches which are spread upon the posterior crico-arytenoid, and the arytenoid muscles : and also upon the lateral crico-arytenoid and the thvro-arytenoid muscles, as well as upon the mem- 340 Recurrent Branch of the Par Vagum. brane which lines the hack part of the larynx and the contiguous surface of the pharynx. There is a difference in the arrangement of the re- currents on the different sides in consequence of one winding round the aorta, while the other winds round the subclavian artery. After sending off the recurrents, each trunk of the par vagum proceeds behind the ramifications of the trachea ; but previously detaches some small branches, which are joined by twigs from the intercostal and from the recurrent, and form a plexus upon the an- terior part of the vessels going to the lungs. This Anterior plexus, after sending off some minute branches to the cardiac nerves and the pericardium, transmits its branches with the bronchia and the blood vessels, into the substance of the lungs. Some of the branches which proceed from the par vagum, pass down on the posterior part of the tra- chea, and enter into the membrane which forms it, and the mucous glands which are upon it ; and some pass to the oesophagus. When the par vagum is behind the great vessels of the lungs, a number of branches go off transverse- ly, and are also joined by some fibres from the sym- pathetic. These form the Posterior pulmonary plexus ; the ramifications from which proceed into the substance of the lungs, and are principally spent upon the ramifications of the bronchia. It has been said,* that the small twigs into which they divide, very generally penetrate into the small ramifications of the bronchia, and are spent upon their internal membrane. Soon after sending off the nerves of the pulmona- ry plexus, the Par Vagum proceed dow’nwards * See Buisson, in the continuation of the Descriptive Anatomy of Bichat. Different Functions of the Laryngeal , fye. 341 upon the oesophagus; the left nerve being situated anteriorly, and the right posteriorly. Each of these nerves forms a plexus so as nearly to surround the oesophagus, as they descend on it; but the net- work is thickest on the posterior side. They pass through the diaphragm with the (esophagus, and unite again so as to form considerable trunks. The Anterior , which is the smallest, proceeds along the lesser curvature of the stomach to the pylorus. Some of its fibres are spread upon the anterior side of the stomach and the lesser omentum. Others of them extend to the left hepatic, and also to the solar plexus. The Posterior trunk sends branches to surround the cardiac orifice of the stomach. Many branches are spread upon the under side of the great curvature of the stomach. Some of them pass in the course of the coronary artery to the cceliac, and unite to the hepatic and splenic plexuses; and one trunk, which is thick, although short, proceeds to the solar plexus. The Accessory Nerve of Willis , Which has been mentioned as associated with the eighth pair of nerves, within the cranium, has a very peculiar origin. It arises by small filaments, which come off from the spinal marrow, between the anterior and posteri- or fasciculi of the cervical nerves, and proceeds up- wards to the great occipital foramen, between these fasciculi. It commences sometimes at the sixth or seventh cervical vertebra, and sometimes about the fourth. It enters the cavity of the cranium through the foramen magnum, and proceeds upwards and out- wards, so as to join the eighth pair of nerves at some distance from its origin, and in this course it receives filaments from the medula oblongata. After approaching very near to the eighth pair of 342 JYinth Pair of jYerves. nerves, it accompanies it to the foramen lacerum, and passes out in its own separate sheath. It then leaves the eighth pair and descends towards the shoulder, proceeding through the sterno-mastoid muscle. Soon after it emerges from the cranium, it sends a ramifi- cation to the pharyngeal branch of the Par Vagurn and another to the par vagum itself. After passing- through the upper and back part of the sterno-mastoid muscle, it terminates in the trapezius. It adheres to the ninth pair of nerves as it passes by it, and sends a twig to the sub-occipital and some of the cervical nerves. It also gives ramifications to the sterno-mastoid muscle as it passes through it. It has already been stated that the Laryngeal and Re- current Nerves appear to answer different purposes in their distribution to the Larynx. — When both of the recurrent nerves are divided in a living animal, the voice seems to be lost. When the laryngeal nerves only are divided, the strength of the voice remains, but it is flatter. The recurrent nerves, therefore, seem essential to the formation of the voice. The laryngeal nerves are necessary to its modulation. The history of the investigation of this subject is contained in Mr. Haighton’s paper in the third volume of Memoirs of the Medical Society of London. THE NINTH PAIR OF NERVES. Each of these nerves arises from the groove in the medulla oblongata, between the corpora pyramidalia and the corpora olivaria. Three or four fasciculi, of distinct filaments, unite to form it. Thus composed, it proceeds to the anterior condyloid foramen of the occipital bpne, and passes through the dura mater. It seems firmly united, by the cellular membrane, to the eighth pair, and to the first ganglion of the sym- pathetic, soon after it passes from the occipital bone. It is either connected to the sub-occipital nerve by a small ramification, or it joins a branch which pro 343 Ninth and Tenth Pair of Nerves . eeeds from the sub-occipital to the cervical, and bends round the transverse process of the atlas. It passes between the internal carotid artery and the internal jugular vein, and crosses the external caro- tid at the origin of the occipital artery. At this place it generally sends downwards a large branch which is called the Descendens Noni. Passing for- wards, it is on the outside of the posterior portion of the digastric muscle, and inclines downwards ; but near the tendon of the muscle it turns upwards, and proceeds on the inside of the mylo-hyoideus, where it divides into ramifications, which, at the anterior edge of the hyo glossus muscle, begin to enter into the substance of the tongue, between the genio-glos- sus and the lingualis muscles. Some of the branches of this nerve unite with those of the lingual branch of the fifth pair. Others are distributed to almost all the muscles connected with the tongue. The branch called descendens noni passes, down in the course of the common carotid artery, and sends branches in its progress to the upper portions of the eoraco-hyoidei and sterno- thy roidei muscles; it unites with ramifications of various sizes from the first, second and third cervical nerves, which form a bow under the sterno- mastoid muscle, from which ramifications go to the lower portions of the sterno- byoidei and thyroidei muscles and of the coraco- hyoidei. OF THE CERVICAL NERVES. The tenth or last pair of the head, commonly call- ed the Sub occipital , may be arranged with these nerves, because they arise* like them, from the me- dulla spinalis, and are distributed to the muscles on the neck. 344 Siib-Oceipitul or , Tenth pair of Serves, The sub-occipital JYet'vcs Arise on each side of the spinal marrow, nearly op- posite to the interval between the great foramen of the os occipitis and the atlas. Each of these nerves consists of an anterior and posterior fasciculus, or bundle of fibres, which pass outwards immediately under the vertebral arteries, and form a ganglion, from which proceeds an ante- rior and a posterior branch. The anterior branch is united to the second cervi- cal nerve below, and to the ninth nerve, or the hy- po-glossal,. above. It also sends filaments to the up- per ganglion of the great sympathetic nerve. The posterior branch is spent upon the Recti> the Obiiqui, and some other muscles of the head. The proper Cervical Nerves consist of seven pair; of which the first six go off between the vertebrae of the neck, and the seventh between the last of the neck and the first of the back. The first cervical JYerve Passes out between the atlas and the Vertebra Den - tata. It originates from two fasciculi, which are connected to each other at a ganglion, and then sepa- rate into an anterior and a posterior branch.* The anterior branch is connected by filaments with the accessory nerve, with the ninth pair of the head, and with the upper ganglion of the sympathetic. It is also connected with the second cervical nerve ; and sends some branches to the muscles on the anterior part of the spine. The posterior branch, after communicating with the posterior branches of the sub-occipital and the * This arrangement is common to the nerves of (he spine. The ganglion is formed by the posterior fasciculus. Cervical and Phrenic JVerves, 345 second nerves of the neck, perforates the complexus muscle, and ascending upon the back of the head, is distributed with the occipital artery. THE SECOND CERVICAL NERVE Sends off, from its Anterior Branch , a twig which descends to the low r er cervical ganglion of the sympa- thetic, and a considerable ramification to the third cer- vical nerve. It also sends off some twigs to the sterno- mastoid muscle, and others to join the accessory nerve. Some of its small ramifications pass down upon the external jugular vein, and others unite with the de- scending branch of the ninth pair of the head. A small branch is also concerned in the formation of the phrenic nerve. Two larger branches of this nerve wind round the posterior edge of the sterno-mastoid, and are spread under the integuments of the anterior, lateral and posterior parts of the neck and lower parts of the head; they have a communication with the portio dura of the seventh pair.*— The posterior branch of this nerve is spent upon the extensor mus- cles of the head and neck. The THIRD CERVICAL NERVE Sends down, from its Anterior Branch , the princi- pal trunk of the phrenic nerve. It also sends twigs to the fourth cervical, to the lower cervical ganglion of the intercostal, and to the descending branch of the ninth of the head. Some of its branches, unite with twigs of the accessory nerve, and others are spent upon the muscles and integuments of the shoul- der and lower part of the neck. A small Posterior Branch is spent upon the muscles of the back of the neck. * These superficial branches have sometimes been described as coming from a plexus ; but they often arise directly from the Second Cervical nerve. VOL, If, 44 346 Cervical JVerve$, THE NERVES OF THE DIAPHRAGM Are generally denominated the Phrenic. The prin- cipal root of each of them is commonly derived from the third cervical nerve, but frequently the second and the fourth cervical nerves contribute to the forma- tion ; and they are sometimes joined by a twig which is derived from the ninth pair. Each nerve proceeds down the neck, between the rectus capitis major and the scalenus anticus, and continues along the fore part of the scalenus anticaS; it descends into the thorax within the anterior end of the first rib, between the subclavian vein and the ar- tery. It sometimes receives a twig from the fifth cer- vical nerve, and a twig passes between it and the great sympathetic. After entering the thorax, they descend, attached to the mediastinum, before the root of the lungs. In consequence of the projection of the point of the heart to the left, the course of the left is a little different from that of the right; that of the right pro- ceeding in a more perpendicular direction. When they arrive at the diaphragm, they divide into many ramifications, which have a radiated arrangement, and terminate on the fibres of that muscle, both on the upper and lower surface. Some fibres from each nerve are continued downward, and communicate in ^ the abdomen with fibres from the intercostal, THE FOURTH, FIFTH, SIXTH, AND SEVENTH CERVI- CAL NERVES, May be comprised in one description. They pass off successively from the Medulla Spinalis* between the vertebrae, like the other nerves. Their Posterior Branches are generally distributed to the back of the neck, and are very small. Their Anterior Branches are principally appropriated to the upper extremi- ties, and are large. They generally send each a Remaining Cervical Nerves.- —^Brachial Plexus . 347 small twig to the lower cervical ganglion of the inter- costal nerve, and a few small branches to some of the contiguous muscles. They are arranged and com- bined so as to form the net-work, now to be described, which is called the Brachial or Axillary plexus ; and, in the formation of this plexus, they are joined by the first dorsal nerve. The BRACHIAL PLEXUS Extends from the lower part of the side of the neck, into the arm-pit. It commences in the following manner. The fourth and fifth cervical nerves pro- ceed downwards, and after uniting to each other about an inch and a half below their egress from the spine, they separate again, almost immediately, into two branches. The sixth cervical nerve, after passing downwards, divides also into two branches, one of which unites with the uppermost branch that proceeds from the union of the fourth and fifth, and the other with the lowermost, and they all proceed downwards. The seventh cervical is joined by the first dorsal, which proceeds upwards, and unites with it at a short distance from the spine. The cord produced by their junction soon unites with one of the cords above described. As these different cords proceed down- wards, they divide, and their branches again unite. The axillary artery, which passes in the same direc- tion, is surrounded by them. In this manner the axillary plexus is often formed. The muscles about the shoulder, both before and behind, are supplied by the axillary plexus. Thus, it sends branches to the Sub -scapular is, Teres Major , and Latissimus Dorsi, behind; and to the Pect ora- lis Major and Minor , and the Mamma, before. It also sends off a branch called the Scapularis , which 348 Nerves of the Arm . commonly arises from the upper part of the plexus, and proceeds through the notch in the upper costa of the scapula, to the supra and infra spinatus, teres minor, &c. Nerves of the Arm. All the great nerves of the arm are derived from the axillary plexus. There are six of them, which are denominated The Musculo Cutaneous ; the Me- dian The Cubital , or Ulnar ; The Internal Cu- taneous ; The Radial or Muscular Spiral ; and the Circumflex or Articular. The Musculo Cutaneous, or Perforating Nerve , passes obliquely through the upper part of the coraco brachialis muscle. Before it enters the muscle, it sends a branch to it. After leaving the muscle, it passes down the arm between the biceps and the brachialis internus, to which it also gives branches. It proceeds to the outside of the biceps, and conti- nues under the median cephalic vein to the anterior and external part of the fore -arm ; along which it passes, under the integuments. On the lower part of the fore-arm it divides into many branches, which extend to the root of the thumb and the back of the hand, and terminate in the integuments. The Median Nerve, which is one of the largest of the arm, often proceeds from the axillary plexus next to the musculo cutaneous ; it passes dow n the arm, very near the humeral artery, within the edge of the biceps flexor muscle, and, during this course, gives off no branches of any importance. After passing the bend of the elbow, it proceeds, under the apo- neurosis of the biceps, between the brachialis inter- ims and the pronator teres, and continues down near Sometimes called Radial. Median and Cubital or Ulnar Nerves. 349 the middle of the fore-arm, between the flexor subli- mis and the flexor profundus. At the elbow it sends branches to several muscles on the anterior side of the fore-arm, and to the integuments. Among these branches is one, called the Interosseal Nerve, which passes down on the anterior surface of the interos- seal ligament, with the artery of that name. This nerve sends branches, in its course, to the long flexor of the thumb and the deep flexor of the fingers. When it arrives at the pronator quadratus, it sends branches to that muscle, and, passing between it and the interosseous ligament, perforates the liganient, and soon terminates on the posterior side of the wrist and hand. As the Median Nerve proceeds downwards, it be- comes more superficial ; and continuing among the tendons of the flexors of the fingers, it gives off a branch which is principally spent upon the integuments of the palm of the hand. This great nerve passes with the tendons under the annular ligament ; and imme- diately after, while it is covered by the Aponeurosis Palmaris, and by that portion of the artery which is called Arcus Sublimis, it divides into branches, which separate from each other at acute angles, and subdi- vide so as to send a ramification to each side of the thumb, of the index, and of the middle finger ; and to the radial side of the ring finger. The Cubital or Ulnar Nerve is also of considera- ble size. It passes down on the inside of the triceps extensor muscle, to the great groove formed by the olecranon process and the internal condyle of the os humeri ; and in this course it often sends a branch to the triceps, and some smaller twigs to the upper part of the fore -arm. From the groove it proceeds on the anterior part of the fore-arm, between the flexor carpi ulnaris and the flexor sublimis, to the wrist. At a small distance above the wrist it sends off a branch, 350 Radial, or Muscular Spiral Nerve. called the Dorsalis, which passes between the flexor ulnaris and the ulna, to the back of the fore-arm and wrist, where, after sending ramifications to the inte- guments and contiguous parts, it divides into branches which pass to the little finger and the finger next to it. Those branches send off, in their course, many twigs which pass to the skin and cellular sub- stance. The ulnar nerve then proceeds with the artery, over the annular ligament, on the radial side of the os pisiforme, and divides into two branches ; one of which is superficial, and the other deep-seated. The Superficial divides into two principal branches, an external and an internal. The external passes under the aponeurosis palmaris ; and, after sending a branch to combine with one from the median, and some twigs to the contiguous muscles, it subdivides into two branches, one of which goes to the ulnar side of the ring finger and the opposite side of the little finger. The other branch sends off some twigs to the muscles, and proceeds along the ulnar side of the little finger. The Deep-seated palmar branch of the ulnar nerve, passes between the muscles of the little finger, under the tendons of the flexors, and accompanies the deep- seated arterial arch in the palm of the hand, giving branches to the interossei, and other contiguous mus- cles. The Radial or Muscular Spiral nerve is one of the largest nerves of the arm. It passes from the axillary plexus downward, backward and outward, under the triceps muscle, to the external side of the os humeri. In this course it gives off several branches to the different portions of the triceps. It also fre- quently gives off a large branch, which passes down- wards on the outside of the olecranon, to the back of the fore-arm. and continues to the back of the Radial , ar Muscular Spiral Nerve. 351 hand, furnishing many branches which terminate in the integuments. It then proceeds downwards be- tween the supinator radii longus and the brachialis interims. Immediately after passing the articulation of the elbow, it divides into two branches, denomi- nated the Superficial and the , Profound. The Su- perficial soon joins the radial artery, and proceeds downwards, sending branches to the contiguous mus- cles. In its course about the middle of the arm, it crosses the tendon of the supinator longus, and pro- ceeds between it and the tendon of the extensor carpi radialislongior ; itsoon after divides the two branches, which are principally distributed to the thumb and fore finger, and also to the integuments. The Profound branch proceeds to the back of the fore-arm under the radial extensor, and continues to the back of the wrist and hand. Into this course it divides into two branches, which are distributed to the contiguous muscles and tendons, and the inte- guments. The Internal Cutaneous nerve is the smallest of the nerves which proceed from the axillary plexus. It descends in the course of the basilic vein, and very near it. Above the elbow it divides into an Internal branch, which proceeds over the Basilic Vein, and separates into branches that pass down on the side of the fore-arm ; and an External Branch that passes under the Median Basilic Vein, and con- tinues down on the anterior part of the fore-arm. The Articular or Circumflex nerve proceeds backwards from the plexus, between the teres major and minor, and passes nearly around the body of the os humeri, at a small distance below its head. It is distributed to the contiguous muscles and to the arti- culation ; but its principal branches terminate in the deltoid muscle. 352 The Dorsal JYerves. THE DORSAL JYERVES Proceed from the cavity of the spine between the dorsal vertebrae. They are sometimes called Inter - costals, because they pass between the ribs, like the blood vessels of that name. There are twelve pair of them, and they are named numerically, beginning from above. These nerves proceed from the medula spinalis by two fasciculi of fibres — one from each of its late- ral portions — the posterior fasciculus is the largest. After passing through the lateral foramen and the dura mater, a ganglion is formed by the posterior fasciculus : the anterior fasciculus unites to this gang- lion at its external extremity: and one nerve is formed, which almost immediately divides into an anterior and a posterior branch, of which the ante- rior is the largest. The posterior branch proceeds backwards, and is distributed to the muscles of the back. The anterior branch passes towards the angle of the rib, in con- tact with the pleura. Soon after its origin, this an- terior branch sends off two ramifications which unite to the intercostal nerve, at the ganglion ; it then pro- ceeds forwards with the blood vessels, between the internal and external intercostal muscles, in the groove near the lower margin of the ribs ; and ter- minates on the anterior part of the thorax. In its course it sends branches, not only to the intercostal muscles and pleura, but to the other muscles and the integuments of the thorax. Some of the dorsal nerves differ from the others, as to the ramifications which they send off. The first nerve , of this order, joins the lower cer- vical nerves in the axillary plexus ; but it sends off the ramifications to the sympathetic ; and also a General Account of the Lumbar Nerves. 353 branch, which passes under the first rib, like the other dorsal nerves. The second nerve, sends off a branch, which passes through the external intercostal muscle into the ax- illa, and combines there with a branch of the cuta- neous nerve, being distributed to the internal and posterior part of the arm. The third dorsal nerve also sends off a branch, which is distributed to the axilla and the back part of the arm. These branches of the second and third dorsad nerves, are called intercosto-humeral nerves. The lower dorsal nerves supply the muscles and integuments of the abdomen. Of the L UMBAR Nerves . There are five pair of these nerves. The first of them passes off between the first and second of the lumbar vertebras, and the others succeed regularly ; so that the last pair is situated between the last lum- bar vertebra and the sacrum. The first lumbar nerves arise from the medulla spinalis, before it forms the cauda equina ; the other four pair are formed by the cauda equina. They commence by anterior and posterior fascicu- li, which are united at a ganglion. From these gang- lion, anterior and posterior branches go off, which are very different in size, the anterior being the largest. The posterior branches are distributed to the mus- cles of the back. The anterior sends branches to the ganglions of the sympathetic nerve, and also commu- nicate with each other to form the Lumbar Plexus , which is situated on the lateral parts of the bodies of the Lumbar Vertebrae, before their transverse pro- cesses, and supplies nerves to the muscles of the thigh. Vol. II, 45 354 Lumbar JYerves. —Lumbar Plexus . THE FIRST LUMBAR NERVE Is connected, by its anterior branch, to the last dor- sal and the second lumbar. From the same branch, ramifications go off to the Quadratus Lumborum, and obliquely across that muscle, to the lower part of the abdominal muscles near the spine of the ilium. THE SECOND LUMBAR NERVE Sends off a muscular branch downwards and out- wards : it also sends off the small branch, called the External Spermatic, which passes down in such a direction, that it perforates the transversalis and the obliquus internus muscles, near their lower margin, at a small distance from the superior anterior spine of the ilium, and then proceeds within the lower edge of the tendon of the external oblique to the abdomi- nal ring, through which it passes. In the male it is distributed to the spermatic cord and scrotum, and in the female, to the labia pudendi. In the female it also sends a branch to the uterus.* The Second Lumbar, after sending off these branches, passes downwards, and joins the Third lumbar nerve. From this union of the second and third nerves, a branch called the Cutaneus Medius, which will be soon de- scribed proceeds downwards. After sending off this branch, the united trunk of the second and third joins the Fourth ; and from this union is sent off the Obturator Nerve, which passes through the aperture in the membrane that closes the foramen thyroideum ; the Crural Nerve, which passes under Poupart’s ligament ; and a third branch that proceeds downwards, and joins the Fifth lumbar nerve. The Fifth lumbar nerve, with this acces- sion from above, descends into the pelvis, and unites with the sacral nerves. The external spermatic often comes off from the first lumbar nerve 355 Obturator Nerve.— Crural Nerve . This arrangement of the lumbar nerves constitutes the Lumbar Plexus , which, as has been already sta- ted, furnishes three nerves to the lower extremity, viz. the Cutaneus Medius , the Obturator , and the Crural Nerve . The Cutaneus Medius , which arises from the union of the second and third nerves, as has been already observed, proceeds downwards, and fre- quently adheres to the crural nerve, for a short dis- tance, near Poupart’s ligament, but soon leaves it, and descends on the inside of the thigh, supplying the integuments as low as the knee. THE OBTURATOR NERVE Descends into the pelvis, and passes out of it at the upper part of the foramen thyroideum ; proceeding downwards in an internal direction, to be distributed on the inside of the thigh. This nerve is generally accompanied by the obtu- rator artery and vein ; the artery being above, and the vein below it. When it has arrived at the fora- men ovale or thyroideum, it sends off a branch to the internal and external obturator muscles, and, after passing these muscles, divides into two branches which are distributed to the muscles on the inside of the thighs, the adductors, the pectineus, the graci- lis, &c. THE CRURAL NERVE Is situated at first behind, and then on the outside of the psoas muscle. It passes under Poupart’s liga- ment with the great femoral vessels, being on the outside of the artery. It is distributed to the integuments, and also to the muscles, which are situated on the anterior and internal parts of the thigh. Some of its ramifications go off before it passes under Poupart’s ligament* 336 Crural Nerve. — Sacral Nerves. Several of them are spent upon the integuments, and are therefore denominated Cutaneas. — They are distinguished by the terms Cutaneus Anterior , Cu- taneous Interims, &c. according to their situations. The deep-seated branches are the largest. They are principally spent upon the muscles on the ante- rior and the internal side of the thigh, viz. the four extensors, the adductors, the pectineus, the sar tori us, and the gracilis. Among these nerves there is one, called the Saphenus, which has a different destina- tion. It accompanies the great artery of the thigh to the place where it perforates the Adductors : it there separates from the artery, and passes over the tendon of the Adductors, under the sartorius muscle ; thence it continues, with the great saphena vein, on the in- side of the leg, to the internal ankle ; sending branches to the integument, in its course. It terminates in skin and cellular substance on the upper and internal sur- face of the foot. The SACRAL Nerves Are composed of those cords of the cauda equina, which remain after the formation of the lumbar nerves. They are frequently stated to consist of five or six pair, four of which pass through the fo- ramina of the sacrum, and the fifth between the sa- crum and the os coccygis.* The cords of which they are respectively composed arise by anterior and posterior fasciculi. When they have arrived oppo- site to the foramina of the sacrum, through which they are to pass, a ganglion is formed, at which they unite, and then divide into anterior and posterior branches.! The uppermost of the anterior branches * The sixth pair, when they exist, ’proceed in a groove in the os coccygis. t The ganglions of the fourth and fifth nerves are extremely small and not so near the foramina as those of the others. Sciatic Plexus.— Pudic Nerve. 337 are large and pass through the anterior foramina of the sacrum. The posterior are small, and go through the posterior foramina. The Posterior branches are generally spent upon the muscles which lie on the sacrum, and posterior parts of the pelvis, externally. The anterior branches of the. three first nerves send ramifications to the sympathetic. They unite to each other, and are joined by the last lumbar nerve, and by a branch of the fourth sacral, in the forma- tion of the great sciatic nerve. This union constitutes the Sciatic plexus. The anterior branch of the fourth nerve transmits branches to the sympathetic: it also sometimes sends a branch to the united nerves above, or the sciatic plexus. It sends branches to the hypogastric plexus, and to the contiguous muscles. The fifth and sixth pairs, which are very small, terminate also in the contiguous muscles and in the integuments. From the Sciatic plexus , or the nerves which com- pose it, several smaller branches go off. There are generally two which pass off backwards through the ischiatic notch, and are denominated Gluteal, as they are distributed to the glutei muscles. From the low- ermost of these a branch descends on the thigh. The Pudic nerve, which is appropriated to the or- gans of generation, also passes off from this plexus, and appears to consist of fibres which are derived from each of the nerves that compose it. It proceeds between the sacrosciatic ligaments, and divides into two branches — an inferior and a superior. The in- ferior passes between the erector penis and the ac- celerator urinse muscles, and is distributed to those muscles, to the bulb of the urethra and the interior of that canal, to the scrotum and dartos. The superior proceeds along the os pubis to the U5& Course of the Great Sciatic Nerve. symphisis, and passes between the bone and the body of the penis to the dorsum. A considerable branch accompanies the artery on the dorsum, and terminates, by many ramifications, on the glans penis; after send- ing branches in its course to the integuments gene- rally, and to the prepuce. In females, the Inferior pudic nerve proceeds along the external labia pudendi to the mons veneris, send- ing off many ramifications in its course. The Superior pudic nerve proceeds, as in males, along the branch of the pubis to the superior surface of the clitoris, and terminates principally upon the extremity of that organ. The sacral nerves unite in the sciatic plexus to form the great nerve of the lower extremity, which is next to be described. The GREAT SCIATIC JYerve Proceeds from the pelvis through the ischiatic notch, between the pyramidalis and the superior gemellus muscle : it then passes down to the back part of the thigh, between the tuberosity of the ischium and the great trochanter of the os femoris ; and continues downwards, inclining from within outwards, to the ham, where it is situated between the tendons of the semi-tendinosus and semi-membranosus on the inter- nal side, and the tendon of the biceps on the external. In this course it sends off branches to the muscles on the posterior part of the thigh. As the great nerve passes down the thigh, it sends off obliquely downwards and outwards, a large branch which is called the Fihular , that passes across the head of the fibula to the external and anterior part of the leg. The place where this branch separates from the main nerve is different in different subjects. It continues in contact with it Distribution of the Fibular Nerve » 359 for some distance, connected only by cellular mem- brane. THE FIBULAE Oil PERONEAL NERVE Proceeds downwards on the inside of the tendon of the biceps, and crosses obliquely to the outside of the external head of the gastrocnemius: it then passes inwards between the long peroneus muscle and the fibula; and descending between the muscles on the front of the leg, divides into two branches, one of which inclines to the exterior side of the leg, and the other preserves an internal situation. In its course from the great sciatic nerve to the fibula, it sends off some superficial ramifications. The two branches into which it divides, after passing over the fibula, continue downwards. The Internal , after supply- ing the muscles on the anterior part of the leg, passes under the annular ligament like the anterior tibial artery; and on the upper part of the foot, di- vides into two ramifications, one of which proceeds forwards near the internal edge of the foot, and the other near the external ; they divide again, and are distributed to the parts on the upper surface of the foot, one of their ramuli descending with the conti- nuation of the anterior tibial artery to the sole of the foot. The External Branch of the fibular nerve, as it proceeds downwards, supplies ramifications to the contiguous muscles, and passing through the fascia on the outside of the leg, continues between it and the skin towards the foot. In this course it gene- rally divides into two branches which are spent upon the upper surface of the foot. The GREAT SCIATIC Nerve , after the fibular nerve leaves it, continues down the thigh, between the tendons of the flexors, behind the great blood vessels, and of course exterior to them. 360 Distribution of the Tibial Nerve. In the ham, this great nerve takes the name of POPLITEAL, and proceeds across the articulation of the knee, between the heads of the gastrocnemii, to the posterior side of the tibia : here it passes through the upper portion of the soleus or gastroc- nemius internus, and continues between it and the long flexor of the toes, near the Posterior Tibial Ar- tery ; descending with that artery to the hollow of the os calcis. In this situation it has the name of POSTERIOR TIBIAL NERVE. At the commencement of this course, a small dis- tance below the internal condyle of the os femoris, it sends off a branch of considerable size called the Communieans Tibas, or Saphena Externa, which passes down behind the gastrocnemii, and gradually inclines externally, so that it is situated on the ex- ternal edge of the tendo Achillis, soon after the com- mencement of that tendon, and proceeds behind the external ankle, near the outer side of the foot, to the smaller toes: distributing branches to the contiguous parts. In its course on the back of the leg, it sends off a branch which unites with one of the superficial ramifications of the fibular nerve, and descends to the outer part of the foot. The Tibial Nerve, in its course downwards, sends branches to the contiguous muscles ; and a few twigs which form a species of net-work on the artery. In the hollow of the os calcis it sends off a superficial branch to the integuments of the sole of the foot, which proceeds on the outside of the aponeurosis plantaris: it there also divides into branches, which are denominated the Internal and External Plantar Nerves. The Internal Plantar Nerve proceeds forwards, along side of the tendon of the long flexor muscle of the great toe, givin'g off small branches in its course. Commencement of the Sympathetic Nerve, 361 About the middle of the foot it divides into four branches, one of which proceeds to the inside of the great toe ; and a second to the angle formed by thb great toe and the toe next to it, where it divides and sends a branch to the opposite sides of those toes : the other two branches are distributed in a similar manner, to the succeeding toes. These digital branches are connected with each other by small ramifications. The External Plantar Nerve proceeds with the external plantar artery towards the externel side of the foot, between the short flexor of the toes and the flexor accessorius. Near the external edge of the foot, about the posterior end of the metacarpal bones, it divides into three branches. One proceeds to thd outside of the little toe; another passes to the angle between the fourth toe and the little toe, and divides into branches which are distributed to the correspond- ing sides of these toes. The third branch proceeds more deeply in the foot, from the external towards the internal edge of it, and is spent upon the deep- seated contiguous muscles. THE GREAT SYMPATHETIC OR INTERCOSTAL NERVE Commences in the cranium with those small ramifica- tions of the pterygoid branch of the upper maxillary nerve, and of the sixth pair, which accompany the carotid artery through the canal in the petrous portion of the temporal bone. These small nerves from a net-work which surrounds the artery in the canal, and gives rise to the incipient sympathetic, a small cord which passes down close- to the nerves of the eighth and ninth pair of the neck. Opposite to the second cervical vertebra, this nerve is swelled or dilated, so as to form a body of a light red colour,, which is more than an inch in length, and has the form of two cones united to each other at their bases. This is the Su- perior Cervical Ganglion of the Sympathetic Nerve* Vol. ii, ^46 " 362 First Ganglion, and other Cervical and from it tlie nerve descends ? behind the Par Va- guin, on the front part of the neck. This ganglion receives twigs from the first, second, third and fourth pairs of cervical nerves, and also from the eighth and ninth nerves of the head. It sends off several twigs, which pass behind the carotid artery, at its bifurcation, and are joined by twigs of the Por- tio Dura and the Glosso- Pharyngeal nerves. From these united twigs proceed very small ramifications, which accompany several branches of the external carotid artery, and some of them pass down with the Common Carotid. This superior ganglion also furnishes small twigs which accompany theGlosso-Pharnygeaho the tongue and pharynx. Sometimes a twig from it passes on the back part of the thyroid gland to communicate with the recurrent nerve. From this ganglion go off some small branches, which, uniting with others from the superior laryngeal nerves, form the supe- rior or superficial cardiac nerve, which will be soon described. The trunk of the Sympathetic Nerve descends, on the front of the neck, front this ganglion, as has been already stated. In its course it receives very small twigs from the fourth and fifth cervical nerves, and sends some very small twigs which appear to go to the oesophagus, and some which unite to the la- ryngeal nerve and go to the thyroid gland. Some twigs, which are larger, proceed from it into the thorax, and go to the cardiac plexus hereafter to be described. Opposite to the interval between the fifth and sixth cervical vertebrae it forms another ganglion, of an irregular shape, much smaller than the first. This ganglion, in different subjects, differs in size as well as in several other respects. Sometimes it is en- tirely wanting, and sometimes it is doubled. It is Ganglions and Branches of the Sympathetic. 363 denominated the Middle Cervical , or Thyroid Gan- glion. When the fourth, fifth, and sixth cervical nerves do not send ramifications to the sympathetic nerve, this ganglion receives twigs from them. The Middle Cervical , or Thyroid Ganglion sends many ramifications downwards. Some of them enter the thorax and contribute to the formation of the Cardiac Plexus ,* others accompany the inferior thyroid artery, and, with twigs from the recurrent nerve, form a plexus which extends towards the thyroid gland. Some proceed downwards before, and others behind, the subclavian artery, to the next ganglion ; and among them is generally one which may be regarded as the trunk of the Sympathetic. This third Ganglion is denominated the Inferior. Cervical , or the First Thoracic. It is almost con- stantly found in the same situation, viz. between the transverse process of the last cervical vertebra and the head of the first rib, and is partly covered by the origin of the vertebral artery. It is generally larger* than the middle ganglion. It receives branches from the sixth and seventh cervical, and the two first dor- sal nerves. Ramifications pass from it to the par va- gum and recurrent nerve, and also to the cardiac and pulmonary plexus. From this ganglion the Sympathetic Nerve pro- ceeds downwards on the side of the spine, as will be described hereafter. The Nerves of the Heart , Being derived from branches which have already been mentioned, are now to be described. They arise principally from an arrangement of nerves denominated the Cardiac Plexus , or Plex- uses, which is situated above the curve of the aorta, and extends, on the posterior side of it, from the root of the arteria imiominata to the bifurcation of i >64 Branches of the Sympathetic . the pulmonary artery. This plexus is composed of nerves which are principally formed by the union of small ramifications that are derived from the three above mentioned ganglions of the Sympathetic Nerve , and the nerve itself ; and also from the Par Vagum and some of its branches* These nerves are denominated the Cardiac . — - They descend on their respective sides of the neck, but are somewhat different on the different sides. — On the right side three nerves have been described as particularly entitled to this name, and on the left side but two. The first on the right side is denominated Supe- rior , or Superficial Cardiac NeiPve . It generally arises by several fine threads, which unite into one delicate cord that passes down by the side of the common carotid. When it has arrived on a line with the middle ganglion, it sends a twig to the thy- roid plexus, and another that communicates with a twig from the par vagum, which continues downwards on the carotid artery. After passing beyond the gan- glion, it divides into several branches, which unite themselves to branches of the recurrent nerve that are going to the middle ganglion. The second, which is denominated the Middle Cardiac , the Great Cardiac, or the Deep Cardiac , is the largest of the three. It arises from the Mid- dle Cervical, or Thyroid Ganglion, by five or six fine fibrils, which finally form one, that passes before and across the subclavian; and at that place as well as lower down, it receives twigs from the par vagum : below this, it is joined by a considerable twig from the recurrent, and terminates in the Cardiac Plexus , to which it contributes largely. The third cardiac nerve of the right side is called the Inferior, or the Small Cardiac Nerve. It ori- ginates from the third, or lower cervical ganglion, by 365 Branches of the Sympathetic . many fibrils which unite into a smaller number that form a plexus. It crosses behind the subclavian, and proceeds on the outside of the Arteria Irmomi- nata to the curve of the aorta ; continuing between it and the pulmonary artery, to the anterior corona- ry plexus. In this course it receives several fibres from the recurrent and the par vagum. On the left side the first cardiac nerve arises from the upper ganglion. The second derives its origin from the two lower ganglions. The left superior or superficial cardiac nerve arises like the right, by many distinct fibres, and proceeds downwards in the same way. It descends betweed the carotid and the subclavian, and when it has ar- rived at the place where they originate from the aorta, it divides into a great number of small ramifications. Some pass before the aorta, either to join the branches of the inferior cardiac, or to unite with the cardiac branches of the left nerve of the par vagum. The others proceed behind the aorta, and enter into the common cardiac plexus. The second cardiac nerve of the left side may be called the Great Left Cardiac, and has a double origin as above mentioned. The principal branch in its composition arises from the lowest cervical ganglion, and passes behind the transverse portion of the subclavian artery. , Where the inferior thyroid arises from the subclavian, this branch receives a considerable number of ramifications, which arise from the upper ganglion, and are interwoven with each other before they unite to it. It passes behind the curve of the aorta, and terminates in the great cardiac plexus, which it particularly contributes to form. Here it is joined by many fibres from the par vagum. • 366 Plexus formed by the Nerves of the Heart . The Cardiac Plexus Is situated principally behind the curve of the aorta, at a small distance above the heart. It commences as high as the origin of the Arteria Innominata, and extends downwards to the bifurcation of the pulmo- nary artery. As has been already mentioned, it is principally composed of branches from the middle cardiac nerve of the right side,, and the inferior cardiac nerve of the left ; but it receives branches from the superior cardiac of the left, and sometimes of the right side. Some fibres of the inferior cardiac of the right are also united to it. Many branches proceed from this plexus. A small number pass upon the aorta, and seem to enter into its texture.* Some of them also combine with the ramifications of the Par Vagum in the anterior pulmonary plexus. The majority proceed to the basis of the heart, near the origin of the pulmonary artery and the aorta, and constitute the proper nerves of that organ . They accompany the coronary arteries, and are so arranged around them that, by some anatomists, they have been said to form plexuses, which have been deno- minated Coronary. The Sympathetic Nerve, as has been stated above, proceeds from the ganglion, called the Lower Cer- vical, or the First Thoracic, before the neck of the first rib. It continues to descend, in the same direc- tion, along the spine, exterior to the pleura, to the infe- rior part of the thorax. Near the head of each rib it forms a ganglion, which unites with the intercostal nerve behind it, by two branches, and thus forms an indirect communication with the medula spinalis. * It has been asserted, that some of the anatomists of Paris have traced tbese.nerves on the aorta, to a great distance from the heart. Sympathetic Nerve, fyits Splanchnic Branches. 367 From several of the uppermost of these ganglions, small twigs proceed to the pulmonary plexus, and also to the great trunk of the aorta, below the curve, forming a species of net-work, or plexus, upon it. From the ganglions near the heads of the fifth and sixth ribs, and from four or five of the ganglions tfhich succeed them, small nerves arise, which pro- ceed downwards on the sides of the bodies of the vertebrae, and unite into one trunk that is denomi- nated the Splanchnic Nerve, because it is distributed to the viscera of the abdomen. — This nerve proceeds behind the crus of the diaphragm, on its respective side, into the abdomen. A second and smaller nerve, of the same destination, called the Lesser Splanchnic Nerve, arises lower down, from two or three of the lowermost dorsal ganglions, and penetrates separate- ly into the cavity of the abdomen : it then generally divides into two branches, one of which unites to the great splanchnic nerve, and the other proceeds to the renal plexus soon to be described. As soon as the great splanchnic nerve has entered the abdomen, it divides into many branches, which commonly form small ganglions on each side of the c celiac artery, but above it. • These ganglions are generally contiguous ; but sometimes they are at a small distance from each other, and united by nerves. They are, however, commonly spoken of as one, and called the Semilunar Ganglion. They are of ir- regular forms, and very different from each other in size, as well as form. Those formed by the splanch- nic nerve on one side are sometimes different from those on the other. From this assemblage of ganglions proceed many small nerves, which are woven together so as to form a net-work denominated the Solar Plexus. This plexus is situated anterior to the spine and the crura of the diaphragm ; behind the stomach and 368 Arrangement of the Nerves of the above the pancreas ; and is extended upon the coeliae arid superior mesenteric arteries. Some ramifications from the par vagum and the phrenic also join it. The lower part of the solar plexus, which sur- rounds more immediately the coeliae artery, is termed the Coeliae Plexus. From it net-works of nerves ex- tend upon the great branches of the artery to the organs which they go to. They extend to the stomach, (although it is sup- plied by the par vagum,) along the superior coronary or gastric branch of the coeiiac ; and the fibres in their composition being spread upon the coats of the sto- mach, unite with the branches of the par vagum, which are also spread upon them. A similar net- work, denominated the Hepatic Plexus , extends upon the Hepatic Artery , and from it to the Vena Portarum ,* and accompanies those vessels into the substance of the liver. It also sends branches to the biliary duct and gall bladder ; to the stomach by the arteria gastrica dextra ; and to the omentum. The Sple?iic Artery is invested by a similar but smaller arrangement of nerves, denominated the Splenic Plexus. In its course .to the spleen, this plexus sends some nerves to the pancreas ; and also to the stomach and omentum, with the left gastric artery. The superior mesenteric artery is surrounded by a net-work, which extends to it directly from the solar plexus, and is the largest of all which proceed from that plexus. The Mesenteric Plexus at first nearly surrounds the artery, and proceeds with it between the lamina of the mesentery. In this course it sends branches, with the arteria dolica dextra, to the trans- verse portion of the colon. Between the lamina of the mesentery, it sends ramifications with all the branches of the artery, to the small intestines gene- Abdominal Viscera „ 369 rally; to the ececum, and the right portion of the colon ; as well as to the mesenteric glands. From the lower part of the solar plexus a net-work proceeds, on the front of the aorta, to the inferior mesenteric artery, and surrounds it. Nerves from this plexus accompany the artery to the left portion of the colon and the rectum. Some of their ramifi- cations combine with those of the hypogastric plexus. The Emulgent Artery is attended by nerves, which are arranged like a net-work on its anterior and poste- rior surfaces, and are denominated the Renal Plexus . They are derived from the solar plexus, and fre- quently contain small ganglions. They proceed with the artery to the fissure of the kidney, and are distributed with its different ramifications, in the substance of the organ. Some branches pass from them to the renal gland with the capsular artery. Before the renal plexus arrives at the kidney, it sends off, from its inferior part, some new fibres, which, after joining some others from one of the lumbar nerves, accompany the spermatic arteries, and are, therefore, called the Spermatic Plexus. In the male these fibres proceed through the abdominal ring, and many of them go to the testis, but they are followed with great difficulty, on account of their small size. In the female, they go to the ovary and the fallo- pian tube. From the great plexuses above, a small net-w T ork continues downwards on the aorta, receiving fibres from the intercostals on each side ; at the great bifur- cation of the aorta it divides, and is joined on each side by many ramifications from the third lumbar nerves, which thus form a plexus of considerable extent, that sends nerves to the bladder, rectum, and vesiculse seminales in males; and to the uterus and Yol, ii. 47 370 Termination of the Sympathetic Nerve, vagina, as well as the bladder and rectum, in fe- males.* This is called the Hypogastric Plexus . The plexuses above mentioned are derived from the splanchnic nerve, which came off from the Sym- pathetic in the thorax. The Sympathetic Nerve, after giving off the lesser splanchnic^ is diminished in size, and approaches nearer to the bodies of the vertebra. It passes through the crura of the diaphragm, and then pro- ceeds forwards and downwards upon the spine, be- tween the tendinous crura of the diaphragm and psoas muscle ; near the vena cava on the right side, and the aorta on the left. In this course, it gene- rally receives one or two small cords from the ante- rior branch of each of the lumbar nerves: these cords proceed downwards and forwards, between the bo- dies of the vertebrae and the psoas muscle, and a ganglion is generally formed at the plaee where they join the nerve. In its descent on the lumbar vertebrae, the Sym- pathetic sends off several nerves that unite to the net-work nvhich descends on the aorta from the plexus above. After passing over the lumbar vertebrae, it descends into the pelvis, close to the sacrum, on the inner side of the great foramina: here it also forms ganglions, and communicates with the sacral nerves, and likewise with the hypogastric plexus. It termi- nates on the os coccygis, where its minute fibres join those of the opposite side. * Although the testicle receives nerves which are derived from the Sym* pathetic, the penis and other external parts of the organs of generation do not: the nerves which accompany the pudic artery being derived from those which unite to form the great Sciatic, SYSTEM of anatomy. PART XI. OF THE ABSORBENT VESSELS.* The absorbent vessels are small transparent tubes, of a delicate structure, which exist in considerable numbers in almost every part of the body. These tubes originate upon the surfaces of all the cavities of the body; and of the cellular membrane, in all the various parts into which it penetrates ; upon the internal surface of the stomach and the intes- tines ; and probably upon the skin. Those which originate in the Lower Extremities and the Cavity of the Abdomen, unite and form a large trunk called the Thoracic Duct , which proceeds through the thorax, and terminates in the left Sub- clavian Vein, at its junction with the Internal Jugu- lar. Those of the Left Upper Extremity, the Left Side of the Head, and the contiguous parts, form a trunk which terminates in the same place. While the remaining absorbents, or those of the Right Up- per Extremity, and the Right side of the Head , 8,'C. also form a trunk, which terminates in the cor- responding part of the Right Subclavian Vein. The absorbent vessels of the middle size, which * Discovered at Leyden in 1650.. by Olaus Rndbeck. and at Copenhagen in 1651. by Bartholine. — E d ' 372 Structure of the JibsQrbent Vessels. arise from the union of the small vessels, and unite to form the larger, in their progress to these large vessels, pass through certain bodies which have been denominated Conglobate Glands, and may be consi- dered as appendages of the absorbent system. The absorbent vessels are composed of two coats, v hich are thin, but dense and firm, and also elastic. The coats of the thoracic duct may be separated from each other. The internal surface of the exte- rior coat is fibrous. The internal coat is a delicate but strong membrane. — There is great reason to be- lieve that the above mentioned fibres are muscular, or at least irritable : for the absorbent vessels have been observed, by Haller, to contract upon the ap- plication of strong sulphuric acid. They have also been observed to propel their contents with consider- able rapidity, by their own contraction, independent of pressure, or of motion communicated by any other body. Blood vessels are sometimes observable in the coats of the larger absorbents, in injected subjects. The vascularity of these tubes may also be inferred from the inflammation which frequently takes place in them. Nerves have not been traced into their texture ; but the absorbents seem to be painful when they are inflamed, and, therefore, it is probable that they are supplied with nerves. The absorbent vessels are very generally supplied with valves, which are much more numerous in some of them than in others ; and are different in their number, in the same vessels, in different subjects. Very frequently there are several valves in the course of an inch : sometimes a valve will not appear in the course of several inches. In the Thoracic Duct , the number of valves is very different in different subjects. These valves are folds or plaits of the in- Commencement of the Absorbents. 373 ternal membrane, and are of a semi-circular form. There are commonly two of them together origina- ting from opposite sides of the vessel. The absorbents are generally somewhat dilated on the side of the valve which is next to their termi- nation, and this occasions their knotted appearance when they are injected. The object of this valvular structure seems to be the prevention of retrograde motion of the contained fluid, in consequence of la- teral pressure. • • Where the different trunks of the absorbents open into the veins, there are one or two valves to prevent the regurgitation of the blood into them. The valves of course prevent the injection of the branches of these vessels from their trunks. —In some animals the valves have sometimes been ruptured, or forced back; and the absorbents have been injected in a retrograde direction. There are but two or three instances upon record where this has been practicable in the Human Subject. In consequence of the impracticability of injecting the small branches from the larger, the absorbent vessels cannot, generally, be demonstrated at their commencement, or origin. It is, however, to be ob- served, that the Lacteals , or Absorbents of the Intes- tines, appear no way different from other absorbents; and they have been seen distended with chyle, from their commencement, in certain subjects w’ho had died suddenly. Their origins have been described very differently by different observers. Mr. Cruikshank describes them as originating on the surfaces of the villi, by a number of very small radiated branches with open orifices; which branches soon unite to form a trunk. Lieberkuhn believed them to commence in the form of an ampullula . — See page 105 of this volume. The second Monro also believes that the absorb- 574 Conglobate Glands . ents begin by very small tubes, with open orifices, in several species of fish.* It is stated by Dr. Soemmering upon the authority of Haase, a German anatomist, that when mercury is forced backwards in the absorbent vessels of tbe foot and the heart, it has sometimes escaped on the sur- faces of those parts. The probable inference from these facts is, that those vessels originate by open orifices on the surfaces of the heart and foot. The bodies connected with the absorbent vessels, which are called Conglobate Glands, are generally of a roundish, or irregular oval form, and somewhat flattened. They are of various sizes, from two lines in diameter to more than twelve. Their colour is frequently whitish, but sometimes it is slightly in- clined to red. They are invested with a covering of cellular membrane, which appears like a membranous coat ; and they are connected to the contiguous parts by a loose cellular substance. When the absorbent vessels connected with these bodies approach near to them, they divide into a number of ramifications, most of which enter into the substance of the gland, while some of them run over it. On the opposite side of the gland a number of branches go out, which unite and form trunks similar to those which entered the gland. The vessels which enter the gland are called Vasa infer entia , and those which go out of it Vasa effer entia. These vessels are generally much convoluted in the substance of the glands, so that those bodies sometimes appear like a mere convolution of absorb- ent vessels. There has been much diversity of sen- timent respecting the structure of these organs.! * See his work on the Structure and Physiology of Fishes, p. 34. f Mr. Abernethy states, that the mesenteric gland of the Whale con- sists of large spherical bags into which a number of the lacteals open. — Numerous blood vessels are ramified on the surfaces of these cysts ; / Fluid contained in the Absorbents, 375 The absorbent vessels, in the different parts of the body, generally contain fluids resembling those which are found in those parts. Mr. Hewson opened the large absorbents in many living animals of different kinds, and found that they contained a transparent fluid, which coagulated when exposed to the air. The arrangement of these vessels resembles that of the veins in several respects. Many of them are superficial ; but there are also deep-seated absorbents which accompany the blood vessels. and injection passes from them into the cyst. He also found cells in the glands of the absorbent vessels, in the groin and the axilla of the horse— See Philosophical Transactions, for 1796, Part I. CHAPTER I. OF THE ABSORBENTS OF THE LOWER EXTREMITIES, THE ABDOMEN, AND THE THORAX, Under this head are arranged the ramifications of all the vessels which unite to form the Thoracic Duct . SECTION I. Of the Absorbents of the Lower Extremities . These absorbents, like the veins, are superficial and deep-seated. The Superficial lie in the cellu- lar membrane, very near the skin ; and form an irre- gular net-work which extends over the whole limb. They are, however, most numerous on the internal side. The Deep-seated accompany the arteries like the veins, and there are two at least to each artery. The Superficial Absorbents Have been injected from the toes so as to form a net-work, which occupies the upper surface of the foot. They have also been injected in a similar manner on the sole. Those on the upper surface of the foot generally proceed upward on the anterior and inner side of the leg; but some of them pass on the external side of it. Those on the sole are con- tinued on the back of the leg; but communicate very frequently with the anterior vessels. Some of the absorbents from the outside of the foot and leg enter into some of the popliteal glands, soon to be de- scribed; but they are not numerous; and the princi- pal number continues up to the glands of the groin. Absorbents of the Lower Extremity. 377 The absorbents which originate on the surface of the thigh, as we!! as those which pass over it from below, incline gradually a!ong the auterior and pos- terior surface, to the internal side of it; on which they proceed, in great numbers, and very near to each other, to the inguinal glands. Superficial absorbents proceed also from the buttock, and lower part of the back, from the lower part of the abdomen, the peri- neum and the exterior of the genital organs, to these glands. The Deep-seated Absorbents Are named from the arteries they accompany. The Anterior Tibial Absorbents. The anterior tibial artery is generally attended by one which comes with it from the sole, and by another which commences on the upper surface of the foot. The first mentioned absorbent continues with the ar- tery. The last often passes through an aperture in the interosseal ligament, about one-third of the dis- tance from the ankle to the knee, and accompanies the fibular artery, while the anterior tibial artery is join- ed by other absorbents about the same place. In some instances a small absorbent gland occurs in this course, at a short distance below the knee. The Posterior Tibial Absorbents Have been injected from the under side of the toes. They accompany the ramifications on the sole of the foot ; and after unitiug, continue with the main trunk up the leg, where they enter the popliteal glands. The Peroneal Absorbents arise also from the sole of the foot, and its external side. They accompany the peroneal artery, and terminate in the popliteal glands, which receive also the absorbents from the knee and ham. Yol. ii. •I 1 : -I 378 * absorbents of the Lower Extremity . From these glands four or five absorbent vessels proceed, which accompany the great blood vessels of the lower extremity; and, proceeding with them through the aperture in the tendon of the adductors, continue upwards until they enter some of the glands of the groin. The glands of the ham and groin, which are so in- timately connected with the absorbents of the lower extremity, are very different from each other. The Popliteal Glands , or those of the Ham , are but three or four in number, and very small in size. They are generally deep-seated, and very near the artery. The Inguinal Glands vary in number, from eight to twelve or more. They are superficial and deep- seated. The superficial communicate principally with the superficial absorbents. The lowermost of them are at some distance below Poupart’s ligament, and the uppermost are rather above it. They are exterior to the fascia of the thigh. Their number is generally six or eight, while that of the deep-seated is but three or four. The superficial absorbents from below, approach very near to each other, and enter these glands. — They are commonly distributed among three or four of the lowermost ; but some of them pass by these, and proceed to one that is higher up; and some- times there are absorbent vessels which pass to the abdomen without entering into any of the glands of the groin. The deep-seated absorbents pass into the deep- seated glands, which, as has been already observed, are but few, and lie very near the artery under the fascia of the thigh. The two sets of glands are connected to each other by many absorbent vessels that pass between them. The vessels which finally go out of these glands are considerably less in num- Inguinal and External Iliac Glands. 379 ber than those which enter into them. They pro- eeed under Pou part’s ligament, and, in some in- stances, a large proportion of them passes through three glands which lie below this ligament* and are often so arranged, that they lie on each side of the great femoral vessels, and above them. One very frequently is found on the inside of the femoral vein, in the vacuity between it and the internal part of the ligament. All the absorbents of the lower extre- mity, however, do not enter these glands. Some pass along the great vessels and enter other glands near the margin of the pelvis. Some also descend a short distance into the pelvis, and unite with vessels that are passing from the pelvis to the plexus and the glands that surround the external iliac. The absorbents which proceed from the glands last mentioned, joined to those which pass under Poupart’s ligament, without entering these glands, and some which come from the pelvis, form a large plexus, which almost surrounds the external iliac ves- sels, aud contains many glands. These External Iliac Glands vary in their number from six to ten or twelve. They lie on the side of the pelvis, in the course of the external iliac vessels, and some of them are of considerable size. These glands and the plexus of absorbents, extend in the track of the iliac vessels, to the first lumbar vertebra. In this course they are joined by the plexus which comes from the pelvis ; and soon after they arrive at the Lumbar Glands , which form a very large assem- blage, that extends from the bifurcation of the aorta to the crura of the diaphragm. These glands lie irregularly, on the aorta, the vena cava, and the lumbar vertebrae. Most if not all the absorbents above mentioned pass through some of them ; and from the union of these absorbents, some 380 Absorbents of the Testicles, fyc. of the great brandies, which unite to form the tho- racic duct, are derived. In this course from the thigh to the lumbar glands, these absorbent vessels are joined by several others. The Superficial Absorbents of the scrotum commonly enter into the upper inguinal glands, and thus unite to the great body of absorbents. The Absorbents of the Testicles, originate in the body, and the coats of the testicle, and in the epi- didymis, and are remarkably large and numerous. — They proceed along the spermatic cord, through the abdominal ring, to the lumbar glands. These ves- sels are remarkable for the little communication they have with each other. The Deep-seated Absorbents of the Scrotum ac- company the absorbents of the testicle to the lumbar glands; but those which are superficial enter the up- per inguinal glands. The Absorbents of the Penis are also deep-seated and superficial. The deep-seated arise from the body of the penis, and accompany the internal pudic artery into the pelvis. The superficial absorbents arise from the prepuce, and pass along the dorsum of the penis. There are frequently several trunks which receive branches from the lower surface of the penis in their course. At the root of the penis they generally separate to the right and left, and pass to the glands on the respective sides. In females, the absorbents of the interior of the clitoris accompany the internal pudic artery. Some, which arise about the vagina, pass through the abdo- minal ring with the round ! 1 and ’ rs pro- ceed to the inguinal- glands. Absorbents of the Pelvis and the Kidneys. 381 SECTION ir. Of the Absorbents of the Abdomen and Thorax. The Absorbents of the lower portions of the pa- rietes of the Abdomen and the Pelvis unite into trunks that follow the epigastric, the circumflex and the iliac, as well as the lumbar and sacral arteries, &c. They proceed to some of the glands which are in the groin ; or in the external iliac, the hypogas- tric, or some of the contiguous plexuses. The Absorbents of the Womb are extremely nu- merous; and, in the gravid state, are very large. Those which are on the neck and anterior part of the uterus, join the hypogastric plexus. Those which are on the posterior part of the body, accompany the spermatic vessels. The Absorbents of the Bladder pass to small glands on its lateral and inferior parts, and finally join the hypogastric plexus. The Absorbents of the Rectum are of considerable size. They pass though glands that lie upon that intestine, and unite with the lumbar plexus. The Absorbents of the Kidney are superficial and deep-seated. '1 hey are very numerous, but, in a healthy state of the parts, are discovered with dilli- culty. Cruikshank describes them as they appeared, filled with blood, iri consequence of pressing upon the kidney when its veins were full of blood. Mas- cagni did not inject the superficial vessels with mer- cury ; but describes them as they appeared when filled with colourless size, after he had injected the blood vessels of the organ with the coloured fluid. — The deep-seated absorbents pass out of the fissure of the* kidney with the blood vessels, and unite with the superficial ; they proceed to the lumbar plexus, and pass into different glands. 382 Lacteals, or absorbents of the Intestines. Absorbent vessels can be proved to proceed from the pelvis of the kidney, and the ureters, by artifices analogous to those above mentioned. The Gian dulse Renales are also supplied with ab- sorbents, which are numerous in proportion to the size of the organs. They commonly join those of the kidney. The Absorbents of the Intestines Have generally been called LACTEALS, from the white colour of the chyle which they contain : but there seems no reason for believing that they are dif- ferent in their structure and nature from the absorb- ents in other parts of the body. A small number of them appear as if they formed a part of the struc- ture of the intestines, and originated from their ex- ternal surface, as they do in other parts of the abdo- men ; while the principal part of them are appro- priated to the absorption of the contents of the cavity of the intestines. The first mentioned absorbents run between the muscular and peritoneal coats, and proceed for some distance lengthways on the intestine, while the others proceed for some distance within the muscular coat, with the arteries ; and after passing through it, con- tinue between the lamina of the mesentery. Branches of these different absorbents are frequent- ly united in one trunk so as to prove that there is no essential difference between them. The absorbents which come from the internal sur- face of the intestines commence in the villi. The manner in which they originate has been the subject of considerable inquiry, as has been stated in the ac- count of the intestines.* The lacteals or absorbents of the intestines are rr See page 105. Lac teals, or Absorbents of the Intestines. 383 very numerous. They pass between the lamina of the mesentery to glands which are also seated be- tween those lamina. The number of these glands is very considerable,* and they are various in size — - some being very minute, and others eight or ten lines in diameter. They are generally placed at a small distance from each other, and are most numerous in that part of the mesentery which is nearest to the spine. They are almost always at some distance from the intestines. They appear to be precisely like the absorbent glands in other places. These absorbent vessels, in their course frequently divide into branches; which sometimes go to the same gland, sometimes to different glands, and sometimes unite with other absorbent vessels. As they proceed, they frequently enlarge in size. When they have ar- rived near the spine, they frequently form three or four trunks, and sometimes one or two ; which pro- ceed in the course of the superior mesenteric artery, until they have arrived near to the aorta. Here they either pass into the thoracic duct, or descend and join the trunks from the inferior extremities, to form the thoracic duct. The absorbents of the great intestines are not equal in size to those of the small; but they are numerous. They enter into glands, which are very near, and in some places, in contact with the intestine; and are commonly very small in size. The vessels which arise from the caecum, and the right portion, as well as the arch of the colon, unite with those of the small intestines ; while the vessels from the left side of the colon, and the rectum, pro- ceed to the lumbar glands. The absorbents of the intestines are frequently injected with mercury ; but the injection does not proceed to their termination with so much facility as They have been estimated between 130 and 150. 384 Absorbents of the Stomach. it does in others vessels cf the same kind. They have, however, very oftt n been seen in animals, who were killed for the purpose after eating miik ; and in several human subjects who died suddenly during digestion. — The description of the origin of the lactoals, quoted in page 108, from Mr. Cruikshank, was taken from a subject of this kind, of which an account is given in his work on the absorbing ves- sels, p. 59. It is worthy of note, that in several instances, in which the lac teals were thus found distended with chyle, the glands in the mesentery were also uniform- ly white. The Absorbents of the. Stomach Are of considerable size, and form three divisions. The vessels of the first set appear upon both sides of the stomach, and pass through a few glands on the small curvature near the omentum minus. — From these glands they proceed to others, which are larger, and which also receive some of the deep- seated absorbents of the liver. The vessels from these glands pass to the thoracic duct, near the ori- gin of the eoeliac artery. The second arise also on both sides of the stomach, and pass to the left extre- mity of the great curvature to unite with the absor- bents of that side of the great omentum. They then proceed with the lymphatics of the spleen and pan- creas, to the thoracic duct. The last set, pass off from the right extremity of the great curvature, and unite also with absorbents from the right portion of the omentum. They proceed near the pylorus, and go to the thoracic duct, with some of the deep- seated absorbents of the liver. Although the absorbents of the stomach are deep- seated, as well as superficial, it is a general senti- ment, that they do not contain chyle in the' human Absorbents of the Liver . ,jsj subject ; notwithstanding chyle has been found in the absorbents on the stomach of dogs, and some other animals. It ought, however to be remembered, that Sabatier has, in some instances, seen white lines on the stomach, which he supposed to be lacteals. The Absorbents of the Liver Are especially interesting, because they have been more completely injected than those of any other viscus. They are deep-seated and superficial . The superficial it has been already observed admit of in- jection in a retrograde direction, and, therefore, can be exhibited most minutely ramified. They commu- nicate freely with each other, and also with the deep- seated vessels, by their small ramifications ; so that the whole gland has been injected from one large vessel. The gland is so large, that the absorbents of the superior and inferior surfaces proceed from it in dif- ferent directions. A large absorbent is generally found on the sus- pensory ligament. This is formed by the union of a great many branches that arise both on the right and left lobes, but principally on the right. It often passes through the diaphragm at an interstice which is anterior to the xiphoid cartilage, and then proceeds through glands on the anterior part of the pericar- dium. Several absorbents proceed to the lateral ligaments on each side, and then pass through the diaphragm. Some of these branches return again into the abdo- men, and the others generally run forwards in the course of the ribs, and join those which passed up from the suspensory ligament. The trunk, or trunks, formed by these vessels, either pass up between the lamina of the mediastinum, and terminate in the up- Vol. il 49 386 Absorbents of the Liver and Spleen . per part of the thoracic duct ; or they accompany the internal mammary arteries, and terminate on the left side in the thoracic duct, and on the right in the trunk of the absorbents of that side. The Absorbents on the concave side of the Liver are as numerous as those on the convex side ; they are also very abundant on the surface of the gall bladder. The greatest part of them join the deep- seated vessels. The Deep-seated Absorbents proceed in consider- able numbers front the interior of the liver through the portae. They accompany the billiary ducts and the great blood vessels of the organ ; and, after passing through several glands, near the vena porta- rum, terminate in the thoracic duct, near the com- mencement of the superior mesenteric artery. Mascagni states, that the absorbents of the liver will be distended, by injecting warm water into the biliary ducts, or the vena portarum. He also observes, that in those preparations in which the superficial vessels are completely injected, in the retrograde direction, the peritoneal coat of the liver appears to be composed entirely of absorbent vessels ; and to be connected to the membrane within, by many filaments which are also absorbent vessels. The Absorbents of the Spleen Are composed of superficial and deep-seated vessels; but they differ greatly from those of the liver, in this respect, that the superficial vessels are remark- ably small in the human subject. Mascagni however asserts, that when the blood vessels of the spleen are injected with size, colour- ed with vermilion, these absorbents will be filled with colourless size. In the spleen of the calf the superficial absorbents are remarkably large. 887 Absorbents- of the Pancreas. In the human subject the superficial absorbents of the spleen proceed from the convex to the concave surface, and there communicate with the deep-seated absorbents., which proceed from the interior of the organ with the blood vessels. These Beep-seated Absorbents are very numer- ous, and also large. They accompany the splenic artery; and in their course pass through many glands, some of which are said to be of a dark colour. The glands lie on the splenic artery, at a short distance from each other. The absorbents of the spleen re- ceive the absorbents of the pancreas in their course; they unite with the absorbents of the stomach and the lower surface of the liver, and pass with them to the thoracic duct. Little has been latterly said by practical anato- mists respecting The Absorbents of the Pancreas, Mr. Cruikshank once injected them in the retrograde direction ; he found that they came out of the lobes of the pancreas in short branches like the blood ves- sels, and passed at right angles into the absorbents of the spleen, as they accompanied the artery in the groove of the pancreas. THE THORACIC DUCTf Or common trunk of the absorbent system, is formed by the union of those absorbent vessels which are collected on the lumbar vertebrae. These vessels, as it has been already observed, are derived from various sources, viz. The Lower Extremities ; the lower part of the Trunk of the Body; the Organs of Generation; the * First discovered by Eustachius in the horse, 15G4, but he considers it a vein for the nourishment of the thoracic viscera. — Ed. 3SS Commencement of the Thoracic Duct. Intestines, with the other Viscera of the abdomen and Pelvis, except a part of the liver. Their num- ber is proportioned to the extent of their origin : for, with the numerous glands appropriated to them, they form the largest absorbent plexus in the body, and are spread over a considerable portion of the aorta and the vena cava. The manner in which these vessels unite to form the thoracic duct, is very different in different sub- jects; but in a majority of cases it originates imme- diately from three vessels, two of which are the trunks of the absorbents of the lower extremities, and the other is the common trunk of the lacteals and the other absorbents of the intestines. These vessels generally unite on the second or third lumbar vertebrae ; and, in some instances, the trunk which they form dilates considerably, soon af- ter its commencement ; in consequence of which it was formerly called the RECEPTACLE of the CHYLE. At first it lies behind the aorta, but it soon inclines to the right of it, so as to be behind the right crus of the diaphragm. In the thorax, it ap- pears on the front of the spine, between the aorta and the vena azygos, and continues between these vessels until it has arrived at the fourth or third dor- sal vertebra. It then inclines to the left, and pro- ceeds in that direction until it emerges from the thorax, and has arisen above the left pleura, when it continues to ascend behind the internal jugular, nearly as high as the sixth cervical vertebrae : it then turns downward and forward, and, after descend- ing from six to ten lines, terminates in the back part of the angle formed by the union of the left internal jugular with the left subclavian vein. Sometimes, after rising out of the thorax, it divides into two branches, which unite before they terminate. Some- times it divides, and one of the branches terminates Absorbents of the Lungs. 389 at the above mentioned angle, and the other in the subclavian vein, to the left of it. The orifice of the thoracic duct has two valves, which effectually prevent the passage of blood into it from the vena cava. There are sometimes slight flexures in the course of the duct; but it generally inclines to the left, in the upper part of the thorax, as above mentioned: and is then so near the left larnen of the mediastinum that if it be filled with coloured injection, it can be seen through that membrane, when the left lung is raised up and pressed to the right. The duct sometimes varies considerably in its dia- meter in different parts of its course. About the mid- dle of the thorax it has often been found very small. In these cases it generally enlarges in its progress upwards, and is often three lines in diameter, in its upper part. Many anatomists have observed it to divide and to unite again, about the middle of the thorax. Absorbents of the Lungs. The absorbents of the lungs are very numerous, and, like those of other viscera, are superficial and deep-seated. The large superficial vessels run in the interstices between the lobuli, and therefore form angular figures of considerable size. In successful injections, the vacancies within these figures are filled up with small vessels, and the whole surface appears minutely in- jected. Mascagni observes, that the superficial vessels are very visible when any fluid has been effused into the cavity of the thorax ; or when warm water is in- jected, either into the blood vessels of the lungs, or the ramifications of the trachea. Cruikshank de- monstrated them by inflating the lungs of a still born 390 Absorbents of the Lungs. child ; in which case the air passes rapidly into them. The deep-seated absorbents accompany the blood vessels and the ramifications of the bronchi®. They pass to the dark coloured glands, which are situa- ted on the trachea at its bifurcation; and on those portions of the bronchi® which are exterior to the lungs. The injection of the absorbents, which pass to and from these glands, seems to prove that they are of the same nature with the absorbent glands in general notwithstanding their colour. They are nu- merous, and they vary in size; from a diameter of two lines, to that of eight or ten. From these glands, some of the absorbents of the left lung pass into the thoracic duct, while it is in the thorax, behind the bifurcation of the trachea; others proceed upwards and enter into it near its ter- mination; while those of the right lung terminate in the common trunk of the absorbents of the right side 0 CHAPTER II. OF THE ABSORBENTS OF THE HEAD AND NECK ; OF THE UPPER EXTREMITIES, AND THE UPPER PART OF THE TRUNK OF THE BODY. The absorbents from the various parts of the head pass through glands, which are situated on the neck, or the lower part of the head. Those on the head are the least numerous, and also the least in size. — Some of them, which are generally small, lie about the parotid gland. Several of them, which are also small, are on the occiput, below and behind the mastoid process. Sometimes there are two or three on the cheek, near the basis of the lower jaw, about the anterior edge of the masseter muscle. Below the lower jaw, in contact with the sub-maxillary gland and anterior to it, there are always a number of these glands, which are generally small, but often swelled during infancy. The Glands on the JYech are the most numerous. Many of them are within the sterno- mastoid muscle, and accompany the internal jugular vein and the carotid artery down to the first rib. Many also lie in the triangular space between the sterno- mastoid muscle, the trapezius, and the clavicle ; therefore it has been truly said thpt the glands of the neck are more numerous than those of any other part, except the mesentery. They are frequently called Glan- dulse Concatenatae. It has already been mentioned that the various absorbents, which are connected with these glands, unite on each side into a trunk, which on the left passes into the thoracic duct, and on the right into the common trunk of the absorbents of that side. 392 Absorbents of the Head and JYeck . SECTION i. Of the Absorbents of the Head and Neck. There is the greatest reason to believe that the brain and its appendages are supplied with absorb- ents like the other parts. Some of these vessels have been discovered in the cavity of the cranium ; but very little precise information has as yet been obtain- ed, respecting the extent, or arrangement of the ab- sorbent system, in this part of the body. The absorbents on the exterior of the head are as numerous as in other parts of the body. On the oc- ciput they pass down, inclining towards the ear, and continue behind it to the side of the neck ; behind the ear they pass through several glands. — From the middle or temporal region of the cranium, they pass with the carotid artery before the ear, and enter some small glands that lie on the parotid ; from which they continue to the neck. They are on every part of the face, and unite, so that their principal trunks, which are very numerous, pass over the basis of the lower jaw, near the facial artery. They enter into glands, which are also very numerous, immediately under the jaw, or which are sometimes to be found on the cheek, at the anterior edge of the masseter muscle. All the absorbents of the exterior part of the head pass to the glands on the side of the neck, already described. Those from the interior of the nose accompany the ramifications of the internal maxillary artery, and proceed to glands behind the angle of the lower jaw; into which glands also enter the absorbents of the tongue and inner parts of the mouth. The absorbents of the thyroid gland, on the left side, pass down to the thoracic duct; those on the right, unite to the trunk of the absorbents on that 393 Absorbents of the Hand and Ann , side, near its termination. It has been remarked that they can be readily injected, by thrusting the pipe into the substance of the gland. SECTION II. Of the Absorbents of the Arm and Upper Part of the Trunk . The absorbents of the arm are superficial and deep- seated, like those of the lower extremity. The superficial absorbents have been injected on the anterior and posterior surfaces of the fingers and the thumb, near their sides. On the back of the hand they are very numerous, and increase consi- derably in their progress up the fore-arm. As they proceed upwards, they incline towards the anterior surface of the fore-arm ; so that by the time they have arrived at the elbow, almost all of them are on the anterior surface. The absorbents on the anterior part of the hand are not so numerous as those on the back. Sometimes there are digital branches from the fin- gers, and an arcus in the palm ; but this bow is not formed by one large absorbent, analogous to the ulnar artery. On the contrary, its two extremities are con- tinued over the wrist, and pass on the fore-arm like the absorbents. At the elbow, some of them often pass into one or two small glands, which are very superficial; but the whole of the absorbents, somewhat reduced in number, as some of them unite together, pass along with the bloodvessels into the hollow of the arm-pit; where they enter the axillary glands. There are generally one or more vessels which pass in the course of the cephalic vein, between the pectoral and the deltoid muscle, and enter into some of the glands under the clavicle. VOL, II, 50 394 Absorbents of the Upper Part of the Trunk. There are almost always several glands in and near the axilla. Some of them are very near the great blood vessels ; sometimes one or more of them are much lower; sometimes they are to be found under the pectoral muscle. They are commonly not so large as those of the groin, and are surrounded with fat. The deep-seated absorbents originate also at the fingers, and soon accompany the branches of the ar- teries. Those which attend the radial artery, origi- nate on the back of the hand, and also in the palm, where they are associated with the arcus profundus. They go up with the radial artery to the elbow, and sometimes pass through a small gland about the mid- dle of the fore-arm. Those which attend the ulnar artery, commence under the aponeurosis palmaris, and go with the ar- tery to the elbow; at the bend of the elbow they are generally joined by one or more, which accompany the interosseal artery ; there they unite, so as to form several trunks which pass up to the axillary with the humeral artery. They sometimes pass through one or two glands, which are near the elbow ; and they receive in their course, deep-seated branches from the muscles on the humerus. The absorbents from the anterior and external part of the thorax, and the upper part of the abdo- men, also proceed to the axilla, and enter into the glands there ; those which are deep seated, joining the deep-seated vessels. The absorbents of the mammae pass to the same glands; and when they are affected with the virus of cancer, can often be perceived, in their course, in the living subject. The absorbents of the uppermost half of the back, and those of the back of the neck, go likewise to the axilla. The absorbent vessels, collected from these van- Facts relating to Cutaneous Absorption. 395 ous sources, proceed from the exterior to the inner- most glands, but with a considerable diminution of their number; they accompany the subclavian vein, and are reduced to one or two trunks, that gene- rally unite before their termination. On the left side , the absorbents of the head and neck generally open into the thoracic duct, as has been already ob- served; and those of the left arm also open into the thoracic duct, or into the subclavian vein very near it. On the right side the absorbents from each of these parts empty into the common trunk; which often is formed by the union of large vessels, from four sources; viz. the Head, the Thyroid gland, the right arm, and the right cavity of the Thorax, &c. The diameter of the trunk is very considerable ; but it is often not more than half an inch in length. It generally opens into the right subclavian vein, at the place where it unites to the right internal ju- gular. Two respectable physiologists of Europe, (M. Seguin, of Paris, and the late Dr. Currie, of Liverpool,) have doubted whether absorption takes place on the exter- nal surface of the skin.* This question has been ex- amined in a very interesting manner by several gra- duates of the University of Pennsylvania, who chose it for the subject of their inaugural theses ; viz. Drs. Rousseau, Klapp, Daingerfield, Mussey, and J. Brad- ner Stewart. The three first of these gentlemen state, that when spirit of turpentine, and several other substances which are commonly supposed to be absorbed by the skin, were applied to it in a way which prevented their volatile * I believe that M. Seguin’s Memoir on this subject was read to the Academy of Sciences a short time before the meetings of that body were suspended. It was published by M. Fourcroy, in La Medicine Eclairee par les Sciences Physiques, vol. iii. An extract from M. Fourcroy’s pub- lication may be se*n in the 19th chapter of the first volume of Dr. Currie’s $ likeness. Cuboides. A bone of the foot ; from a cube, and enJ'uj likeness ; because it resembles a cube. Cuneiform. Some bones are so called ; from cuneus a wedge, and forma likeness ; being shaped like a wedge. D. Deltoid. A muscle resembling the Greek letter A : from A, and eih 5 resemblance. Diaphragm. The muscle which separates the thorax from the abdomen ; from to divide. Diarthrosis. A moveable connexion of bones; from f$icto6goa to articulate. Digastric. From <5Vs twice, and ng a bell y ; having two bellies. Diploe. The spongy substance between the two tables of the skull ; from fttrXttt to double. Duodenum. The first portion of the small intestines ; so call- ed because the ancients supposed that it did not exceed the breadth of twelve fingers ; from. duodenUs, consisting of twelve. Dura Mater. The outermost membrane of the brain ; called dura , because it is much harder than the other membranes, and mater , from the idea of the ancient^ that it was the source of all the other membranes. E. Embryo. The child in the womb is so called before the fifth month, after which it is termed foetus ; from e^v&> to bud forth. Enarthrosis. An articulation of bones ; from -a in, and a joint or articulation. Enteric. Belonging to the intestines ; from evregov an entrail or intestine. Epidermis. The scarf or outermost skin ; from tm upon, and thg/tii the skin. . .. Epididymis. The small oblong body which lies above the tes- ticles ; from upon, and JiS'v /. a testicle. Epigastric. The superior part of the abdomen ; from svt upon, and the stomach. Epiglottis , A cartilage of the larynx so called; from em upon, and yxarhi the aperture of the larynx, being situated upon the glottis. VOL, II. 53 418 Glossary. Epiphysis , A portion of bone growing upon another bone, but separated from it by a cartilage : from em upon, and pva to grow. Epiploor i. The membranous viscusof the abdomen, which co- vers the intestines, and hangs to the bottom of the stomach; from eKin-Xsa to swim upon. Ethmoid. From a sieve, and e/^05 resemblance; being perforated like a sieve. F. Fascia. An expansion, enclosing other parts, like a band ; from fascis a bundle. Falciform. Shaped like a si the ; from falx, a si the. Fasciculus. A little bundle, dim. of fascis a bundle. Fauces. The plural of faux , the top of the throat. G. Ganglion. Tv/y^iot, a knot in the course of a nerve. Gastrocnemius. The muscle which forms the thick of the leg; from a belly, and the leg. Genio. Names compounded with this word belong to mus- cles which are attached to the chin, as geno-glossus — geno- hyoideus, &c. ; from yeve;ov the chin. Ginglymus. An articulation ; from yifyXv^ot; a hinge. Glenoid cavity. From y*m a cavity, and a Sat resemblance. Glosso. Names compounded with this word belong to muscles which are attached to the tongue ; as glosso-pharyngeus — glosso-staphilinus, &c. ; from yXawu, the tongue. Glottis. The superior opening of the larynx at the bottom of the tongue ; from yXcaria the tongue. Glutseus. The name of a muscle ; from yA&rog the buttocks. Gomphosis. To^paicng, a species of immoveable connexion of bones ; from yo/tnpet a nail, because one bone is fixed in an- other bone like a nail in a board. II. , Helix. The outward circle of the ear; from uXta to turn about. Hepar. The liver. an abdominal viscus. Hyaloid. From glass, and uS 'as likeness ; the capsule of the vitreous humour of the eye is so called, from its trans- parent and glassy appearance. Hymen. The membrane situated at the entrance of the vir- gin vagina ; from Hymen, the god of marriage. Hyoides. A bone of the, tongue, so called from its resem- blance to the Greek v ; from v, and resemblance. 419 Glossary. Hypochondrium. That part of the body which lies under the cartilages of the spurious ribs ; from uxa under, and a cartilage. ! Hypogastric. The lower region of the fore part of the abdo- men ; from v7ro under, and y« the stomach. I. lleon. A portion of the small intestines ; from uXea to turn ; being always convoluted. Ischium. The part of the os innominatum upon which we sit; from nr%va to sustain. L. Lacuna. The excretory duct of the glands of the urethra and vagina ; from lacus a channel. Lambdoidal suture. So called because it is shaped like the letter A ; from A, and etfos resemblance. Larynx. The superior part of the windpipe; the larynx. , M. Masseter. A muscle of the face, which assists in the action of chewing; paro-cupai to chew. Mastoid. From yM?o to, extend. Thorax. The breast or chest. Thyroid. From Stress a shield, and uctog likeness; shaped like a shield. Trachea. The windpipe ; so called from its roughness ; from TQv.yju', rough. Trochanter. A process of the thigh-bone, so called from r^oyag a wheel. U. Ulna. A name for the cubit ; from aXew the cubit. Ureter. The canal which conveys the urine from the kidney to the bladder; from xgov urine. Urethra. The passage through which the urine passes from the bladder ; from the urine. Uvea. The posterior lamen of the iris, so called because in many animals it is of the colour of unripe grapes ; from uva an unripe grape. Uvula. The conical substance which hangs down from the middle of the soft palate : so called from its resemblance to a grape. A dim. of uva a grape. y. Valves. From valvse folding doors. Vertebrae. The bones of the spine are so called ; from verto to turn. X. Xiphoid. So called from the resemblance to a sword ; from | i my y Pla t Explanation of the Plates of Osteology. 43? Fig. 4. The Bones of the Right Hand, with the Palm in view. a The radius, b, The ulna, c, The scaphoid bone of the carpus, d, The os lunare- e, The os cuneiforme. f, The os pisiforme. g, Trapezium, h, Trapezoides. i, Magnum, k, Unciforme. 1, The four metacarpal bones of the fingers, m, The first phalanx, n, The second phalanx, o, The third phalanx, p, The metacarpal bone of the thumb, q, The first joint, r, The second joint. Fig. 5 . The posterior View of the bones of the Left Hand. The explication of Fig. 4. serves for this figure; the same letters pointing out the same bones, though in a different view. Fig. 6. The Upper Extremity of the Tibia, with the Semilunar Cartilages of the Joint of the Knee, an*l some Ligaments. a, The strong ligament which connects the rotulato the tu- bercle of the tibia, bb, The parts of the extremity of the ti- bia, covered with cartilage, which appear within the semilu- nar cartilages, cc, The semilunar cartilages, d, The two parts of what is called the cross ligament. Fig. 7. The Posterior view of the Joint of the Right Knee. a, The os femoris cut. b, Its internal condyle, c, Its ex- ternal condyle, d, The back part of the tibia, e, The superior extremity of the fibula, f, The edge of the internal semilunar cartilage, g, An oblique ligament, h, A large perpendicular ligament, i, A ligament connecting the femur and fibula. Fig. 8. The Anterior view of the Joint of the Right Knee. b, The internal condyle, c, Its external condyle, d, The part of the os femoris, os which the patella moves, e, A per- pendicular ligament. * fit 1 , The two parts of the crucial liga- ments. gg, The edges of the two moveable semilunar carti- lages. h, The tibia, i, The strong ligament of the patella. k, The back part of it where the fat has been dissected away, l, The external depression, m, The internal one. n, The cut tibia. Fie. 9. A View of the inferior part of the Bones of the Right Foot. a, The great knob of the os calcis. b, A prominence on its outside, c, The hollow for the tendons, nerves and blood ves - sels. d, The anterior extremity of the os calcis. e, Part of the astragalus, f, Its head covered with cartilage, g, The internal prominence of the os naviculare. h, The os cuboides. i, The os cuneiform internum ; k, — Medium : 1, — Externum, m, The metatarsal bones of the four lesser toes, n, The first — o, The second — p, The third phalanx of the four lesser toes, q, 438 Explanation of the Plates, of Osteology . The metatarsal bones of the great toe. r, Its first — s, Its se- cond joint. Fig. 10. The Inferior Surface of the two large Sesamoid Bones, at the first Joint of the Great Toe. Fig. 11. The Superior View of the Bones of the Right Foot, a, b, as in Fig. 9. c, The superior head of the astragalus. d, &c. as in Fig. 9. Fig. 12. The view of the Sole of the Foot, with its ligaments. a, The great knob of the os calcis. b, The hollow for the tendons, nerves, and blood vessels, c, The sheaths of the flexores pollicis and digitorum long! opened, d, The strong cartilaginous. ligament supporting the head of the astragalus. e, h, Two ligaments which unite into one, and are fixed to the metatarsal bone of the great toe. f, A ligament from the knob of the os calcis to the metatarsal bone of the little toe. g, A strong triangular ligament, which supports the bones of the tarsus, i, The ligaments of the joints of the five metatarsal bones. Fig. 13. a, The head of the thigh bone of a child, b, The lmamentum rotundum connecting it to the acetabulum, c, The capsular ligament of the joint with its arteries injected, d, The numerous vessels of the mucilaginous gland injected. Fig. 14. The Back-view of the Cartilages of the Larynx, with the Os Hyoides. a, The posterior part of the base of the os hyoides. bb, Its cornua, c, The. appendix of the right side, d, A ligament sent out from the appendix of the left side, to the styloid process of the temporal bone, e, The union of*the base with the left cornu, ff, The posterior sides of (g) the thyroid cartilage, hh, Its superior cornua, ii, Its inferior cornua, k, The cri- coid cartilage. 11, The arytenoid cartilages, m, The entry into the lungs, named glottis, n, The epiglottis, oo, The superior cartilages of the trachea, p, Its ligamentous back part. Fig. 15. The superior Concave surface of the Sesamoid Bones at the first joint of the Great Toe, with their Ligaments, a, Three sesamoid bones, b, The ligamentous substance in which they are formed. / Explanation of the Plates of the Muscles. 439 EXPLANATION OF PLATES VI. and VII. PLATE VI. Fig. I. The Muscles immediately under the common te- guments on the anterior part of the body are represented on the right side; and on the left side the Muscles are seen which come in view when the exterior ones are taken away. A, The frontal muscle. B, The tendinous aponeurosis which joins it to the occipital; hence both named occipitofrontalis . C, Attollens aurem. D, The ear. E, Anterior auris. FF, Or- bicularis palpebrarum. G, Levator labii superioris alaeque nasi. H, Levator anguli oris. I, Zygomaticus minor. K, Zy- gomaticus major. L 5 Masseter. M, Orbicularis oris. N, De- pressor labii inferioris. O, Depressor anguli oris. P, Bucci- nator. QQ, Platysma myoides. RR, Sterno-cleido-mastol- deus. S, Part of the trapezius. T, Part of the scaleni. Superior Extremity. — U, Deltoides. V, Pectoralis ma- jor. W, Part of the latissimus dorsi. XX, Biceps flexor cu- biti. YY, Part of the triceps extensor. ZZ, The beginning of the tendinous aponeurosis, (from the biceps,) which is spread over the muscles of the fore-arm. aa, Its strong tendon insert- ed into the tubercle of the radius, bb, Part of the brachialis in- ternus. c, Pronator radii teres, d, Flexor carpi radialis. e, Part of the flexor carpi ulnaris. f, Palmaris longus. g, Apo- neurosis palmaris. 3, Palmaris brevis. 1, Ligamentum carpi annulare. 2 2, Abductor minimi digiti. h, Supinator radii lon- gus. i, The tendons of the thumb, k, Abductor -pollicis. 1, Flexor pollicis longus. mm, The tendons of the flexor sublimis perforatus, profundus perforans,|andlumbricales. The sheaths are entire in the right hand, — in the left cut open to show the tendons of the flexor profundus perforating the sublimis. Muscles not referred to — in the left superior extremity. — n, Pectoralis minor, seu serratus anticus minor, o, The two heads of (xx) the biceps, p, Coraco-brachialis. qq, The long head of the triceps extensor cubiti. rr, Teres major, ss, Sub- scapularis. tt, Extensores radiales. u, Supinator brevis, v, The cut extremity of the pronator teres, w, Flexor sublimis perforatus. x, Part of the flexor profundus, y. Flexor pol- licis longus. z, Part of the flexor pollicis brevis. 4, Abduc- tor minimi digiti. 5, The four lumbricales. Trunk.— 6 , Serrated extremities of the serratus anticus ma- 440 Explanation of the Plates of the Muscles , jor. 7 7, Obliquus externus abdominis. 8 8, The linea alba. 9, The umbilicus. 10, Pyramidalis. 11 11, The spermatic cord. On the left side it is covered by the cremaster- 12 12, Rectus abdominis. 13, Obliquus internus. 14 14, &c. In- tercostal muscles. Inferior Extremities. — a a , The gracilis, b b. Parts of the triceps, c c, Pectinalis. d d, Psoas magus, e e, Iliacus in- ternus. /, Part of the glutasus medius. g, Part of the glutseus minimus. li, Cut extremity of the rectus femoris. ii, Vastus externus. k, Tendon of the rectus femoris. 1 1, Vastus inter- nus. # Sartorius muscle. **. Fleshy origin of the tensor vaginas femoris or membranosus. Its tendinous aponeurosis covers («) the vastus externus on the right side, m in, Patella, nn, Ligament or tendon from it to the tibia, o, Rectus femoris. p, Crurasus. q q, The tibia, r r, Part of the Gemellus, or gas- trocnemius externus. sss , Part of the soleus or gastrocnemius internus. t, Tibialis auticus. u , Tibialis posticus, v v , Pero- naei muscles, tv w, Extensor longus digitorurn pedis, a; a;, Ex- tensor longus pollicis pedis, y, Abductor pollicis pedis. Fig. 2. The Muscles, Glands, &c. of the Left Side of the face and neck, after the common Teguments and Platys- ma myoides have been taken off. a, The frontal muscle, b, Temporalis and temporal arte- ry. c, Orbicularis palpebrarum, d, Levator labii superioris alaeque nasi, e, Lavator anguli oris, f, Zygomaticus. g, Depressor labii inferioris. h, Depressor anguli oris, i, Buc- cinator. k, Masseter. 1 1, Parotid gland, m, Its duct, n, Sterno-cleido-mastoideus. o, Part of the trapezius, p, Ster- no-hyoideus. q, Sterno-thyroideus. r, Omo-hyoideus. f, Le- vator scapulae, tt, Scaleni. u, Part of the splenius. Fig. 3. The Muscles of the Face and Neck in view after the exterior ones are taken away, aa, Corrugator supercilii. b, Temporalis, c, Tendon of the levator palpebrae superioris. - d, Tendon of the orbicula- ris palpebrarum, e, Masseter. f, Buccinator, g, Levator anguli oris, h, Depressor labii superioris alseque nasi, i, Or- bicularis oris, k, Depressor anguli oris. 1, Muscles of the os hyoides. m, Sterno-cleido-mastoideus. Fig. 4. Some of the Muscles of the Os Hyoides and Sub- maxillary Gland. a, Part of the masseter muscle, b, Posterior head of the digastric, c, Its anterior head, d d, Sterno-hyoideus. e, Omo- hyoideus. f, Stylo-hyoideus. g, Submaxillary gland in situ f; c A i v';c Anatomy Explanation of the Plates of the Muscles. 441 Fig. 5. The Submaxillary Gland and Duct. a, Musculus mylo-hyoideus. b, Hyo-glossus. c, Submax- Illary gland removed from its place, d, Its duct. Plate VII. Fig. 1. The Muscles immediately under the common te- guments on the posterior part of the body, are represented on the right side ; and on the left side the Muscles are seen which come in view when the exterior ones are taken away. Head. — AA, Occipito-frontalis. B, Attollens aurem. C, Part of the orbicularis palpebrarum. D, Masseter. E, Ptery- goideus interims. Trunk. — Right side. FFF, Trapezius seu cucullaris. GGGG, Latissimus dorsi. H, Part of the obliquus externus abdominis. Trunk. — Leftside. I, Splenius. K, Part of the cotnplexus. L, Levator scapulas. M, Rbomboides. NN, Serratus posticus inferior. O, Part of the longissimus dorsi. P, Part of the sacro-lumbalis. Q, Part of the semi-spinalis dorsi. R, Part of the serratus anticus major. S, Part of the obliquus internus abdominis. Superior Extremity. — Right side. T, Deltoides. U, Triceps extensor cubiti. V, Supinator longus. WW, Ex- tensores carpi radialis longior and brevior. XX, Extensor carpi ulnaris. YY, Extensor digitorum communis. Z, Ab- ductor indicis. 12 3, Extensores pollicis. Superior Extremity. — Left side, a, Supra Spinatus. b, Infraspinatus, c, Teres minor, d, Teres major, e, Tri- ceps extensor cubiti. ff, Extensores carpi radialis. g, Supi- nator brevis, h, Indicater. 12 3, Extensores pollicis. i, Ab- ductor minimi digiti. k, Interossei. Inferior Extremity. — Right side. 1, Glutseus maximus. m, Part of the Glutseus medius. n, Tensor vaginae femoris. o. Gracilis, pp, Adductor femoris magnus. q, Part of the vastus internus. r, Semimembranosus, s, Semitendinosus. t, Long head of the biceps flexor cruris, uu, Gastrocnemius externus seu gemellus, v, Tendo Achillis. w, Soleus seu gastrocne- miusfeternus. xx, Peronasus longus and brevis, y, Tendons of the flexor longus digitorum pedis; — and under them *flexor brevis digitorum pedis, z, Abductor minimi digiti pedis. Inferior Extremity. — Left side, m, n, o, pp, q, r, s, t , v, ww , xx, y, z, Point the same parts as in the right side, a, Pyri- formis. bb, Gemini, cc, Obturator internus. d, Quadratus fe- moris. e, Coceygaeus. /, The short head of the biceps flexor cru- ris. gg, Plantaris. /i 3 Popliteus. i, Flexor longus pollicis pedis , Vol. n. 56 442 Explanation of the Plates of the Muscles „ Fig. 2. The Palm of the Left Hand after the common Tegu- ments are removed, to show the Muscles of the Fingers, a, Tendon of the flexor carpi radialis. b, Tendon of the flex- or carpi ulnaris. c, Tendons of the flexor sublimis perforatus, profundus perforans and lumbricales. d, Abductor pollicis. ee, Flexor pollicis longus. f. Flexor pollicis brevis, g, Palma- ris brevis, h, Abductor minimi digiti. i, Ligamentum carpi- annulare. k, A probe put under the tendons of the flexor di~ gitorum sublimis; which are performed by 1, the flexor digito- rum profundus, mmmm, Lumbricales. n, Abductor pollicis. Fig. 3. A fore-view of the foot and Tendons of the flexores Digitorum. a, Cut extremity of the tendo Achillis. b, Upper part of the astragalus, c, Os calcis. d, Tendon of the tibialis anticus. e, Tendon of the extensor pollicis longus. f, Tendon of the peroneus brevis, g, Tendons of the flexor digitorum longus, with the non us Vesalii. hh, The whole of the flexor digitorum brevis. Fig. 4. Muscles of the Anus, aa, An outline of the buttocks, and upper part of the thighs, b, The testes contained in the scrotum, cc, Sphincter ani. d, Anus, e, Levator ani. ff, Erector penis, gg, Accelera- tor urinae. h, Corpus cavernosum urethrae. Fig. 3. Muscles of the Penis. aa, b, d, ee, ff, h, point the same as in fig. 4. c, Sphincter ani. gg, Transversalis penis. EXPLANATION of PLATES VIII. IX. and X. Plate VIIL Fig. 1. Shows the Contents of the Thorax and Abdomen In situ. l, Top of the trachea, or windpipe. 2 2, The internal ju- gular veins. 3 3, The subclavian veins. 4, The vena cava de- scendens. 5, The right auricle of the heart. 6, The right ven- tricle. 7, Part of the left ventricle. 8, The aorta descendens. 9, The pulmonary artery. 10, The right lung, part of which, is cut off to show the great blood vessels. 11, The left lung entire. 12 12, The anterior edge of the diaphragm. 1313, The two great lobes of the liver. 14, The ligamentum rotun- dum. 15, The gall bladder. 16, The stomach. 1717, The jejunum and ilium. 18, The spleen. ANj\ T oMY Explanation of the Plates of the Muscles. 443 Fig. 2. Shows the organs subservient to the Chylopoietic Vis- cera, — with those of Urine and Generation. 11, The under side of the two great lobes of the liver, a, Lobulus spigelii. 2, The ligamentum rotundum. 3, The gall- bladder. 4, The pancreas. 5, The spleen. 6 6, The Kid- neys. 7, The aorta descendens. 8, Vena cava ascendens. 9 9, The renal veins covering the arteries. 10, A probe un- der the spermatic vessels and a bit of the inferior mesentric artery, and over the ureters. 1 1 11, The ureters. 12 12, The iliac arteries and veins. 13, The intestinum rectum. 14, The bladder of urine. Fig. 3. Shows the Chylopoietic Viscera, and Organs subser- vient to them, taken out of the body entire. AA, The under side of the two great lobes of the liver. B, Ligamentum rotundum. C, The gall-bladder. D, Ductus cysticus. E, Ductus hepaticus. F, Ductus communis chole- dochus. G, Vena portarum. H, Arteria hepatica. I, The stomach. KK, Vense and arterise gastro-epiploicae, dextrae and sinistrae. LL, Venae and arteriae coronariae ventriculi. M, The spleen. NN, Mesocolon, with its vessels. 000, In- testinum colon. P, One of the ligaments of the colon, which is a bundle of longitudinal muscular fibres. QQQQ, Jejunum and ilium. RR, Sigmoid flexure of the colon with the liga- ment continued, and over. S, The intestinum rectum. TT, Levatores ani. U, Sphincter ani. V, The place to which the prostate gland is connected. W, The anus. Fig. 4. Shows the Heart of a Foetus at the full time, with the Right Auricle cut open to show the Foramen Ovale, or passage between both Auricles. a, The right ventricle, b, The left ventricle, cc, The out- er side of the right auricle stretched out. dd, The posterior side, which forms the anterior side of the septum, e, The foramen ovale, with the membrane or valve which covers the left side, f, Vena cava inferior passing through, g, A por- tion of the diaphragm. Fig. 5. Shows the Heart and Large Vessels of a Foetus at the full time. a, The left ventricle, b, The right ventricle, c, A part of the right auricle, d, Left auricle, ee, The right branch of the pulmonary Artery, f, Arteria pulmonalis. gg, The left branch of the pulmonary artery, with a number of its largest branches dissected from the lungs, h, The canalis arteriosus, i, The arch of the aorta, kk, The aorta descendens. 1, The left sub- 444 Explanation of the Plates of the Muscles. clavian artery, m, The left carotid artery, n, The right caro- tid artery. o, The right subclavian artery, p, The origin of the right carotid and right subclavian arteries in one common trunk, q, The vena cava superioror descendens. r, The right common subclavian vein, s, The left common subclavian vein. N. B. All the parts described in this figure are to be found in the adult, except the canalis arteriosus. Plate IX. Fig. 1 . Exhibits the more superficial Lymphatic Vessels of the Lower Extremity. A, The spine of the os ilium. B, The os pubis, c, The iliac artery. D, The knee. E, E, F, Branches of the crural ar- tery. G, The musculus gastrocnemius. H, The tibia. I, The tendon of the musculus tibialis anticus. On the outlines, a, A lymphatic vessel belonging to the top of the foot, b, Its first division into branches, c, c, c, Other divisions of the same lymphatic vessel, d, A small lymphatic gland, e, The lymphatic vessels which lie between the skin and the muscles of the thigh, ffj Two lymphatic glands at the upper part of the thigh below the groin, gg, Other glands, h, A lympha- tic vessel which passes by the side of those glands without communicating with them; and bending towards the inside of the groin at (i,) opens into the lymphatic gland (k.) 11, Lymphatic glands in the groin, which are common to the lym- phatic vessels of the genitals and those of the lower extremi- ty. m, n, A plexus of lymphatic vessels passing on the in- side of the iliac artery. Fig. 2. Exhibits a Back-view of the Lower Extremity dis- sected so as to show the deeper-seated Lymphatic Vessels which accompany the Arteries. A, The os pubis. B, The tuberosity of the ischium. C, That part of the os ilium which was articulated with the os sacrum. D, The extremity of the iliac artery appearing above the groin. E, The knee. FF, The two cut surfaces of the triceps muscle, which was divided to show the lymphatic vessels that pass through its perforation along with the crural artery. G, The edge of the musculus gracilis. H, The gastrocnemius and so- leus much shrunk by being dried, and by the soleus being se- parated from the tibia to expose the vessels. I, The heel. K, The sole of the foot. L, The superficial lymphatic vessels passing over the knee, to get to the thigh. On the outlines ; M, The posterior tibial artery, a, A lymphatic vessel accom- panying the posterior tibial artery, b, The same vessel cross- ing the artery, c, A small 1 vmphatic gland, through which this 4 Explanation of the Plates of the Muscles. 445 deep-seated lymphatic vessel passes, d, The lymphatic vessel passing under a small part of the soleus, which is left attached to the bone, the rest being removed. e, The lymphatic vessel crossing the popliteal artery, f, g, h, Lymphatic glands in the ham,, through which the lymphatic vessel passes, i, The lym- phatic vessel passing with the crural artery, through the perfo- ration of the triceps muscle, k, The lymphatic vessel, after it has passed the perforation of the triceps, dividing into branches which embrace the artery (1.) m, A lymphatic gland belonging to the deep-seated lymphatic vessel. At this place those vessels pass to the fore part of the groin where they communicate with the superficial lymphatic vessel, n, A part of the superficial lymphatic vessel appearing on the brim of the pelvis. Fig. 3. Exhibits the Trunk of the Human Subject prepared to show the Lymphatic Vessels and the Ductus Thoracicus. A, The neck. BB, The two jugular veins. C, The vena cava superior. DDDD, The subclavian veins. E, The begin- ning of the aorta, pulled to the left side by means of a ligature, in order to show the thoracic duct behind it. F, The branches arising from the curvature of the aorta. GG, The two carotid arteries. HH, The first ribs. II, The trachea. KK, The spine. LL, The vena azygos. MM, The descending aorta. N, The cceliac artery, dividing into three branches. 0, The superior mesenteric artery. P, The right crus diaphragm atis. QQ, The two kidneys. R, The right emulgent artery. SS, The external iliac arteries, g, d, The psoas muscles. T, The internal iliac artery. U, The cavity of the pelvis. XX, The spine of the os ilium. YY, The groins, a, A lymphatic gland in the groin, into which lymphatic vessels from the lower extremity are seen to enter, bb, The lymphatic vessels of the lower extremities passing under Poupart’s ligament, cc, A plexus of the lympha- tic vessels lying on each side of the pelvis, d, The psoas muscle with lymphatic vessels lying upon its inside, e, A plexus of lymphatics, which having passed over the brim of the pelvis at (c,) naving entered the cavity of the pelvis, and received the lymphatic vessels belonging to the viscera contained in that ca- vity, next ascends and passes behind the illiac artery to (g.) f Some lymphatic vessels of the left side passing over the upper part of the os sacrum: to meet those of the right side, g, The right psoas, with a large plexus of lymphatics lying on its inside. M, The plexus lying on each side of the spine. Hi, Spaces occupied by the lymphatic glands, k, The trunk of the lacteals lying on the under side of the superior mesenteric artery. /, The same dividiug into two branches, one of which passes on each 446 Explanation of the Plates of the Muscles. side of the aorta; that of the right side being seen to enter the thoracic duct at ( m .) m, The thoracic duct beginning from the large lymphatics, n, The duct passing under the lower part of the crus diaphragmatis, and under the right emulgent artery, o, The thoracic duct penetrating the thorax, p, Some lymphatic vessels joining that duct in the thorax, q, The tho- racic duct passing under the curvature of the aorta to get to the left subclavian vein. The aorta being drawn aside to show the duct, r, A plexus of lymphatic vessels passing upon the trachea from the thyroid gland to the thoracic duct. Plate X. Fig. 1 . Represents the Under and Posterior Side of the Bladder of Urine, &c. a, The bladder, bb, The insertion of the ureters, cc, The vasa deferentia, which convey the semen from the testicles to dd. The vesiculae seminales, — and pass through e. The pros- tate gland, to discharge themselves into f. The beginning of the urethra. Fig. 2. A transverse Section of the Penis. gg, Corpora cavernosa penis, h, Corpus cavernosum ure- thrae. i. Urethra, k, Septum penis. 11, The septum be- tween the corpus cavernosum urethrae and that of the penis. Fig. 3. A Longitudinal Section of the Penis. mm, The corpora cavernosa penis, divided by o, The sep- tum penis, n, The corpus cavernosum glandis, which is the continuation of that of the urethra. Fig. 4. Represents the Female Organs of Generation. a, That side of the uterus which is next to the os sacrum. 1, Its fundus. 2, Its cervix, bb, The fallopian or uterine tubes, which open into the cavity of the uterus; — but the other end is open within the pelvis, and surrounded by c c, The fim- briae. d d, The ovaria. e, The os internum uteri, or mouth of the womb, ff, The ligamentum rotundum, which passes without the belly, and is fixed to the labia pudendi. gg, The cut edges of the ligamenta lata, which connect the uterus to the pelvis, h. The inside of the vagina, i, The orifice of the urethra, k, The clitoris surrounded by (1,) The praeptium. mm, The labia pudendi. nn, The nympbae. Fig. 5. Shows the Spermatic Ducts of the Testicle filled with Mercury. A, The vas deferens. B, Its beginning, which forms the /< w . Akat omt A N ATOMY Explanation of Vie Plates of the Muscles . 447 posterior part of the epididymis. C, The middle of the epi- didymis, composed of serpentine ducts. D, The head or an- terior part of the epididymis unravelled, e e e e, The whole ducts which compose the head of the epididymis unravelled, f f, The vasa deferentia. g g, Rete testis, h h, Some recti- linear ducts which send off the vasa deferentia. i i, The sub- stance of the testicle. Fig. 6. The right Testicle entire, and the Epididymis filled with Mercury. A, The beginning of the vas deferens. B, The vas deferens ascending towards the abdomen. C, The posterior part of the epididymis, named globus minor. D, The spermatic vessels enclosed in cellular substance. E, The body of the epididy- mis. F, Its head, named globus major. G, Its beginning from the testicle. H, The body of the testicle, enclosed in the tu- nica albuginea. EXPLANATION of PLATE XI. This plate represents the Heart in situ, all the large Arte- ries and Veins, with some of the Muscles, &c. Muscles, &c. — Superior Extremity. — a, Masseter. b, Complexus. C, Digastricus. d, Os hyoides. e, Thyroid gland, f, Levator scapulae, g, Cucullaris. h h, The clavicles, cut. i. The deltoid muscle, k, Biceps flexor cubiti. cut. 1, Coraco- brachialis. m, Triceps extensor cubiti. n, The heads of the pronator teres, flexor carpi radiales, and flexor digitorum sub- limis, cut. o, The flexor carpi-ulnaris, cut at its extremity. p, Flexor digitorum profundus, q, Supinator radii longus, cut at its extremity, r, Ligamentum carpi transversale. s, Extensores carpi radiales. t, Latissimus dorsi. u, Anterior edge of the serratus anticus major, vv, The inferior part of the diaphragm, ww, Its anterior edge cut. xx, The kidneys, y, Transversus abdominus. z, Os ilium. Inferior Extremity. — a, Psoas magnus. b, Iliacus inter- nus. c, The fleshy origin of the tensor vagina femoris. dd } The ossa pubis cut from each other, e, Musculus pectineus cut from its origin, f, Short head of the triceps abductor fe- moris cut. g, The great head of the triceps, h, The long head cut. i, Vastus internus. k, Vastus externus. I, Cru- reus. m, Gemellus, n, Soleus. o, Tibia, p, Peroneus longus. q, Peroneus brevis, r, Fibula. Heart and Blood-vessels. — A, The heart, with the co- ronary artery and veins. B, The right auricle of the heart. C, The aorta ascendens. D, The left subclavian artery. E, The 448 Explanation of the Plates of the Muscles. left carotid artery. F, The common trunk which sends oft' the right subclavian and right carotid arteries. G, The carotis ex- terna. H, Arteria facialis, which sends off the coronary arte- ries of the lips. I, Arteria temporalis profunda. K, Aorta de- scendens. LL, The illiac arteries, which sends oft' MM, The femoral or crural arteries. N. B. The other arteries in this figure have the same distribution as the veins of the same name : — And generally, in the anatomical plates, the descrip- tion to be found on the one side, points out the same parts in the other. 1, The frontal vein. 2, The facial vein. 3, Vena temporalis profunda. 4, Vena occipitalis. 5, Vena jugulads externa. 6, Vena jugularis interna, covering the arteria caro- tis communis. 7, The vascular arch on the palm of the hand, which is formed by, 8, The radial artery and vein, and, 9, The ulnar artery and vein. 10 10, Cephalic vein. 1 1, Basilic vein, that on the right side cut. 12, Median vein. 13, The humeral vein, which, with the median, covers the humeral ar- tery. 14 14, The external thoracic or mammary arteries and veins. 15, The axillary vein, covering the artery. 16 16, The subclavian veins, which, with (6 6) the jugulars, form, 17, The vena cava superior. 18, The cutaneous arch of veins on the fore part of the foot. 19, The vena tibialis antica, covering the artery. 20, The vena profunda femoris, covering the artery. 21, The upper part of the vena saphena major. 22, The fe- moral vein. 2323, The illiac veins. 24 24, Vena cava infe- rior. 25 25, The renal veins covering the arteries. 26 26, The diaphragmatic veins. EXPLANATION of PLATE XII. Fig. 1 . Represents the Inferior part of the Brain; — the An- terior part of the whole Spine, including the Medulla Spina- lis; — with the origin and large portions of all the Nerves. AA, The anterior lobes of the cerebrum. BB, The lateral lobes of the cerebrum. CC, The two lobes of the cerebellum. D, Tuber annulare. E, The passage from the third ventricle to the infundibulum. F, The medulla oblongata, which sends off the medulla spinalis through the spine. GG, That part of the os occipitis which is placed above (HH) the transverse processes of the first cervical vertebra. II, &c. The seven cervical vertebrae, with their intermediate cartilages. KK, &c. The twelve dorsal vertebrae, with their intermediate car- tilages. LL, &c. The five lumbar vertebrae, with their inter- mediate cartilages. M, The os sacrum. N, The os coccygis. Ill* Explanation of the Plates of the Muscles . 449 Nerves. — 1 1, The first pair of nerves, named olfactory 9 which go to the nose. 2 2, The second pair named optic , which goes to form the tunica retina of the eye. 3 3, The third pair, named motor oculi ; it supplies most of the muscles of the eye-ball. 4 4, The fourth pair, named pathetic , — which is wholly spent upon the musculus trochlearis of the eye. 5 5, The fifth pair divides into three branches. — The first, named ophthalmic, goes to the orbit, supplies the lachrymal gland, and sends branches out to the forehead and nose. — The second, named superior maxillary, supplies the teeth of the upper jaw, and some of the muscles of the lips. — The third, named inferior maxillary, is spent upon the muscles and teeth of the lower jaw, tongue, and muscles of the lips. 6 6, The sixth pair, which after sending oft' the beginning of the intercostal or great sympathetic, is spent upon the abductor oculi. 7 7, The seventh pair, named auditory, divides into two branches.-- The largest, named portio mollis , is spent upon the internal ear. The smallest, portio dura , joins to the fifth pair within the internal ear by a reflected branch from the second of the fifth ; and within the tympanum, by a branch from the third of the fifth, named chorda tympuni. — >Vid. fig. 3. near B 8 8, &c. The eighth pair, named par vugum, — which accompanies the intercostal, and is spent upon the tongue, larynx, pharynx, lungs, and abdominal viscera. 9 9, The ninth pair, which are spent upon the tongue. 10 10, &c. The intercostal, or great sympathetic, which is seen from the sixth pair to the bottom of the pelvis on each side of the spine, and joining with all the nerves of the spine; — In its progress supplying the heart, and, with the par vagum, the contents of the abdomen and pelvis. 1111, The accessorius, which is spent upon the ster- no-cleido-mastoideus and trapezius muscles. 12 12, The first cervical nerves; — 13 13, The second cervical nerves; — both spent upon the muscles that lie on the neck, and teguments of the neck and head. 1414, The third cervical nerves, which, after sending off, (15 15, &c.) the phrenic nerves to the dia- phragm, supply the muscles and teguments that lie on the side of the neck and top of the shoulder. 16 16, The brachial plexus, formed by the fourth, fifth, sixth, seventh cervicals, and first dorsal nerves : which supply the muscles and tegu- ments of the superior extremity. 17 17, The twelve dorsal, or proper intercostal nerves, which are spent upon the inter- costal muscles and some of the large muscles which lie upon the thorax. IS 18, The five lumbar pairs of nerves, which supply the lumbar and abdominal muscles, and some of the teguments and muscles of the inferior extremity. 19 19. The Vol. ii, 57 450 Explanation of the Plates of the Muscles, sacro-sciatic or posterior crural nerve, formed by the two in- ferior lumbar, and three superior of the os sacrum. This large nerve supplies the greatest part of the muscles and teguments of the iuferior extremity. 20, The stomachic plexus, formed by the eighth pair. 21 21, Branches of the solar or cceliae plexus, formed by the eighth pair and intercostals, which sup- ply the stomach and chylopoietic viscera. 22 22, Branches of the superior and inferior mesenteric plexuses, formed by the eighth pair and intercostals, which supply the chylopoietic vis- cera, with part of the organs of urine and generation. 23 23, Nerves which accompany the spermatic cord. 24 24, The hy- pogastric plexus, which supplies the organs of urine and gene- ration within the pelvis. Fig. 2, 3, 4, 5, Show different Views of the Inferior part of the Brain, cut perpendicularly through the Middle, — with the Origin and large Portions of all the Nerves which pass but through the Bones of the Cranium,— and the three first Cervicals. A, The anterior lobe. B, The lateral lobe of the cerebrum, C, Onehrf the lobes of the cerebellum. D, Tuber annulare. E, Corpus pyramidale, in the middle of the medulla oblongata, F, The corpus olivare, in the side of the medulla oblongata, G, The medulla oblongata. H, The medulla spinalis. Nerves.— 1 23 45 6 7 8 and 9, Pairs of Nerves. 10 10, Nerves accessorius, which comes from — 11, 12, and 13, The three first cervical nerves. EXPLANATION of PLATE XIII. Fig. 1. Shows the Lachrymal Canals, after the Common Te- guments and Bones have been cut away, a, The lachrymal gland, b, The tivo puncta lachrymalia, from which the two lachrymal canals proceed to c, The lach- rymal sac. d, The large lachrymal duct, e, Its opening into the nose, f, The carunca lachrymal is. g, The eye-ball. Fig. 2, An interior View of the Coats and Humours of the Eye. a a a a, The tunica sclerotica cut in four angles, and turned back, bbbb, The tunica clioroides adhering to the inside of the sclerotica and the ciliary vessels are seen passing over — cc. The retina which covers the vitreous humour, dd, The ciliary processes, which were continued from the choroid coat, a e, The iris f, The pupil. Anatomy plate xm . Explanation of the Plates of the Muscles. 451 Fig. 3. Shows the Optic Nerves, and Muscles of the Eye. a, a. The two optic nerves before they meet, b, The two optic nerves conjoined, c, The right optic nerve, d, Muscu- lus. attollens palpebrse superioris. e, Attollens oculi. f, Ab- ductor. gg, Obliquus superior, or trochlearis. h, Abductor, i, The eye-ball. Fig. 4. Shows the Eye-ball with its Muscles, a, The optic nerve, b, Musculus trochlearis. c, Part of the os frontis, to which the trochlea or pulley is fixed through which, — d, The tendons of the trochlearis pass, e, Attollens oculi. f, Adductor oculi. g, Abductor oculi. h, Obliquus in- ferior. i, Part of the superior maxillary bone to which it is fixed, k, The eye-ball. Fig. 5. Represents the Nerves and Muscles of the Right Eye, after part of the Bones of the orbit have been cut away. A, The eye-ball. B, The lachrymal gland. C, Musculus abductor oculi. D, Attollens. E, Levator palpebras superi- oris. F, Depressor oculi. G, Abductor. H, Obliquus superior, with its pulley. I, Its insertion into the sclerotic coat. K, Part of the obliquus inferior. L, The anterior part of the os frontis, cut. M, The crista galli of the ethmoid bone. N, The posterior part of the sphenoid bone. 0, Transverse spi- nous process of the sphenoid bone. P, The carotid artery, denuded where it passes through the bones. Q, The carotid artery within the cranium. R, The ocular artery. Nerves.— a a, The optic nerve,—- b, The third pair, c, Its joining with a branch of the first branch of the fifth pair, to form 1, — The lenticular ganglion, which sends off the ciliary nerves, d. ee, The fourth pair, f, The trunk of the fifth pair, g, The first branch of the fifth pair, named ophthalmic, h, The frontal branch of it. i, Its ciliary branches, along with which the nasal twig is sent to the nose, k, Its branch to the lachrymal gland. 1, The lenticular ganglion, m, The second branch of the fifth pair, named superior maxillary, n, The third branch of the fifth pair, named inferior maxillary, o, The sixth pair of nerves — which sends oft' p, The beginning of the great sympathetic, q, The remainder of the sixth pair, spent on c, The abductor oculi. Fig. 6. Represents the head of a youth, where the upper part of the cranium is sawed off, —to show the upper part of the brain, covered by the pia mater, the vessels of which are minutely filled with wax. AA, The cut edges of the upper part of the cranium. B. 452 Explanation of ike Plates of the Muscles. The two tables and intermediate diploe. BB, The two hemi- spheres of the cerebrum. CC, The incisure made by the falx, D, Part of the tentorium cerebello super expansum. E, Part of the falx, which is fixed to the crista galli. Fig. 7. Represents the parts of the External Ear, with the Parotid Gland and its Duct. a a, The helix, b, The antihelix, c. The antitragus, d, The tragus, e, The lobe of the ear. f, The cavitas innominata. g, The scapha. h, The concha, i i, The parotid gland, k, A lymphatic gland, which is often found before the tragus. 1, The duct of the parotid gland, m, its opening into the mouth. Fig. 8. A view of the posterior part of the external ear, mea- tus auditorius, tympanum with its small bones and Eusta- chian tube, of the right side. a, The back part of the meatus, with the small ceruminous glands, b, The incus, c, Malleus, d, The chorda tympani. e, Membrana tympani. f, The Eustachian tube, g, Its mouth from the fauces. Fig. 9. Represents the anterior part of the right external ear, the cavity of the tympanum — it9 small bones, cochlea, and semicircular canals. a, The malleus, b, Incus, with its long leg, resting upon the stapes, c, Membrana tympani. d, e, The Eustachian tube covered by part of — f f, The Musculus circumilexus palati. 1, 2, 3, The three semicircular canals. 4, The vestibule. 5, The cochlea. 6, The portio mollis of the seventh pair of nerves. Fig. 10. Shows the muscles which compose the fleshy sub- stance of the Tongue. a a, The tip of the tongue, with some of the papillse mini- mse. b, The root of the tongue, c, Part of the membrane of the tongue, which covered the epiglottis, d d, Part of the mus- culus hyo-glossus. e, The lingualis. f, Genio-glossus. g g. Part of the stylo-glossus. JgT I Wistar 1827 v. 2