-■^O'fll Cornell University Library QM 28.Y73 1913 Handbook of anatomy; being a complete com 3 1924 003 140 864 CORNELL. UNIVERSITY LIBRARY GIFT OF J. W. Papez The original of tiiis book is in tine Cornell University Library. There are no known copyright restrictions in the United States on the use of the text. http://www.archive.org/details/cu31924003140864 HANDBOOK OF ANATOMY BEING A COMPLETE COMPEN0 OF ANATOMY, INCLUDING THE ANATOMY OP THE VISCERA AND NUMEROUS TABLES JAMES K. YOUE'G-. M.D. Professor of Orthopaedic Surgery, Philadelphia Polyclinic; eiinical Professor of Orthopaedic Surgery, Woman's Medical College of Pennsylvania ; Associate Professor of Orthopaedic Surgery, University of Pennsylvania ; Fellow of the College of Physicians, of Philadelphia; Fellow of the Philadelphia Academy of Surgery ; Fellow of the Ameri- can Orthopaedic Association; Member of the American Medical Association, etc., etc. Fourth revised edition WITH 172 ENQRAVINQS, SOME IN COLORS PHILADELPHIA F. A. DAVIS COMPANY, Publishers 1913 Copyright, 1889 Copyright, 1905 Copyright, 1909 Copyright, 1913 BY P. A. DAVIS COMPANY 1 Registered at Stationers' Hall, London, Eng.] Philadelphia, Pa., U. S. A. Press of F. A. Davis Company 1914-1916 Cherry Street PREFACE TO FOURTH EDITION. Although for over twenty years the book has been in con- stant use by large numbers of students and approved by professors and teachers in leading medical colleges and in Xurses' Training Schools generally, comparatively few errors or omissions of im- portance have been discovered; but in the present edition these liave all been noted and corrected. The appreciative thanks of the author are extended to Dr. A. C. Pole, of Baltimore, for several pertinent suggestions in the preparation of this edition. J. K. Y. (iii) PREFACE TO THIRD EDITION. In the preparation of this new edition, the author has taken advantage of the opportunity to correct the few clerical errors which must necessarily be found in a comprehensive though condensed work upon Anatomy, and to make such changes as have become necessary through the advances of the science since the publication of the last edition. In the section upon the Xervous System the greatest changes will be found, the first part having been rewritten. In the description of the Muscular System, the regional grouping has been retained, because it is more practical for the student. Attention is directed to the original colored plates of the Arterial System which offer the most complete classification of all the arteries in the smallest space, and also to the original diagrams of the sensory tracts from spinal cord to brain, and the motor tracts from brain to spinal cord. The author desires to express his gratitude to the student body of America for their continued confidence in this early literary effort of his, and express the hope that the present edition may be as well received as the others. J. K. Y. (iv) PREFACE TO SECOND EDITION. The demand for a new editipn has afforded an opportunity to thoroughly revise the text and illustrations, and to add such changes of description and nomenclature as have been intro- duced into modern methods of teaching. Whenever it seemed necessary or advisable sections have been entirely rewritten, but the same attention to conciseness and accuracy has been observed as in the former edition. The size of the pages has been considerably enlarged and increased in number, and the illustrations increased from 76 to 171. Many improved cuts replace those used in the former edi- tion and full-page engraved plates have been inserted in the text where they will be most serviceable. In its preparation, the last editions of both Morris and Gray have been freely consulted, so that it can be used with either of these standard works. As in the former edition, although prepared particularly for students, sufficient descriptive matter has been added to render it valuable to the busy practitioner, particularly in the sections upon the Viscera, Special Senses, Vascular System, the Nervous System, and Surgical Anatomy. Special attention has been given to the anatomy of the brain and nervous system, which will be found to compare favorably with the descriptions in larger works. The author has been much gratified to observe the tenacity with which students and physicians have retained the work for reference. The author would acknowledge his obligation to Dr. Eichard F. Gerlach, who has distinguished himself by his careful and ' accurate work upon Deaver's "Surgical Anatomy," for valuable assistance upon the revision. J. K. Y. (V) PREFACE TO FIRST EDITION. The object of this little book is, as its title indicates, to furnish a concise though complete synopsis of human anatomy for the use of students of medicine and others. Appreciating, from a personal contact with students, the limited time at their disposal and the unlimited amount of medical material to be digested, the author has endeavored, by well-selected wood-cuts, typographical arrangement, and numer- ous tables, to facilitate the acquisition of a subject as difficult as it is essential, and elegance or diction has therefore been sacri- fieed to conciseness and accuracy. In its preparation the last edition of Gray's "Anatomy" [edited by Keen], as the work most largely employed, has been selected as the standard, but Leidy, Quain, Allen, Holden, Henle, SchafEer, Klein, Brown Aids, and others, have been freely used, and on the special subjects Lusk, Spiegelberg, Savage, Schroeder, Budin, Treve's "Surgical Applied Anatomy," and the "Ameri- can System of Dentistry" have been consulted. Although pre- pared particularly for students, sufficient descriptive matter has been added to render it valuable to the busy practitioner, par- ticularly the sections on the Viscera, Special Senses, Vascular System, and Surgical Anatomy. The aim throughout has been to make it as thoroughly complete and accurate as possible, and at the same time readily accessible for reference or study. The writer would acknowledge his obligations to l^Icssrs. S. Z. Durborow and C. W. Holly, medical students at the "Uni- versity, for assistance in the correction of proof. J. K. Y. (vi) TABLE OF CONTENTS. PAGE Intboductort 1 Osteology 3 The Frontal Bone 8 Parietals 10 The Occipital Bone 11 Temporals 13 The Sphenoid 17 The Ethmoid 21 The Lachrymal 22 The Malar 23 The Superior Maxilla 24 Palate Bones 26 The Inferior Turbinated Bones 28 The Vomer 28 The Inferior Maxilla 29 The Sutures and Fontanelles 31 Fossae of Skull 32 The Orbits 34 Foramina at the Base of the SkuU 38 The Sternum 41 The Ribs 41 The Spine 43 The Upper Extremity 47 The Arm SI The Forearm 53 The Hand 50 The Lower Extremity ' 59 The Thigh 63 The Patella 65 The I*g 65 The Foot 68 Table of Ossification 72 Abticui-ations and Ligaments Ill The Thorax 115 The Pelvis 118 (vii) viii TABLE OF CONTENTS. PAGE The Mtjsctjlab System 132 Muscles of the Head 133 Muscles of the Neck 137 Vertebral Region 140 Muscles of the Back 141 Muscles of the Abdomen 146 Muscles of the Upper Extremity 149 Muscles of the Forearm 152 Muscles of the Hand 155 Muscles of the Lower Extremity 157 Muscles of the Leg 161 Muscles of the Foot 161 The Fascias 166 The Heart and Vascular Ststem 170 The Circulation of the Blood 177 The Arterial System 173, 177 Table of thfe Arterial System 200 The Venous System 206 The Lymphatic System 214 The Alimentary Apparatus 218 The Teeth 219 The Pharynx 224 The CEsophagus 225 The Abdomen 226 The Peritoneum 228 The Stomach 231 The Small Intestine 233 The Large Intestine 235 The Pancreas 237 The Liver 238 The Ductless Glands 242 Vocal and Respiratory Apparatus 246 The Larynx 246 The Trachea and Bronchi 251 The Lungs 252 The Pleurse 255 The Mediastinum 256 The Genito-Urinaey Apparatus 257 The Kidneys 257 The Bladder 261 Male Organs 263 Female Organs 272 The Mammary Gland 281 TABLE OF CONTENTS. ix PAGE The Nebvotts System 283 The Brain 286 The Cranial Nerves 309 The Spinal Cord 317 The Spinal Nerves 318 Table of Spinal Nerves 328 The Sympathetic Nervous System 335 Organs of Special Sense 340 The Nose 340 The Eye 343 Appendages of the Eye 356 The Ear 359 The Tongue 371 The Skin and Appendages 372 Sdegical Anatomy 374 Triangles of the Neck 374 Triangle in Front of Elbow -joint 377 Scarpa's Triangle 377 AxiUa 377 Anatomy of Hernia 379 Ischio-rectal Region and Perineum 384 Anatomy of Lateral Lithotomy 390 Index 391 LIST OF ILLUSTRATIONS. FIS. PAGE 1. Frontal Bone, Outer Surface ^ 2. Frontal Bone, Inner Surlace 8 3. Parietal Bone, Outer Surface 9 4. Parietal Bone, Inner Surface 10 6. Occipital Bone, Outer Surface 11 6. Occipital Bone, Inner Surface 12 7. Left Temporal Bone, Outer Surface 14 8. Left Temporal Bone, Inner Surface 15 9. Sphenoid Bone, Upper Surface 18 10. Sphenoid Bone, Anterior Surface 19 11. Superior Maxillary Bone 23 12. Superior Maxillary Bone, Inner Surface 24 13. Palate Bone 26 14. Inferior Maxillary Bone 29 16. Fossae of Skull 33 16. Facial Portion of Skull 34 17. Floor of the Cranium 39 18. A Vertebra 43 19. Clavicle 47 20. The Scapula, Inner Surface 49 21. The Scapula. Outer Surface 50 22. The Humerus E3 23. The Ulna and Radius 53 24. The Carpal Bones 56 25. Os Innominatum 60 26. The Femur 63 27. The Tibia and Fibula, Anterior Surface 66 28. The Tibia and Fibula, Posterior Surface 66 29. The Tarsus, Metatarsus, and Phalanges 69 30. Frontal Bone, Outer Surface 77 31. Frontal Bone, Inner Surface 77 32. Parietal Bone, Outer Surface 78 83. Parietal Bone, Inner Surface 78 34. Occipital Bone, Outer Surface 79 35. Occipital Bone, Inner Surface 79 36. Left Temporal Bone, Outer Surface 80 37. Left Temporal Bone, Inner Surface 80 38. Left Temporal Bone, Inferior Surface of the Petrous Portion 81 39. Section of Left Temporal Bone 81 40. Sphenoid Bone, Upper Surface 82 41. Sphenoid Bone, Anterior Surface 82 42. Ethmoid Bone «3 43. Nasal Bones 83 44. Lachrymal Bone 83 45. Superior Maxillary Bone 84 46. Superior Maxillary Bone, Inner Surface 84 47. Vomer 85 48. Inferior Maxillary Bone 85 49. Left Half of Interior Maxillary Bone, Inner Surface 85 BO. Malar Bones 86 51. Palate Bone (Viewed Posteriorly) 86 52. Palate Bone (Inner Surface) 86 63. Inferior Turbinated Bones 86 54. Nasal Meatuses 87 B6. The Hyold Bone 87 56. Facial Portion of Skull 88 57. Base of Skull, External Surface 59 68. Sternum and Costal Cartilages 90 69. The Peculiar Ribs 91 60. The Vertebral (Column, or Spine 92 61. A Dorsal Vertebra 93 (^) LIST OF ILLUSTRATIONS. xi FIO, PAGE 62. A Cervical Vertebra 98 63. Seventh Cervical, or Vertebra Prominens 93 64. The Axis Si 65. The Atlas 91 66. Peculiar Dorsal Vertebrae 95 67. A Iiumbar Vertebra 95 68. The Sacrum, Anterior Surface 96 69. The Sacrum, Posterior Surface 96 70. The Left Clavicle, Upper Surface 97 71. The Left Clavicle, Under Surface 97 72. The Scapula, Anterior Surface 98 73. The Scapula, Posterior Surface 99 74. The Humerus, Anterior Surface 100 75. The Humerus, Posterior Surface 100 76. Ulna and Radius, Anterior Aspect 101 77. Radius and Ulna, Posterior Aspect 102 78. Carpus, Metacarpus, and Phalanges, Dorsal Surface 103 79. Carpus, Metacarpus, and Phalanges, Palmar Surface 104 80. Os Innominatum, Outer Surface 105 81.- Os Innominatum, Inner Surface 106 82. The Femur, Anterior Surface 107 83. The Femur, Posterior Surface 107 84. The Tibia and Fibula, Anterior Surface 108 85. The Tibia and Fibula, Posterior Surface 108 86. The Tarsus, Metatarsus, and Phalanges, Dorsal Surface 109 87. The Tarsus, Metatarsus, and Phalanges, Plantar Surface 110 88. The Thorax 117 89. The Pelvis 118 90. The Shoulder 121 91. The Elbow 123 92. The Knee 127 93. The Knee 127 94. Muscles of the Back 142 95. Muscles of the Abdomen 147 96. The Diaphragm 148 97. Muscles of the Shoulder and Arm 151 98. Triceps Muscles 151 99. Posterior Superficial Muscles of the Forearm 153 100. Muscles of the Anterior Femoral Region 159 101. Muscles of the Posterior Femoral Region 160 102. Anterior Muscles of the Leg 162 103. Posterior Muscles of the Leg 162 104. Right Side of Heart 171 106. Diagram of the Foetal Circulation 176 106. Arteries of the Face 181 107. Internal Maxillary Artery 183 108. The Arteries of the Head and Neck Facing 184 109. The Arteries of the Upper Extremity Facing 187 110. Abdominal Aorta 193 111. The Arteries of the Trunk Facing 194 112. The Arteries of the Lower Extremity Facing 196 113. Temporary Teeth 221 114. Permanent Teeth 222 115. Pharynx, Laid Open from Behind 225 116. Regions of the Abdomen 226 117. Abdominal Viscera. Facing 226 118. The Peritoneum 229 lis. The Liver 239 120. Transverse Section of Lobules of Liver 240 121. Vertical Section of Larynx 247 122. Heart and Lungs 254 123. Longitudinal Section of Kidney 258 124. Diagram of the Course of the Uriniferous Tubules 260 125. Male Organs of Generation 265 126. The Testicles and Epididymis 267 127. Vertical Section of Testicle 269 128. Internal Female Genitals 273 129. Female Organs of Generation 275 130. Section of an Ovary 276 131. External Female Genitals 278 132. Surface Anatomy of the Myelencephalon 288 133. Left Side of Human Cerebrum 291 134. Convolutions and Fissures of the Median and Tentorial Surfaces ol the Right Cerebral Hemisphere 292 Xll LIST OF ILLUSTRATIONS. PIG. PAGE 136. Inferior Aspect of Cerebral Hemisphere 293 136. The Lateral Ventricles and Choroid Plexus 297 137. Mesial Section of Brain and Brain Stem 300 138. Interior of Mesencephalon 305 139. Diagram of Sensory Tracts from Spinal Cord to Brain 306 140. Diagram of Motor Tracts from Brain to Spinal Cord 307 141. Scheme of the Nuclei and Root-fibres of the Cranial Nerves 308 142. Base of Brain and Cranial Nerves 310 143. Optic, Oculomotor, Trachlear, and Trifacial Nerves Facing 310 144. Facial and Auditory Nerves 314 145. Glossopharyngeal, Fneumogastric, Spinal Accessory, and Hypoglossal Nerves 315 146. Section of the Spinal Cord and Membranes 318 147. Cervical Plexus and Branches 319 148. Brachial Plexus and Branches 321 149. Lumbar Plexus and Branches Facing 324 150. Sacral Plexus and Branches Facing 326 151. The Sympathetic System Facing 335 152. The Olfactory Nerves and Nerves of Common Sensation to the Nose... 340 153. Nasal Meatuses and Accessory Cavities 341 154. Inner Wall of Orbit and Adjacent Parts 342 155. Horizontal Section of Right Orbit 343 106. Antero-posterior Section of Eyeball 344 157. Vessels of the Eye 347 158. The Structure of the Human Retina 350 159. Diagram of the Retinal Vessels 351 160. The Orbital Muscles 354 161. The Ophthalmic Artery and Branches 335 162. The Lachrymal Apparatus 358 163. External, Middle, and Internal Ear 360 164. The Auditory Ossicles 364 165. Organ of Corti 367 166. Triangles of the Neck 375 167. The Axilla 378 168. Dissection of the Inguinal Canal 380 169. Deep Femoral Region— The Femoral Vessels, etc. (McOrath) 383 170. Male Perineum, Superficial Dissection 385 171. Male Perineum, Deep Dissection 386 172. Female Perineum 388 INTRODUCTORY. The term "anatomy," derived from the Greek avd, "through," and refiveo), "to cut," signifies dissection, but has been appropriated under the general term Morphology, to the science which treats of the apparent properties of organized bodies. It is the science of organization and of form. Anatomy is divided into vegetable, M-hich treats of the structures and prop- erties of vegetal bodies ; comparative, which treats of the anatomy of animals ; human, which treats of tlie anatomy of man in a physiological or healthy state; pathological, which treats of the anatomy of the diseased human body, etc. Human anatomy is subdivided into general, descriptive, and surgical. General anatomy treats of the minute structure or physical elements of the organs of the body. It is also called micro- scopical anatomy, or histology. It comprises four distinct elementary tissues : — Epithelial, including mucous membranes, epidermis, glands, etc. Muscular, including striated and nonatriated muscles. Nervous, including the proper nerve elements. Connective, including white fibrous, elastic, adipose, cartilage, and bone. Descriptive or special anatomy treats of the various proper- ties of the organs arranged into systems; hence the name systematic. It includes : — Osteology, the anatomy of the bones. Syndesmology, the anatomy of the joints. Myology, the anatomy of the muscles. Angiology, the anatomy of the vessels. Neurology, the anatomy of the nerves. Splanchnology, the anatomy of the viscera, etc., etc. Surgical or topographical anatomy treats of the relative position of organs to one another and to the surrounding parts in special regions of the body, as the axilla, neck, or groin. (1) OSTEOLOGY. The frame-work of the body is composed of bones, which constitute the skeleton. This in the adult consists of 306 bones, including the ossieula auditus: Excluding the ear ossicles, the number is 200, or excluding also the two patellse and the hyoid bone the number is 197. Spine (including coccyx and sacrum) 26, cranium 8, face 14, ribs, sternum, and os hyoides 26, upper extremity 64, lower extremity 63, ear ossicles, 6. The bones of the head number 32, consisting of Eight cranial bones, the frontal, 2 parietal, occipital, 2 temporal, the sphenoid and ethmoid bones ; 14 facial bones, 2 lachrymal, 2 nasal, 2 malar, 2 superior maxUlae, 2 palate, 2 turbinal, 1 vomer, and 1 inferior maxilla. The bones of the trunk number 52, comprising Twenty-six vertebrae, 7 cervical, 12 dorsal, 5 lumbar, 1 sacrum and 1 coc.iyx. Twenty-six of the thorax, 7 pairs of true ribs, 3 pairs of false ribs, 2 pairs of floating ribs, the sternum and os hyoidas. The bones of the skull, face, thorax, and vertebral column are Imown as the axial skeleton, and consists of 74 segments. The bones of the upper extremity number 64, comprising on either side Two shoulder, clavicle and scapula ; 1 arm, humerus ; 2 forearm, radius and ulna ; 8 wrist (carpus), scaphoid, semi-lunar, cuneiform, pisi- form, trapezium, trapezoid, os magnum, and unciform ; 19 hand, 5 meta- carpal and 14 phalanges. The bones of the lower extremity number 62, comprising on either side One hip, divided into 3 portions, the ilium, ischium, and pubes ; 1 thigh, the femur; 2 leg, tibia and fibula ; 1 knee, the patella ; 26 foot, 7 tarsal bones, the astragalus, os calcis, scaphoid, cuboid, external, middle, and internal cuneiform ; 5 metatarsal bones, and 14 phalanges. The bones of the shoulder girdle, upper extermities, pelvic girdle, and lower extremities are known as the appendicular skeleton, and consists of 126 segments. They are divided into long, short, flat or tabular, and irregular. Long hones, such as femur, humerus, or radius, consist of shaft or diaphysis, and two extremities or epiphyses. The shaft or diaphysis is a cylindrical tube composed ex- ternally of compact tissue and internally of cancellous tissue, the centre being hollowed out by the medullary canal. The epiphyses, or extremities, are expanded for articulation, and are composed of a mass of solid cancellous tissue with a thin coating of compact tissue. (3) 4 HUMAN ANATOMY. The short bones, as tarsus or carpus, are small, compact, irregular cubes. Flat bones J as those of skull and shoulder-blade, afford extensive surfaces for protection or muscular attachment. Under irregular or mixed bones are classed the vertebral, sphenoid, maxillary bones, and such that could not be placed under either of the other heads. The surfaces of the bones are marked by certain eminences and depressions, which have received the following names : — An apophysis is a prominent excrescence formed directly upon a bone, and is distinguished from An epiphysis, which has been formed from a distinct centre and separated by cartilage, but afterward united to the bone. A head is a rounded, smooth, articular eminence for articulation. A condyle is an irregular prominence for muscular attachment. A trochanter is a large prominence for the attachment of rotator muscles. A tuberosity is a broad, uneven eminence. A tubercle is a small tuberosity. A spine is a sharp-pointed eminence. A line, or ridge, is a rough, narrow elevation, extending some distance. Others, from their fancied resemblance to ordinary objects, have received the following names : — Coronoid, like a crown ; coracoid, like a crow's beak ; unciform, or hamular, hook-like; malleolar, like a mallet; mastoid, nipple-like; zygoma, yoke-like; pterygoid, wing-like; odontoid, tooth-like; spinous, thorn-like; styloid, pen-like; rostrum, a beak; vaginal, enshea thing; squamous, scaly; conoid, cone-like. The cavities of bones are divided into the articular and the non-articular. The articular are named acetabulum, measure-like ; glenoid, hollow; cotyloid, cup-like; facet, smooth; trochlear, pulley- like; alveolar, socket-like. The non-articular cavities are named notches, fissures, grooves, furrows, fossse, hiatus, foramina, canals, sinuses, aque- ducts, cells, depressions, meatuses, etc. Composition of Bone. — Adult human bones have a specific gravity of 1.93, and are composed of about one-third (33.30) organic or animal matter, principally gelatin, and two-thirds inorganic or mineral matter, as follows :— Gelatin and blood-vessels, 33.30; phosphate, carbonate, and fluoride of calcium, 64.34; soda, sodium chloride, and mag- nesium phosphate, 2.36 : total, 100. Structure of Bone. — In structure they consist of an ivory- like compact substance, inclosing a lamellar, reticular cancellous substance. OSTEOLOGY. 5 They are covered with periosteum, and their cavities are Itaed with endosteum and filled with medulla, or marrow. Periosteum is a flbro-vascular membrane, composed of two layers, the outer formed chiefly of connective tissue, the inner composed of several layers of elastic fibres, the deepest of which is the "osteogenic," or bone-producing layer. The tendons and ligaments are attached to the periosteum by a mutual interlace- ment of the fibres. Endosteum^ medullary membrane or internal periosteum, is a delicate, highly vascular membrane lining the cylindrical cavity of long bones. Medulla, or bone-marrow, is of two kinds, the red in the flat and irregular bones, and the yellow in the shafts of adult long bones. The red marrow has few blood-vessels, but many corpuscular elements. These may be divided into three groups : 1, red nucleated trae "marrow-cells;" 2, large pinkish cells, "myelo-plaques" of Eobin, supposed to be the source of the red blood-corpuscles; and 3, giant cells , or "osteoclasts." The yellow marrow contains numerous blood-vessels, a few "marrow-cells," and a large quantity of fat which imparts to it its color. Blood-vessels of Bone. — The arteries consist of three sets: The arteries of the (1) compact and (2) cancellous tissue are derived chiefly from the periosteum, the latter being larger and less numerous. The medullary canal of long bones has (3) one or more nutrient arteries, which penetrate the compact tissue obliquely and divide into two branches, one passing upward, the other downward, in the canal. All the arteries anastomose freely with ?ach other. The veins consist of three sets also: 1. The veins of the compact substance are small and join the periosteum. 2. The veins of the cancellous tissue do not accompany the arteries, but in certain localities form large, tortuous channels, as in the diploic veins of the cranium. In the long bones they emerge at the ends near the articular surfaces. 3. The nutrient arteries are accompanied by one or more large veins, which emerge at the nutrient foramen. Lymphatics and nerves are numerous in the periosteum and are also found in the substance of the bone. Microscopical Appearance. — Microscopically, bone-tissue con- sists of concentric layers, or lamellw, arranged about the course of a vascular or Haversian canal ^/leooo i^ch in diameter. In and between these plates of bone-tissue are minute cavities, or lacunce, each containing a bone-cell, or "osteoblast," and from which diverge in every direction minute canals, or canaliculi. 6 HUMAN ANATOMY. connecting the lacuntE with each other and with the Haversian canals. Each canaliculus contains an artery, vein, and lym- phatic (SchafEer). In this manner every part of the osseous substance communicates. Development. — The development of bone is effected in two ways — (1) from cartilage, the intra-cartilaginous ; and (2) from membrane, the intra-membranous ; the former occurring at the base of the skull for the protection of vital centres, or in the extremities to secure rigidity of the parts. In the intra-cartilaginous the parts are first formed in cartilage and then converted into bone. The process is as fol- lows: The cartilage cells at the "centre of ossification" become enlarged and arranged in rows. The cartilaginous matrix also increases and separates the cells. Lime salts are deposited be- tween the rows of cells, inclosing them in oblong spaces called primary areolce. Blood-vessels from the deep or osteogenic layer of the periosteum, carrying numerous osteoblasts (bone-forming cells), and osteoclasts (bone-absorbing cells), pass into the area of calcified cartilage (centre of ossification). The osteoblasts re- place the calcified cartilage, which is absorbed, with new bone, which latter is absorbed in part (tunneled) by the osteoclasts to form the medullary spaces or marrow cavity. The walls of the spaces are gradually thickened by suc- cessive layers of osteoblasts, forming lamellae of bone, till noth- ing remains but a channel — ^the Haversian canal. The perichondrium having been in the meantime converted into periosteum, the osteogenic, or vascular, layer of this mem- brane furnishes a layer of osteoblasts that form enveloping, cir- cumferential layers of bone. Thus, from the osteoblasts all the structures of bone are produced; the remains of a group of cells constitute an Haver- sian canal, the layers deposited by them and the adjacent cells, the lamellse, and the isolated, persistent cells (osteoblasts) form the contents of the lacunse, or "true bone-cells," with their nu- trient canaliculi diverging from them. In the intra-mcmhranous form the parts are first formed in membrane, and from one or more centres of ossification lime salts are deposited in radiating spiculse, or osteogenic fibres, in- closing the osteoblasts. From these fibres the process spreads, and vessels from the neighboring parts pass into it and form Haversian canals. The formation of the lamelltE and the lacunee is essentially the same as in the intra-cartilaginous, and in most bones both processes go on simultaneously. OSTEOLOGY. 7 Ossification appears first in the clavicle and inferior maxilla (from fifth to seventh week) and last in the pisiform bone (twelfth year) . The epiphyses ossify from birth and unite from the age of puberty on to maturity, in reverse order to the* ap- pearance of ossification, being regulated, apparently, by the direc- tion of the nutrient artery in the upper extremity toward the elbow and in the lower extremity from the knee. The only exception to this rule is the lower end of the fibula, which ossifies and also becomes united before the upper {vide Table of Ossifi- cation) . In bones with only one epiphysis the artery is directed toward the other extremity. 1, frontal eminence; 2, superciliary ridge; 3, supra-orbltal arch; 4, external angular process; 6, internal angular process; 6, supra- orbital notch; 7, nasal eminence; 8. temporal ridge; 9, nasal spine. Growth of Bones. — Long bones grow in thickness from the periosteum and in length from the cartilage between the epiph- yses and the shaft near the extremities. The medullary cavity is at first solid and grows by absorption of the cancellous tissue by the giant cells, or osteoblasts. Elat bones grow in thickness from the periosteum and in surface from the borders. Short bones grow from the centre or centres of ossification in all directions. THE CRANIAL BONES. The cranial bones are eight in number : — 1 Frontal, 2 Parietal, 1 Occipital, 2 Temporal, 1 Sphenoid, 1 Ethmoid. 8 HUMAN ANATOMY. The frontal bone forms the anterior portion of the cranium and consists of two portions, a vertical or frontal por- tion and a horizontal or orbito-nasal portion. .The vertical portion consists of two surfaces, external and internal. The external surface is convex, and presents the frontal emi- nence on either side of the median line, the superciliary ridges, produced by the frontal sinuses, beneath, below which is the supra-orlital arch, forming the upper boundary of the orbit, and terminating on either side in the external angular process, articu- ? r^iyiu, FlQ. 2. 2, foramen csecum; 4, articulates with lesser wing; 6, articulates with ethmoid; 7, anterior and posterior ethmoidal foramina; 9, lachrymal fossa; 10, depression for pulley of superior oblique; 11, frontal sinus. lating with the malar bone, and the internal angular process, ar- ticulating with the lachrymal bone, and having at its inner third a notch or foramen, the supra-orbital foramen, for passage of supra-orbital artery, nerve, and vein. Ascending from the external angular process is the tem- poral ridge for attachment of temporal fascia. Between the internal angular processes is the nasal notch, for articulation with the nasal bone, and nasal process of superior maxilla, terminating below in the nasal spine, and above it projects the nasal eminence, or glabella, marking the location of the frontal sinuses. The posterior or internal surface is concave and lodges the anterior lobes of the brain. OSTEOLOGY. 9 In the median line is the frontal crest, which gives attach- ment to the falx cerebri and terminates below in a foramen, the foramen csecum, for passage of a small vein from the nose to the longitudinal sinus. The crest is grooved for longitudinal sinus, and has on either side of it depressions for the Pacchionian bodies. This surface is also grooved for anterior meningeal ar- teries. The horizontal portion consists of an external and internal surface. The external surface is concave, to form the roof of the orbit, and separated from the opposite side by the ethmoidal notch. I .,,,,»-^^%s» . iiilill:" », ■■■ / Fio. 3. 1, Superior border; 2, Inferior border for squamous portion of tem- poral; 3, anterior border for frontal bone; 4, posterior border for occipital; 5, parietal eminence; 6, parietal foramen; 7, anterior in- terior angle. To the outer side is a deep depression for the lachrymal gland, and to the inner side a slight one for the pulley of superior oblique. On the margin of the orbital plate are two notches con- verted into foramina by articulation with the ethmoid, the an- terior ethmoidal canal, for passage of nasal nerve and anterior ethmoidal vessels, and the posterior ethmoidal canal, for passage of posterior ethmoidal vessels. The internal surface is marked by the convolutions of the anterior lobes of the brain. Between the two plates of the vertical portion are the frontal sinuses, two irregular cavities lined with mucous membrane and opening into the middle meatus of the nose on either side by the infundibulum. It articulates with twelve bones, two parietal, sphenoid, ethmoid, nasal, superior maxillary, lachrymal, and malar. 10 HtlMAN ANATOMY. Its muscular attachments are three pairs, corrugator super- ciliij orbicularis palpebrarum, and temporal. It is developed from membrane by two ossific centres, one for each lateral half. Paeietals. — These are two quadrilateral bones forming the superior and lateral walls of the cranium. Each bone consists of two surfaces, four borders, and four angles. The external surface is convex and presents the parietal emi- nence a little above the middle, marking the original centre of ossification, the temporal ridge, crossing about the centre for at- tachment of the temporal fascia, and the parietal foramen, near the superior border or sagittal suture, for passage of a vein to the longitudinal sinus. *^ ma f* I 4. 'M--.M Fig. 4. 1, 2, 3, 4, superior, Inferior, anterior, and posterior borders; B, su- perior longitudinal sinus; 6, parietal foramen; 7, anterior inferior angle. The internal surface is concave, and presents furrows for the branches of the meningeal arteries, depressions for cerebral convolutions, and Pacchionian bodies; A half groove along the superior border for longitudinal sinus, and A groove near the posterior inferior angle for the lateral sinus. The superior border articulates with its fellow of the oppo- site side, forming the sagittal suture; The inferior border articulates from before backward with the sphenoid, squamous, and mastoid portions of temporal bones ; The anterior border forms with the frontal bone the coronal suture, and OSTEOLOGY. 11 The posterior harder forms with the occipital bone the lambdoid suture. Of the angles the anterior inferior is the only important one, being longer and grooved internally by the middle meningeal artery. It articulates with five bones — frontal, sphenoid, temporal, occipital, and opposite parietal bones. It has but one muscle attached — the temporal. It is developed from an ossific centre. The occipital forms the posterior inferior portion of the cranium. It presents two surfaces, four angles, and four bor- ders. 4 f: * ' TiBlTiifint^, X \ IB?''' IW \ FlQ. 5. 2, occipital protuberance; 3, crest; i, iDterlor curvefl line; 8, fora- men magnum; 6, condyle; 7, posterior condyloid foramen; 8, an- terior condyloid foramen; 9, jugular process; 11, basilar process; 12, tubercles tor check ligaments. The external and posterior surface is irregularly convex, and presents The occipital protuberance (also called external to dis- tinguish it), for insertion of ligamentum nuchas, descending from which to the foramen magnum is The occipital crest, for tendinous attachment, and diverg- ing on either side, above and below, are The superior curved line, for attachment of occipito fron- talis, trapezius, and other muscles; and The inferior curved line for attachment of rectus capitis posticus, major and minor. Anteriorly it presents a large aperture. 12 HUMAN ANATOMY. The foramen magnum, transmitting the medulla oblongata and its coverings, the vertebral arteries, and the spinal accessory nerves. A little in front and on either side of the foramen magnum are the condyles for articulation w^ith the atlas, the inner border of each condj'le presenting a tubercle ior the check ligament. Externally to each condyle are the jugular processes, afford- ing attachment for the rectus capitis lateralis, and forming by its articulation with the fibrous portion of the temporal bone the jugular foramen, or foramen lacerum posterius, transmitting the internal jugular vein, glosso-pharyngeal, pneumogastric, and u nc AM An fittir^/aett K Safa4tteaTia**.JI> ' Inftr.Artie.Pret Fig. 66. Peculiar dorsal vertebraa. St^en Aptie^ Prac. Fio. 67. A lumbar vertebra. 96 HUMAN ANATOMY. J'raTjujntory Fig. 68. The sacrum, anterior surface. Fig. 69. The sacrum, posterior surface. OSTEOLOGY. 97 ^i,.r„,.,i«i&f.'' S/^rTiat ExtrtmUij Fig. 70. The lett clavicle, upper surlaco. FIG. 71. The lett clavicle, under surface. 98 HUMAN ANATOMY. FlQ. 72. The scapula, anterior surface. OSTEOLOGY, 99 Cernema v''"'-* ^a:^*^. Fig. 73. The scapula, posterior surface. noo HUMAN ANATOMY. ';? ATreoAL ■uiumB utvii FIC3. 74. The humerus, anterior surface. Fig. 75. The humerus, posterior surface. OSTEOLOGY. 101 R M.0 t It a IttOII DICITORUH- tUOLIMIB ^rlcxan Stylaui jfrsceat Pig. 76, Ulna and radius, anterior aspect. HETMAWI rsLMCia IHTHNsaiirOUlCIf 102 HUMAN ANATOMY. FLCxon DiotTOBS ^UT. CARPI K D, CJtT.CAftPt ItAD.BKKVIi 1BT. BKOUKDI INTSRNODU NtLlDt^ fir tXt. CARPI IJLNAB. Fig. 77. Radius and ulna, posterior aspect. OSTEOLOGY. 103 'l^Jtuir Fig. 78. Carpus, metacarpus, and phalanges, dorsal surface. 104 HUMAN ANATOMY. fi.niDR.cAIIPl ULnAitia M.EXOR BUEVIS minimi DICITI fLEXen OSSIS MBTftCARPI MINIMI DICITI Metaear/iu* PiQ. 79. Carpus, metacarpus, and phalanges, palmar surface. OSTEOLOGY. 105 aueujis Mvcjuai Fig. 80. Os innominatum, outer surface. 106 HUMAN ANATOMY. Fig. St. Os innominatum, inner surface. OSTEOLOGY. 107 Via. 82. Tbe temur, anterior surface. Fio. 83. Tbe lemur, posterior surlace. 108 HUMAN ANATOMY. ^e aa Bl^M>l^ XKUraal M«It,Btuw Fio. 84. The tibia and fibula, anterior surface. '^3jlTKUt Fig. 85. The tibia and fibula, posterior surface. OSTEOLOGY. 109 ruxBi UHen rouicii Ifetataratefi .IRCVIl OICITonul* iiy«# tXT. lANDHS rOLblGIB Fig. 86. The tarsus, metatarsus, and phalanges, dorsal surface. 110 HUMAN ANATOMY, riaijiui tHTiouM Fig. 87. Thn tarsus, metatarsus, and phalanges, plantar surface. ARTICULATIONS AND LIGAMENTS. The bones of the skeleton are connected together by articu- lations or joints. These consist essentially of the expanded extremities of bones, covered with cartilage, often separated by interarticular fibro-cartilage, held together by ligaments, and lined by synovial membrane. The bone entering into the articular lamella differs from ordinary bone by its extreme density, w^ithout Haversian canals, its lacunae being much larger, and without canaliculi. It is not perforated by blood-vessels. Cartilage is a whitish, highly elastic, non-vascular structure, forming in the foetus the greater part of the skeleton, and found in the adult chiefly in the joints, the walls of the thorax, and certain orifices, as the nostrils, ears, etc. It is either temporary, becoming ossified later, or permanent, remaining unossifled. The latter is divided into three varieties : articular, in joints covering the ends of the bones ; costal, form- ing part of the thorax; and reticular, arranged in plates or lamellae to maintain the shape of parts. Fibro-cartilage consists of a mixture of cartilaginous with white fibrous tissue. There are four varieties : — (a) Interarticular, interposed between the joint surfaces; (i) Connecting, binding bones together as in pubes; (c) Circumferential, deepening cavities, as glenoid cavity of shoulder; and (dj Stratiform, lining grooves for tendons. Synovial membranes are of three kinds: articular, lining the cavities of movable joints throughout except the surface of the cartilage ; bursal, irregular cavities interposed at convenient posi- tions to alleviate friction; from their contents they may be either mucous or sjmovial ; and vaginal synovial membranes, or sheaths, surrounding tendons and diminishing friction. Synovia is a transparent, viscid liquid, albuminous in :ts nature. The articulations consist of three (3) classes: diarthrosis, movable; synarthrosis, immovable; and amphiarthrosis, mixed. (Ill) 112 HUMAN ANATOMY. 1. The diarthrosis are subdivided into: — Arthrodia, gliding-joint, as superior tibio-fibular ; Enarthrosis, ball-and-socket joint, as shoulder and hipj Ginglymus, hinge-joint, as knee and ankle; Trochoides, a ring surrounding a pivot, as atlo-axoid joint, and superior radio-ulnar. Condyloid, elliptical cavity receiving an ovoid head, as wrist-joint. Reciprocal reception, a concavo-convex articulation, as carpo-meta- carpal joint of thumb. 2. Synarthrosis^ surface, immovably connected by fibrous membrane without synovial membrane. They are divided into: — Sutura, bones interlocking with one another; Schindylesis, a fissure in one bone receiving a plate of bones, as between vomer and sphenoid; Gomphosis, a socket with a pivot inserted, as in alveolar cavities for teeth. Synchondrosis, a temporary joint in which the connecting medium is cartilage. The sutura may be either true, sutura vera, or false, sutura notha, the former having three divisions: dentata, tooth-like processes, as interparietal suture; serrata, saw-like edges, as iu- terf rental suture; limbosa, dentated processes and beveled mar- gins, as f ronto-parietal ; the latter two divisions: — Squamosa, overlapping beveled margins, and Harmonia, by union of roughened surfaces, as intermaxillary suture. 3. AmpMarthrosis, bony surfaces connected by fibro-carti- lage, with or without synovial membrane, as between vertebral bodies, and pubic symphysis. Motions in Joints. — Joints admit of four (4) distinct varieties of motion : — (a) Gliding movement, between contiguous surfaces; (■ftj Angular movement, as flexion, extension, adduction, and ab- duction ; (c) Circumduction, as in true enarthrosis; (d) Rotation on its own axis, as between the atlas and axis. Temporo-maxillart is formed by condyle of lower jaw, below articulating with glenoid cavity of the temporal bone and eminentia articularis above. It is a double arthrodial joint. Ligaments. — External Lateral. — Origin, tubercle on outer edge of zygoma; insertion, outer side of neck of condyle. Internal Lateral. — Origin, spinous process of sphenoid; in- sertion, lower circumference of inferior dental foramen. ARTICULATIONS AND LIGAMENTS. lib Capsular. — Origin, circumference of articulation ; insertion, neck of condyle. Interarticular fibro-cartilage is within cavity; external pterygoid muscle attached to its inner edge. Synovial membrane is divided into two by cartilage. Intermaxillary ligament passes from external pterygoid process to corouoid. Stylo-maxillary Ligament. — Origin, styloid process; inser- tion, inner surface of angle. of jaw. Nerves are derived from the auriculo-temporal and masseteric branches of the inferior maxil- lary. Articulations of Vertebral Column. — These are formed between the contiguous surfaces of the vertebral bodies and ar- ticular processes, inclosed lq capsular ligaments, and connected with the following ligaments : — The anterior common ligament extends from the front of the body of the axis down the anterior surface of the spine to the sacrum, being expanded opposite, and attached to each inter-' vertebral connecting fibro-cartilage; The posterior common ligament descends along posterior surfaces of bodies from axis to sacrum within spinal canal. It also expands opposite and adheres to intervertebral substance ; Intervertebral substance, disks of varying size, twenty-seven in number, composed of decussating fibres of fibro-cartilage, ar- ranged in crescentie laminae with central, semi-pulpy substance, separate the vertebrae ; Ligamenta subflava descend in pairs, twenty-three on either side, from one lamina to another,«from axis to sacrum, inclosing spinal canal; Supraspinous ligament, strong cord descending from one vertebra to another, from vertebra prominens (seventh cervical) to sacrum. Its continuation in cervical region forms ligamentum nuchce; Interspinous stretch between spinous processes throughout dorsal and lumbar regions ; Intertransverse, between transverse processes in lower dorsal and lumbar regions. The arteries are derived from ascending cervical and verte- bral in cervical, iatercostals in dorsal, and lumbars in lumbar regions. The nerves are from spinal nerves in each region. Atlo-axoid articulation consists of four joints, the two lateral joints being arthrodia, and that between the odontoid process and arch of atlas and transverse ligament — a diarthrosis 114 HUMAN ANATOMY. rotatoria — ^being double, one in front, atlo-odontoid, and one be- hind, odonto-transverse. The ligaments are: — Anterior atlo-axoid, two — superficial and deep, connecting anterior borders together; Posterior atlo-axoid, connecting the arches of the atlas and axis posteriorly, and pierced laterally by the second spinal nerve ; Capsular, two, connecting the articulating processes later- ally, and lined with synovial membrane; Transverse, or cruciform, arises from the tubercle on one side of the lateral mass, passes across the back of odontoid proc- ess to be inserted into the opposite tubercle. The arteries are from vertebral, nerves from second cervical, or from loop between it and suboccipital. OcciPiTO-ATLOiD. — Anterior occipito-atloid — two, super- ficial and deep, connecting the tubercle and upper border of an- terior arch of atlas with basilar process and margin of foramen magnum of occiput. Posterior occipito-atloid (membranous), connects the upper border of posterior arch of atlas with posterior margin of fora- men magnum. Lateral ligaments, two bands connecting transverse process of atlas with jugular process of occipital. Capsular ligaments connect the articular processes of atlas with condyles of occiput. ' The arteries are from vertebral; nerves, from suboccipital. OcciPiTO-AxoiD. — Occipito-axoid is a broad band, the con- tinuation of posterior common ligament, extending from the body of the axis to basilar grooye of occipital. Odontoid, OT check, pass upward and outward from the sum- mit of the odontoid process, to be inserted into the inner side of the occipital condyles. Ligamentum suspensonum, or central occipito-odontoid, arises from the centre of the transverse ligament at the apex of the odontoid process, and passes upward, to be inserted into the anterior margin of the foramen magnum. Occipito-cervical, or cervico-hasilaris, arises from the bodies of the third cervical and axis, and is inserted into the basilar groove of occipital. The arteries are from vertebral; the nerves from suboc- cipital. Capsular ligaments. — Throughout the spine the articula- tions between the facets of the articular processes are surrounded by capsular ligaments, making twenty-five pairs in all. CosTO-VEKTEBRAL ARTICULATIONS. — The costo-vertcbral have each a double arthrodia between the head of the rib and the ARTICULATIONS AND LIGAMENTS. 115 bodies of the two adjacent vertebrse, except the first, tenth, eleventh, and twelfth ribs, which have each but a single joint. Anterior Gosto-vertehral, or Stellate Ligament. — Origin, head of the rib ; insertion, into body of vertebra above and below, and interarticular cartilage between. Interarticular costo-vertehral is within the capsular liga- ment, from the crest on the head of the rib to the intervertebral substance. Capsular ligament surrounds the articular surfaces. Gosto-transverse articulations are united by three ligaments — anterior, middle, and posterior cos to-transverse ligaments — extending between the tubercles of the ribs to the transverse proc- ess below. Arteries, the intercostals ; nerves, anterior branches of spinal. CosTO-STEENAL ARTICULATIONS. — These are connected by arthrodia excepting the first, which is a synarthrodia. Anterior chondro-sternal ligaments connect the chondral and sternal surface in front; Posterior chondro-sternal connect them behind. Capsular ligaments surround all, and synovial membranes are present in all but the first, while the second has an inter- articular cartilage interposed. The chondro-xiphoid ligament connects the cartilage of the seventh, and sometimes the sixth rib with the xiphoid appendix. Interchondral and Costo-chondral Articulations. — The ribs are connected with their cartilages by the periosteum covering them. The cartilages of the sixth, seventh, and eighth ribs, and sometimes the fifth and ninth, are connected by anterior and posterior inter-chondral ligaments, held together by capsular ligaments and lined with synovial membrane. Ligaments of Sternum. — The first and second portions of the sternum are articulated by an amphiarthrodial joint, some- times a diarthrodial joiat, connected by the anterior intersterrml and posterior intersternal ligaments^ and lined with synovial membrane. The third portion, or ensiform cartilage, is united to the gladiolus by a synarthrodial joint. TiiE thorax. The thorax is a conical osteo-cartilaginous framework, formed by the dorsal vertebree behind, the ribs, intercostal mus- cles, and costal cartilages laterally, and the sternum in front. The apex, or superior opening, is bounded behind by the first dorsal vertebra, laterally by the first rib, and in front by the upper border of the sternum. 116 HUMAN ANATOMY. It transmits the following structures: — CEsophagus, trachea, thoracic duct (on left side), lymphatic ves- sels of right side of chest and of surface of liver, innominate artery (right side), left common carotid artery, left subclavian artery, right and left superior intercostal arteries, right and left internal mammary arteries, thyroidea ima artery (if present), right and left inferior thy- roid veins, right and left innominate veins, right and left phrenic nerves, right and left pneumogastric nerves and their cardiac branches, left recurrent laryngeal nerve, right and left first dorsal nerves, right and left sympathetic nerves and their cardiac branches, apices of lungs and pleurae, remains of thymus, and deep cervical fascia passing to peri- cardium. The longus colli, sterno-hyoides, and sterno-thyroides on both sides also pass through it. The inferior opening is formed by the last dorsal vertebra behind, the last rib laterally, and the ensiform cartilage in front. It is filled in by the diaphragm. The cavity of the thorax contains the following structures : the heart and great vessels, trachea, bronchi, and lungs, azygos and bronchial veins, internal mammary arteries, pneumogastric, phrenic, and splanchnic nerves, thoracic duct, cESophagus, lym- phatic vessels, and glands {vide mediastinum). Spine and Pelvis. — Sacra-vertebral articulation. In addi- tion to those of the spine, given above, there are two ligaments connecting the last lumbar vertebra with the sacrum: — - Lumbosacral ligament, arising from the transverse process of the last lumbar vertebra, and inserted into the base of the sacrum ; Ilio-lumbar ligament, arising from the apex of the trans- verse process of the last lumbar vertebra, and inserted into the crest of the ilium. The arteries are from last lumbar, ilio- lumbar, and lateral sacral. The nerves are branches of fourth and fifth lumbar and sympathetic. Sacro-iliac articulation, between the articular surfaces of the sacrum and ilium, is an amphiarthrodial joint connected by The anterior sacro-iliac ligament, crossing between anterior surfaces of the two bones; Posterior sacro-iliac ligament, the stronger, passes in dif- ferent directions between the posterior surfaces of the two bones, firmly binding them together. One of these, stronger than the others, passes from the posterior superior spinous process to the third sacral transverse process. It is called The oblique sacro-iliac ligament. The arteries are from gluteal, sacral, spinal, and ilio-lum- bar; the nerves from lumbo-sacral and posterior sacral cords. ARTICULATIONS AND LIGAMENTS. 117 Sacro-iscMatic Articulation. — The os innominatum and sa- crum are united by two important ligaments — ^the great sacro- sciatic (posterior) ligament, and the lesser sacro-sciatic (or an- terior) ligament. The great sacro-sciatic ligament arises from the fourth and fifth transverse tubercles of the sacrum, from spine of ilium, sacrum, and coccyx, and passes downward, outward, and forward, to be inserted into the inner margin of tlie tuberosity of the ischium, being prolonged forward as the falciform ligament protecting the internal pndic veins and nerves. FIQ. 88. 1, manubrium; 2, gladiolus; 3, xiphoid appendix; 4. first dorsal vertebra; 6, last dorsal vertebra; 6, first rib. It converts the sacro-sciatic notch into the lesser sacro- sciatic foramen, transmitting the obturator internus muscle and nerve, the internal pudic vessels and nerves. The lesser sacro-sciatic ligament arises from the lateral margin of the sacrum and coccyx, and is inserted into the spine of the ischium. It converts the sacro-sciatic notch inio the greater sacro-sciatic foramen, transmitting the pyriformis mus- cle, gluteal vessels, superior gluteal nerve, sciatic vessels and nerves, and the internal pudic vessels and nerves. Sacro-coccygeal articulation is an amphiarthrodial joint, resembling the vertebral, having an interarticular £bro-cartilage, and connected together by 118 HUMAN ANATOMY. An anterior sacro-coccygeal ligament, and A posterior sacro-coccygeal ligament, continuations respect- ively of the anterior and posterior common ligaments. Lateral sacro-coccygeal, intertransverse^ and intercornual ligaments connecting the rudimentary transverse processes and the cornua together. The arteries are from lateral and median sacral; The nerve branches of coccygeal, fifth sacral, and posterior divisions of fourth sacral. Pubic Aeticulation. — This is an amphiarthrodial joint, known also as the symphysis pubis. It is connected by the Anterior pubic ligament in front ; _^s-yL Fig. 89. A A, antero-posterior diameter; B B, transverse diameter; C C, oblique diameter; 1, sacro-iliac ligament; 2, anterior or lesser sacro- Bciatic ligament; 3, posterior or great sacro-sciatic ligament. Superior pubic ligament above; Posterior pubic ligament behind ; Subpubic ligament below, forming the boundary of the pubic arch. Within the joint an interposed fibro-cartilage separates the bones. The obturator membrane, or ligament, closes the obturator foramen, and affords attachment for the external and internal obturator muscles and allows passage at its upper outer part, for the obturator vessels and nerves. THE PELVIS. The pelvis is a strong, bony basin, formed at the lower end of the vertebral column by the articulation of the sacrum and coccyx posteriorly with the ossa innominata laterally. ARTICULATIONS AND LIGAMENTS. 119 It is divided into a false and true pelvis by tlie ilio-pectineal line. The false pelvis is situated above the ilio-pectineal line, and corresponds to the expanded iliac fossae, being bounded laterally by the ossa ilii, and having in front and behind wide intervals. The true pelvis includes all that portion below the linea ilio- pectinea, and presents a cavity, superior circumference or inlet, and an inferior circumference or outlet. The inlet is heart-shaped, being bounded behind by the promontory of the sacrum, laterally by the linea ilio-pectinea, and in front by the crest and spine of the pubes. Its principal diameters in the female are: — Anteroposterior, from symphysis to vertebral angle, four and three-fourths inches; transverse, across the widest part, five and one- fourth inches; and oblique, from the ilio-pectineal eminence in front to the sacro-iliac symphysis behind, five inches. In the male, these meas- urements are about three-fouiths of an inch less in every diameter. It transmits the following structures : — Viscera and Ducts. 1. Small intestine. 5. Urachus and obliterated hypo- 2. Rectum. gastric arteries. 3. Vasa deferentia (in male), 6. Uterus (pregnant). round ligaments (in female). 7. Bladder (distended). 4. Ureters. 8. Peritoneal coverings. 1. Superior hsemorrhoidal. 6. Obturator (from deep epigas- 2. Internal iliac. trie). 3. Sacra media. 7. Ovarian. 4. Ilio-lumbar. 8. Pubic branches from external 5. Branches of vasa intestini ten- epigastric aiteries. uis. - 9. Lymphatic vessels. Nerves. 1. Sympathetic. 3. Obturator. 2. Branches from hypogastric 4. Lumbosacral cord. plexus. The co.vity of the pelvis is bounded behind by the sacrum and coccyx, in front by the symphysis, and laterally by the body of the ischium, forming a curved canal, widest in the middle. It contains, in the recent state, the organs of generation, rectum, bladder, and their vessels, nerves, ducts, etc. Its axis corre- sponds to the concavity of the sacrum and coccyx. Its depth in front, at the symphysis, is about one and a half m the middle axial line, three and a half behind, in males four and a half to possibly five and a half. 120 HUMAN ANATOMY. The outlet of the pelvis is bounded. by three prominences, the tuber ischii laterally and the coccyx behind; separated by three notches, the subpubic arch in front and the sacro-sciatic notches posteriorly. Its principal diameters are : transverse, three and a half in male, four and three-fourths in female; antero-posterior and oblique, three and one-fourth in male, five in female. In the erect position the pelvis is placed at an angle of from sixty degrees to sixty-five degrees with the ground. Differences of Sex. — The male pelvis is characterized by the strength of the bones, by prominent muscular impressions, by the depth and narrowness of the cavity, the large obturator foramen, and the acute angle of the subpubic arch. The female pelvis is characterized by the lightness of the bones, the slight muscular impressions, by the shallow expanded iliac fossae, the increased size of the cavity in every diameter, and the obtuse angle of the subpubic arch. ARTICULATIONS OF THE UPPER EXTREMITY. The sterno-clavicular articulation is an arthrodial joint, formed between the sternal end of the clavicle and the sternum and first rib, being divided into two cavities by an interartieular fibro-cartilage. Its ligaments are: — Anterior sterno-clavicular^ is a broad band passing between the articular margins, attached to the interartieular cartilage and two synovial membranes, and forming the front part of the capsular ligament; Posterior sterno-clavicular, passes over the posterior aspect of the joint and corresponds to the former; Interclavicular, passes from the sternal end of the' clavicle on one side across the supra-sternal notch to the clavicle of the opposite side; Gosto-clavicular, or rhomioid, passes from the rhomboid impression on the inferior surface of the clavicle to the upper sur- face of the first costal cartilage. The synovial membranes form two sacs separated by the in- terartieular cartilage. The arteries are from muscular branch of supra-scapular and contiguous muscular branches; The nerves, from descendens noni. AcEOMio-CLAvicuLAR ARTICULATION is an arthroidal joint formed between the acromial extremity of the clavicle and the acromial process of the scapula. Its ligaments are: — ARTICULATIONS AND LIGAMENTS. 121 The superior acromio-clavicvlar, connect the adjoining parts of the superior surfaces of the claTicle and acromion process; Inferior acromio-clavicular, connecting the inferior sur- faces ; Goraco-clavicular, consists of two parts : — Trapezoid, broad, square, arises from upper surface of cora- coid process, and is inserted into oblique line on under surface of clavicle; and Conoid, behind and internal, arises from base of coracoid process, and is inserted into conoid tubercle on under side of clavicle and a line internal to it. Fig. 90. 1, superior acromio-clavicular ligaments; 2, coraco-clavicular liga- ments; 3, coraco-acromial ligaments; 4, transverse ligament; 6, capsular ligament; 6, coraco-bumeral ligament; 7, tendon of biceps. The capsular ligament covers the anterior and posterior sur- faces of the joint, uniting above and below with the superior and inferior acromio-clavicular ligaments. The arteries are from anterior circumflex, acromial thoracic, and supra-scapular. The nerves are anterior circumflex and supra-scapular. The scapula has, also. The coraco-acromial, arising from the outer border of the coracoid process and inserted into the summit of the acromion, and Transverse, a ligamentous band bridging the supra-scapular notch, converting it into a foramen for supra-scapular nerve. 122 HUMAN ANATOMY. The shoulder-joint is an enarthrodial joint, formed above by the glenoid cavity of the scapula and below by the head of the humerus. Its ligaments are — glenoid, coraco-humcral, and capsular. The glenoid surrounds the edge, deepens the glenoid cavity, and is continuous above with the long head of the biceps tendon. The capsular ligament, extensive and loose, arises above it from circumference of glenoid cavity behind the ligament, is attached below to the anatomical neck of humerus, and is pierced by tendons of two or three muscles. The coraco-humeralj or accessory, is a fibrous band which extends obliquely downward and outward from the coracoid proc- ess to the anterior part of great tuberosity, strengthening the capsular ligament. The synovial membrane is extensive and reflected upon the tendons of biceps, infra-spinatus, and sub-seapularis muscles. Its arteries are derived from the subscapular, supra-scapular, dorsalis scapulas, anterior and posterior circumflex ; Its nerves, from the subscapular, supra-scapular, and cir- cumflex. The elbow-joint is a ginglymoid articulation formed above by the lower extremity of humerus, below by upper ex- tremities of ulna and radius. Its ligaments are external and internal lateral, anterior and posterior ligaments. External lateral arises from external condyle of humerus and is inserted into outer margin of ulna. Internal lateral, much stronger, consists of two portions; anterior arises from fore part of internal condyle to be inserted into coronoid process, and posterior from back part of condyle to inner margin of olec- ranon. Anterior ligament arises above coronoid fossa, and is in- serted into coronoid process of ulna and orbicular ligament. Posterior ligament, attached above olecranon fossa, and be- low to olecranon process of ulna. The anterior and posterior ligaments become continuous with the lateral to encircle the joint. The arteries are derived from superior and inferior pro- funda, anastomotica magna, posterior interosseous recurrent, an- terior and posterior ulnar recurrent, and radial recurrent. The nerves, from median, ulnar, musculo-cutaneous, and musculo-spiral. The eadio-ulnar articulations are three — the superior, middle, and inferior. The superior radio-ulnar is a diarthrosis rotatoria, formed by the inner side of the head of radius and lesser sigmoid cavity ARTICULATIONS AND LIGAMENTS. 123 of ulna. Its synovial cavity comnmnicates with the elbow-joint and it is held together by the orbicular, or annular ligament, at- tached in front and behind to the lesser sigmoid cavity and sur- rounding the head of radius. Its arteries and nerves are the same as fqr elbow. The middle radio-ulnar articulation consists of the oblique or round ligament, passing from tubercle of ulna to below tuberosity of radius, and the interosseous passing between the bones, deficient above for passage of posterior interosseous ves- FiG 91. 1, anterior ligament; 2 and 2, anterior and posterior portions of Internal lateral; 3, orbicular ligament; 4. oblique; 5. interosseous membrane. sels, and perforated below for anterior vessels. Its arteries are from anterior interosseous; its nerves, from anterior and pos- terior interosseous. Inferior radio-ulnar consists of anterior and posterior radio- ulnar ligaments, passing from one to the other in front and back of wrist, and triangular interarticular fibro-cartilage, interposed between head of ulna and cuneiform bone, and binding the radius and ulna firmly together.. Its arteries are from anterior and posterior interosseous and carpal; its nerves, from median and posterior interosseous. 124 HUMAN ANATOMY. The radio- carpal, or wrist-Joint, is a condyloid joint con- sisting of an elliptical cavity formed by the radius and triangular cartilage, into which fits the convex surfaces of the scaphoid, semi-lunar, and cuneiform below. Its ligaments are external and internal lateral, anterior and posterior radio-carpal. External lateral passes from the styloid process of radius to the scaphoid, os magnum, and trapezium. Internal lateral passes from the styloid process of uhia to the cuneiform, pisiform, and annular ligament. The anterior radio-carpal is a broad membrane attached above to the anterior margia of the radius, its styloid process and ulna, and below to cuneiform, semi-lunar, scaphoid, and os mag- num, strengthened by a band extending from styloid process of ulna to semi-lunar and cuneiform. The posterior radio-carpal is attached to dorsal margin of radius, and below to cuneiform, semi-lunar, and scaphoid. The anterior and posterior radio-carpal belong to tlie annular liga- ment (mrfe fascias). Its arteries are the anterior and posterior carpal, derived from the radial and ulnar, anterior and posterior interosseous, and ascending branches from the deep palmar arch. Its nerves are derived from the median, ulnar, and posterior interosseous. The carpal articulations are arthrodial, and consist of three sets. The first row is held together by two palmar, two dorsal, and two interosseous; the second row by three dorsal, three palmar, and three interosseous; and the two rows are held to- gether by a palmar, dorsal, internal and external lateral ligament. First Eow. — The palmar and dorsal ligaments connect the scaphoid and semi-lunar, and semi-lunar and cuneiform together on the front and back. The interosseovs connect the semi-lunar with the scaphoid and cuneiform. The pisiform bone has a separate capsular ligament and an interosseous ligament connecting it to the fifth metacarpal bone and cuneiform. Second Eow. — The palmar and dorsal ligaments connect the trapezium with the trapezoid, and the os magnum with the trapezoid and unciform, on the front and back of the wrist. The interosseous connect the adjacent surfaces of the trape- zium and trapezoid, and the os magnum with the trapezoid and unciform. Two Rows Together. — The palmar and dorsal connect the bones of the first with the second rows on the front and back AETTCULATIONS AND LIGAMENTS. 125 of the wrist. The internal lateral passes on the ulnar side be- tween the cuneiform and unciform. The external lateral, stronger, passes on the radial side between the scaphoid and trapezium. The anterior annular ligament contributes strength to these joints. The arteries are: Carpal of anterior interosseous, anterior and posterior carpals of radial and ulnar, carpal of deep palmar arch, interosseous recurrent, and terminal branches of anterior and posterior interosseous. The nerves are from median, ukiar, and posterior inter- osseous. The carpo-metacarpal articulations are each arthrodial joints, held together by dorsal, palmar, interosseous, and capsular ligaments, except the thumb, which has only capsular. The capsular ligament of the thumb surrounds the margins of the articular surfaces of the first metacarpal and trapezium, and is lined by a separate synovial sac. The palmar and dorsal carpo-metacarpal connects the carpus and metacarpus on the palmar and dorsal surface. The interosseous connect the adjoining inferior angles of OS magnum and imciform with the contiguous surfaces, fourth and fifth metacarpal bones. The synovial membranes of the wrist consist of five distinct sacs : First, memirana sacciformis lines lower end of ulna, sig- moid cavity of radius, and upper surface of triangular cartilage; second, lower surface of radius and cartilage and upper surface of first row of carpus; third, between margins of carpus and carpo-metacarpal joints ; fourth, between carpo-metacarpal joint of thumb ; and fifth, between pisiform and cuneiform. Intermetacarpal Articulation. — The bases of the metacarpal bones, except the thumb, articulate with one another by ar- throdial joints, lined by synovial membrane continuous with the carpal sac and connected together by palmar, dorsal, and inter- osseous ligaments. The digital extremities are connected by a transverse ligament across the anterior surface, continuous with anterior metacarpo-phalangeal ligament. The metacarpo-phalangeal and phalangeal are of the condyloid variety, like the wrist-joint, allowing of motion in every direction except laterally. They are each connected by one anterior and two lateral ligaments, the posterior ligament being substituted by the extensor tendon, which crosses the dorsum of the joint. The arteries and nerves are from the digitals. 126 HUMAN ANATOMY. AETICULATIONS OF THE LOWER EXTREMITY. The hip-joint is a true enarthrodial articulation, formed above by deep cup-like cavity of acetabulum, below by promi- nent spherical head of femur. Its ligaments are the capsular, ilio-femoral, teres, cotyloid, and transverse. The capsular ligament, dense and strong, is attached above to margin of acetabulum and cotyloid ligament, and below to the spiral line in front and to the neck behind. It is strength- ened by several accessory bands, the pubo-femoral, ilio-trochan- teric, ischio-capsular, and ilio-femoral. Of these the latter is the most important, arising above from anterior inferior spine, and attached below to spiral line, and has received the name of Y-ligament of Bigelow. The ligamentum teres, or round ligament, is a strong tri- angular band, its base arising from the bottom of acetabulum and margins of cotyloid notch externally, its apex inserted below and behind centre of head of femur. The cotyloid ligament encircles and deepens the acetabulum, and at the inner side, under the name of the transverse ligament, it bridges over the cotyloid notch, converting it into a foramen for the passage of nutrient vessels to the joint. There are numerous bursa about the joint, with one of which, beneath the ilio-psoas muscle, the synovial membrane often communicates. Its arteries are derived from the sciatic, internal and ex- ternal circumflex, obturator, and gluteal. Its nerves are derived from the great sciatic, obturator, accessory obturator, and the sacral plexus. The knee-joint is a ginglymoid articulation, formed above by condyles of femur, below by head of tibia, and in front by patella. Its ligaments are divided into two sets, the external, con- sisting of anterior, posterior, internal lateral, two external lat- erals, and capsular; and internal, consisting of anterior, pos- terior, two semi-lunar fibro-cartilages, transverse, coronary, ligamentum mucosura, and ligamenta alaria. Its joint surface is the most extensive in the body. External Set. — Anterior, or ligamentum patella, is the tendinous portion of extensors of thigh between patella and tubercle of tibia. The posterior, or ligamentum posticum Wins- lowii, derived principally from the tendon of semi-membranosus, arises from inner tuberosity of tibia, passes upward and outward to be inserted into external condyle of femur. ARTICULATIONS AND LIGAMENTS. 127 The internal lateral is a flat and broad ligament arising from back part of inner condyle of femnr, descends forward to be inserted into inner tuberosity of tibia, covering in its course the inferior internal articular artery and nerve, part of tendon of semi-membranosus, and attaching itself to the internal semi- lunar fibro-cartilage. The long external lateral ligament, round and strong, arises from tuberosity on outer part of external condyle, and descends forward to the head of fibula. The short external lateral liga- ment is an accessory, not very constant, band, descending pos- teriorly to the preceding. Fig. 93. 2, anterior crucial ligament; 3, 2, quadriceps extensor: 3, patella: posterior crucial ligament; 4, trans- 4, ligamentum patelto; 6, bursa; 7, verse ligament; 6, 7, internal and ex- 8, 9, synovial membrane of iinee-joint: ternal semi-lunar cartilages; 8, liga- 10, anterior crucial ligament, menta patellse; 9, bursa patellae ; 10. anterior superior tibio-flbular liga- ment. The capsular ligament fills up the intervals between the preceding ligaments, and is strengthened by bands from fascia lata, vasti, crureus, semi-membranosus, biceps, and sartorius tendons. Internal Set. — Crucial ligaments are two strong, short, interosseous, crossing each other from before backward. The anterior, or external, arises from inner posterior part of condyle of femur, and descends forward and inward to be inserted into front of spine of tibia and internal semi-lunar cartilage. The posterior, or internal, arises from outer fore part of inner condyle of femur, descends downward, backward, and out- ward to spine of tibia. 128 HUMAN ANATOMY. The semi-lunar fibro-cartilages consist of two crescentic laKiinse of interarticular cartilage, resting upon the upper ar- ticular surface of tibia, which serve to deepen its surface. The internal is attached by its inner border to internal lateral ligament and to head of tibia by coronary ligaments, its extremities attached in front of anterior crucial ligament and behind the spine. The external, more circular, is connected to edge of tibial head by coronary ligaments, its extremities being inserted behind and in front of the tibial spine. In other words, the ends of the semi-lunar cartilages are all, except the anterior end of internal, attached to the tibia between the crucial ligaments. A band of fibres passing from the anterior margin of ex- ternal cartilage to the internal has received the name of tram- verse ligament. The knee is lined by the most extensive synovial membrane in the body, covering both surfaces throughout and extending up between quadriceps tendon and surface of femur. Below the patella in front is a duplicature of synovial mem- brane, inclosing some adipose tissue, which has received the name of ligamentum mucosum, and extending from it are two fringes — the ligamenta alaria. On either side in the popliteal space behind are bursse, which often communicate with joint. There are also bursse over the patellse, above and beneath the ligamentum patellee, and between the inner hamstring and head of tibia. The arteries of the knee-joint are derived from the anasto- motica magna, the articular branches of the popliteal (five), and the recurrent branch of the anterior tibial. Its nerves are derived from the anterior crural, obturator, external and internal popliteal nerves. The tibio-fibulae articulations are three— superior, middle, and inferior. The superior is an arthrodial joint between fibular head and outer tuberosity of tibia, and consists of an- terior and posterior ligaments stretching on either side of the outer tuberosity, downward and backward, to the head of the fibula. The arteries are some of the knee; the nerves, from external popliteal. The middle consists of an interosseous membrane between the bones, connecting them firmly together, deficient above, for passage forward of anterior tibial artery and below for anterior peroneal vessels. The inferior is an arthrodial joint between contiguous in- ARTICULATIONS AND LIGAMENTS. 129 ferior surfaces of tibia and fibula, and consists of four: the interosseous, continuous with that above; the anterior inferior tihio-fihular, extending downward and outward between tibia and fibula in front; the posterior inferior tibio-fibular, occupying a similar position behind; and the transverse, a long narrow band, passing between external malleolus and tibia. The syno- vial membrane is continuous with that of ankle-joint. The arteries and nerves are the same as those to the ankle. The AiTKLE-JOiNT is a ginglymoid articulation, formed be- tween the lower extremity and malleolus of tibia on the inner side, the malleolus of the fibula on the outer side, and the as- tragalus below. Its ligaments are anterior, posterior, external, and internal lateral. The anterior tilio-tarsal ligament consists of a broad set of irregular fibres, attached above to lower margin of tibia, below to astragalus. The posterior tibio-tarsal ligament passes trans- versely between back part of tibia and astragalus. The internal lateral, or deltoid, has two layers — ^the super- ficial triangular, its apex arising from malleolus, its base spread out from before backward to be attached to scaphoid and infe- rior calcaneo-scaphoid ligament, the sustentaculum, and inner side of astragalus; and the deep, a short, round cord passing from internal malleolus to astragalus. The external lateral ligament consists of three distinct fasciculi, anterior, middle, and posterior, arising from near sum- mit of external malleolus, and being inserted respectively into the front of astragalus, the outer surface of os calcis, and the back of astragalus. The arteries of the ankle-joint are derived from the malleo- lar branches of the peroneal and anterior tibial. Its nerves are branches of the anterior and posterior tibial nerves. The articulations of the taksal bones consist of those of the first row, those of the second row, and of the two rows with each other. First Eow. — The astragalus articulates with the calcaneum by two distinct surfaces separated by the interosseous groove, and firmly connected by the external and posterior calcaneo-astraga- loid ligaments, and the interosseus. The external calcaneo- astragaloid ligament passes vertically downward from the outer surface of astragalus to outer surface of calcaneum. The pos- terior is a short, narrow, oblique band connecting the posterior border of astragalus to the os calcis. The interosseus is a strong, short ligament passing vertically and obliquely downward between the two bones, the principal bond of union. 9 130 HUMAN ANATOMY. The arteries of the first row are tarsal from dorsalis pedis, external malleolar froni anterior tibial, and branches of peroneal from posterior tibial. Nerves, from posterior tibial, or plantar. The second row are firmly held together by dorsal, plantar, and interosseous ligaments. The two former unite the dorsal and plantar surfaces and the interosseous, four strong bands connect the scaphoid and cuboid, the internal and middle, the middle and external cuneiforms, and the external cuneiform and cuboid. The arteries of the second row are from plantars and meta- tarsals. Nerves, from anterior tibial, and internal and external divisions of posterior tibial nerves. The two eovsts of the taesus are connected with each other by three sets of ligaments, first between os calcis and cuboid, consisting of superior and internal calcaneo-cuboid, long and short calcaneo-cuboid; second, between os calcis and sca- phoid, consisting of superior and inferior calcaneo-scaphoid ; and third, between astragalus and scaphoid, consisting of a single superior astragalo-scapTioid ligament, a band passing from neck of astragalus to upper surface of scaphoid. Calcaneo-cuboid. — The superior passes between dorsal sur- faces of OS calcis and cuboid. The internal, passing from os calcis to inner side of cuboid, is one of the main bonds of union between the two rows of tarsus. The inferior firmly connects the bones below, the ligamen- tum longcB plantce passing from os calcis forward to the plantar surface of the cuboid, and sending fibres forward to bases of second, third, and fourth metatarsal bones; the ligamentum breve plantce passing from fore and under surface of os calcis to under surface of cuboid. The two preceding are important in preserving the arch of the foot. The superior and inferior calcaneo-scaphoid pass above and below these bones, firmly connecting them. The arteries of the mediotarsal joints are derived from the anterior tibial, tarsal, metatarsal, or external plantar. Nerves, from external branch of anterior tibial. The takso-metataesal articulations are firmly held to- gether by dorsal and plantar ligaments, and connected with one another by interosseous bands, three in number, passing between internal cuneiform and second metatarsal bone, external cunei- form and second metatarsal, and external cuneiform and third metatarsal. The arteries are from metatarsal, dorsalis pedis, and deep plantar arch; ARTICULATIONS AND LIGAMENTS. 131 The nerves, from anterior tibial, and internal and external plantars. Intermetatarsal Articulations. — The bases of all except the first are connected by dorsal plantar and interosseous ligaments, and the distal extremities are connected by the transverse meta- tarsal ligament. The synovial membeanes concerned in these articulations are six: (1) between calcaneo-astragaloid articulation, posterior to interosseous membrane; (2) anterior calcaneo-astragaloid and astragalo-scaphoid ; (3) ealcaneo-cuboid; (4) between mid- dle and external cuneiform, scaphoid, and three cuneiforms, middle and external cuneiform, and second and third metatarsal, and between cuboid and external cuneiform and scaphoid; (5) between internal cuneiform and first metatarsal; and (6) be- tween cuboid and fourth and fifth metatarsal bones. The metataeso-phalakgeal articulation's are each con- nected by two lateral and a plantar ligament, and the phalangeal articulations are the same, the extensor tendons acting as dorsal ligaments. They resemble in every respect the articulations in the hand {vide hand) . THE MUSCULAR SYSTEM. Ti-iB muscles constitute 45 per cent, of the bodj'-weight and are the active agents of locomotion. They consist of two kinds — the non-striated or involuntary, and the stbiated, or voluntary. The latter, being usually under the control of the will (the heart being a notable exception), have also received the name of voluntary, and the former, not under the control of the will, involuntary. Microscopically, their structure is as follows : — N on-striated muscular tissue is made up of elongated, con- tractile, nucleated fibre-cells. They consist of minute fibres in bundles, inclosed in extremely delicate sheaths of connective tissue. They are held together by an albuminous cement and col- lected by connective tissue, into groups or masses, one-tenth to one five-hundredth of an inch in length. Non-striated muscular tissue is highly vascular and is sup- plied with nerves from the sympathetic. Striated muscular tissue is made up of spindle-shaped fibres of from one and a half to two inches in length and one two- hundredth to one six-hundredth of an inch in diameter. They are held together in bundles by the endomysium, a delicate fibro- connective tissue. These bundles are collected into groups by the perimysium, a stronger connective-tissue band, forming the fasciculi of the fully formed muscle. Striated muscular tissue is also highly vascular. The parts of an individual fibre are — first, the sarcolemma, a very delicate, transparent, elastic sheath; second, the mem- branes of Krausc, which appear as dark lines stretching across the fibre at regular intervals, forming the third, or the compart- ments of Krause, which contain the muscular substance. The latter contain the multi-nucleated muscle-cells. The striated muscular fibres of the heart differ somewhat from the preceding, being branched, and dividing and subdivid- ing longitudinally to form an intricate net-work. The muscles are connected to cartilages, ligaments, bones, and skin, either directly or by aponeuroses or tendons. Aponeuroses are dense, white, fibrous membranes, serving to connect the muscles with the structures to be acted upon. Tendons are white, fibrous, glistening cords. They are com- posed of white, fibrous tissue, arranged into bauds or bundles. (132) THE MUSCULAR SYSTEM. 133 They are attached to the perichondrium, periosteum, ligaments, and subcutaneous tissue by a mutual interlacement of fibres. Classification. — Muscles are classified according to region of the body, their physiological action, or from their embryo- logical development. The former arrangement is retained here for practical purposes. Names of Muscles. — They have received their names : — 1. From the arrangement of their fibres they have received the names of radiated, penniform, bipenniform, etc. 2. From their uses they are called extensors, adductors, etc. 3. From their direction, oblique, rectus, transversalis, etc. 4. From the number of insertions, as triceps, biceps, etc. 5. From their form, as rhomboid, deltoid, etc. 6. From their attachment, as occipito-frontalis, sterno-hyoid, etc. The origin refers to the fixed extremity; the insertion, to the movable point. Muscles of the Head. cranial eegion. Occipito-frontalis. — Origin, by two bellies, one from outer tv7o-thirds of superior curved line of the occipital bone and base of the mastoid process of the temporal, the other from corrugator supercilii, orbicularis palpebrarum, and pyramidalis nasi fibres; insertion, into fibrous aponeurosis covering the ver- tex of the skull ; action, raises the eyebrows and used chiefly as a muscle of facial expression ; nerves, supraorbital, small occipital, facial, and posterior auricular branch of facial. AURICULAE REGION. Attollens Aurem. — Origin, from aponeurosis of occipito- frontalis ; insertion, into the surface of the pinna ; action, raises the ear ; nerve, occipitalis minor. Atteahens Aurem. — Origin, from edge of occipito-fron- talis aponeurosis ; insertion, into anterior part of helix ; action, draws the ear upward and forward ; nerve, facial. Eetrahens AuREii. — Origin, from mastoid portion of tem- ' poral bone ; insertion, into lower part .of concha ; action, draws the ear backward ; nerve, posterior auricular branch of facial. palpebral region. Orbicularis Palpebrarum. — Origin, from nasal process of superior maxilla, internal angular process of frontal, and from front and sides of tendo-palpebrarum ; insertion, into skin of the cheek, eyelids, forehead, and temple, blending with the cor- rugated supercilii and the occipito-frontalis ; action, sphincter of the eyelids ; nerve, facial. 134 HUMAN ANATOMY. Tbndo Palpebeakum (Tendo Oculi). — Origin, nasal proc- ess of superior maxilla; insertion^ inner part "of tarsal cartilage. CoKEUGATOE SuPEEOiLii. — Origin, superciliary ridge; in- sertion, into orbicularis about the middle of the orbital arch; action, draws eyebrows inward and downward and wrinkles the forehead; nerve, facial. Tensor Taesi. — Origin, from crest and orbital surface of lachrymal gland ; insertion, into tarsal cartilages near the puncta lachrymalia; action, draws the lachrymal canals inward and against the globe of the eye; nerve, facial. ORBITAL REGION. Levator Palpebr^ Supeeioeis. — Origin, from lesser wing of the sphenoid, near the optic foramen ; insertion, superior bor- der of tarsal cartilage; action, elevates the upper eyelid; nerve, third cranial, or motor oculi. Eectds Superior. — Origin, sheath of the optic nerve and upper margin of optic foramen; insertion, into upper surface of sclerotic coat ; acitow, rotates the eyeball upward ; nert)e, third cranial. Eegtus Inferior. — Origin, from lower and inferior part of optic foramen (ligament of Zinn) ; insertion, into lower sur- face of sclerotic; action, rotates the eyeball downward; nerve, third cranial. Eegtus Inteenus. — Origin, same as rectus inferior; inser- tion, into inner surface of sclerotic; action, rotates the eyeball inward; nerve, third cranial. Eectus Externus. — Origin, by two heads — lower from liga- ment of Zinn and lower margin of sphenoidal fissure, upper from outer margin of optic foramen; insertion, into outer surface of sclerotic; nerve, abducens, or sixth cranial. Passing between the two heads are the ophthalmic vein, the third, nasal branch of fifth, and sixth nerves. Superior Oblique. — Origin, from inner margin of optic foramen: its tendon passes through a pulley near the internal angular process of the frontal bone; insertion, into sclerotic, between external and superior recti, midway between entrance of optic nerve and the cornea; action, rotates the eyeball on its axis; nerve, fourth, or patheticus. Inferior Oblique. — Origin, orbital plate of superior maxilla; insertion, near that of superior oblique, between ex- ternal and superior recti; action, rotates the eyeball on its axis; nerve, third cranial. THE MUSCULAR SYSTEM. 135 NASAL EEGION. Pteamidalis Xasi ok Peoceeus. — Origin, from the occipi- to-f I'ontalis ; insertion, into the compressor nasi-; action, lowers the inner angle of the eyebrows; nerve, facial. Levatoe Labii Supeeioeis Al^qdb Nasi. — Origin, nasal process of superior maxilla; insertion, the ala of the nose and upper lip, blending with the levator labii oris proprius and or- bicularis; action, dilates the nostril and elevates the upper lip; nerve, facial. DiLATOK Naeis Posteeioe.- — Origin, nasal notch of superior maxilla; insertion, into skin at the margin of the nostril. Dilator Naris Anteeioe. — Origin, from cartilage of the ala ; insertion, into the skin of nose ; action, dilates the nostrils ; nerve, facial. CoMPEESSOE Nasi. — Origin, superior maxilla, near, the in- cisive fossa ; insertion, into fibro-cartilage of the nose, continuous with the pyramidalis nasi aponeurosis and its fellow; action, dilates the nostril ; nerve, facial. CoMPEESSOR Narium Minor. — Origin, from alar cartilage; insertion, into the skin of the end of the nose ; action, dilates the nostril; nerve, facial. Depressor Alje Nasi. — Origin, incisive fossa of superior maxilla ; insertion, into the ala of the nose ; action, contracts the nostril ; nerve, facial. maxillary REGION. Levator Labii Supeeioeis (Proprius). — Origin, above in- fraorbital foramen to malar and superior maxilla ; insertion, into the upper lip ; action, elevates the upper lip ; nerve, facial. Levator Anguli Oris. — Origin, from canine fossa; inser- tion, into the angle of the mouth, blending the depressor anguli oris, orbicularis, and zygomatici; action, draws the angle inward and raises it; nerve, facial. Ztgomaticus MxsoiL.— Origin, from malar bone ; insertion, into angle of mouth, blending with depressor anguli oris and orbicularis. Ztgomaticus Minor. — Origin, from malar bone ; insertion, at the angle of the mouth, blending with the levator superioris; action, draws the lip outward and upward; nerve, facial. Levator Labii Infeeioeis. — Origin, from incisive fossa of lower jaw; insertion, into the skin of the chin; action, raises the lower lip; nerve, facial. 136 HUMAN ANATOMY. Depressor Labii Inferioris. — Origin, from external ob- lique line of lower jaw; insertion, into skin of lower lip, blending with its fellow and the orbicularis; action, lowers the angle of the mouth ; nerve, facial. Depressor Anguli Oris. — Origin, external oblique line of lower jaw; insertion, into the angle of the mouth, continuous with the orbicularis and risorius at its insertion and the platysma at its origin; action, depresses the angle of the mouth; nerve, facial. Orbicularis Oris. — Origin, by accessory fibres (accessorii orbicularis superioris and inferioris and naso-labialis), from superior and inferior maxillary borders and nasal septum ; inser- tion, into the buccinator and adjoining muscles, forming the sphincter of the mouth ; action, closes the lips ; nerve, facial. Buccinator. — Origin, from pterygo-maxillary ligament and the posterior alveolar processes of the upper and lower jaw; insertion, into orbicularis oris; action, compresses and con- tracts tiie cheeks; nerves, facial and buccal branch of the in- ferior maxillary nerve. Risorius (Santoeini). — Origin, from fascia of masseter muscle , insertion, at angle of the mouth ; action, draws back the angles of the mouth, — ^the "smiling" muscle; nerve, facial. Masseter: Superficial Portion. — Origin, inner surface of zygoma and malar process of superior maxilla; insertion, into the ramus and angle of the lower jaw. Deep Portion. — Origin, posterior border and inner surface of the zygoma ; insertion, into the ramus and coronoid process of the jaw; action, raises the lower jaw, and the superficial portion assists in drawing it forward ; nerve, inferior maxillary. Temporal. — Origin, from the temporal fascia and the tem- poral fossa ; insertion, into the coronoid process of the lower jaw ; action, raises and draws backward the lower jaw ; nerve, inferior maxillary. Internal Pterygoid. — Origin, from pterygoid fossa and the tuberosity of the palate-bone; insertion, into the inner side of the ramus and angle of the lower jaw; action, draws forward and raises the lower jaw; the accessory triturating muscle of mastication ; nerve, inferior maxillary. External Pterygoid. — Origin, by two heads — the lower from the tuberosities of the palate and superior maxilla and from the external pterygoid plate, the upper from the pterygoid ridge on the greater wing of the sphenoid ; insertion, into the front of the neck of the lower jaw and inner side of interarticular car- tilage; action, draws the jaw forward; triturating muscle of mastication; nerve, inferior maxillary. THE MUSCULAR SYSTEM. 137 Muscles of the Eak {vide Ear). Muscles of the Neck. SUPEEFICIAL cervical REGION. Plattsma Mtoides. — Origin, from the deep fascia over the trapezius, deltoid, and pectoral muscles, and from the clavicle and acromion ; insertion, into the lower jaw and skin of the face ; action, wrinkles the skin of the neck and protects the air-passages and blood-vessels from external pressure, — rudimentary in man; nerves, branch of superficial cervical plexus and the facial. Sterno-cleido-mastoid. — Origin, by two heads — sternal portion from the fore and upper part of the manubrium sterni, the clavicular portion from the inner third of the upper border of the clavicle, leaving a triangular interval; insertion, into the mastoid process and outer two-thirds of the superior curved line of the occipital bone; action, rotates and depresses the head; nerves, spinal accessory and branches of the cervical plexus. INFRA-HTOID REGION. Stbrno-htoid. — Origin, from upper and back part of the manubrium sterni and inner extremity of the clavicle ; insertion, into the lower border of the os hyoides ; action, 'depresses the hyoid bone; nerve, branch from the loop of communication be- tween the descendens and communicans noni. Sterno-thykoid. — Origin, posterior surface of manubrium sterni; insertion, into oblique line of the surface of the thyroid cartilage; action, depresses the larynx; nerve, branch from the communicating loop above. Thtro-htoid. — Origin, from the oblique line on the sur- face of the thyroid cartilage; insertion, into the greater cornu and body of the hyoid bone; action, elevates the larynx; nerve, hypoglossal. Omo-htoid. — Origin, from the upper border of the scapula and the transverse ligament; insertion, into the lower border of the OS hyoides; its centre is tendinous and bound down to the cartilage of the first rib by a process of the deep cervical fascia; action, depresses the hyoid bone and draws it backward; nerve, from the communicating loop above. SUPRA-HTOID region. Digastric. — Origin, by two bellies — posterior from digas- tric groove of mastoid process of the temporal bone, anterior 138 HUMAN ANATOMY. from a depression in tlie lower border of the ]"aw near the sym- physis; the tendon is held to the hyoid bone by an aponeurotic loop and pierces the stylo-hyoid; action, raises the tongue and hyoid bone ; nerve, mylo-hyoid branch of the inferior dental and facial. Stylo-hyoid. — Origin, from outer surface of styloid proc- ess ; insertion, into the body of the hyoid bone ; action, retracts and elevates the hj'oid bone ; nerve, facial : near its insertion it is perforated by the tendon of the digastric. MYLO-HYOiD.^Ori^in, from the mylo-hyoid ridge of the inferior maxilla from last molar to symphysis ; insertion, into a fibrous raphe in the median line, extending from the hyoid bone to the chin and into the body of the hyoid bone; action, draws forward and elevates the hyoid bone and forms part of the floor of the mouth; nerve, mylo-hyoid branch of the inferior dental. Genio-hyoid. — Origin, from inferior genial tubercle of the internal surface of the symphysis of the jaw; insertion, into the front of the body of the hyoid bone; action, same as the mylo- hyoid; nerve, hypoglossal. MUSCLES of the TONGUE. Gbnio-hyoglossus. — Origin, from superior genial tubercle of the interiTal surface of the symphysis of the jaw; insertion, by fan-like expansion into the whole length of the inferior sur- face of the tongue, the side of the pharynx, and the body of the hyoid bone; action, retracts and protrudes the tongue; nerve, the hypoglossal. Hypoglossus. — Origin, body and greater comu of the hyoid bone; insertion, between the lingualis and styloglossus into the side of the tongue; action, renders the tongue convex from side to side ; nerve, hypoglossal. Styloglossus. — Origin, from stylomaxillary ligament and styloid process of the temporal; insertion, into the side of the tongue, blending with the hyoglossus and lingualis; action, re- tracts and elevates the tongue; nerve, hypoglossal. Lingualis. — Consists of four portions — superficial, inferior, transverse, and vertical. It lies between the genio-hyoglossus and the hyoglossus, extending from the apex to the base of the tongue; action, renders the tongue convex from before back- ward ; nerve, the chorda tympani. palato-glossus (constrictor isthmi faucium) (vide pala- tal region). THE MUSCULAR SYSTEM. 139 PHARYNGEAL AND PALATAL EEGION. CoNSTEiCTOE SUPERIOR. — Origin, from margin of internal pterygoid plate and its hamular process ; from pterygo-maxillary ligament, part of the alveolar process of the lower jaw and side of the tongue, tendon of the tensor palati, and part of the palate- hone ; insertion, into the median raphe and the pharyngeal spine of the basilar process of the occipital bone ; action, constricts the pharynx; nerves, pharyngeal plexus and glosso-pharyngeal. Constrictor Medius. — Origin, from the stylo-hyoid liga- ment, greater and lesser cornna of the hyoid bone; insertion, into the median fibrous raphe, blending with its fellow of oppo- site side; action, constricts the pharynx; nerves, pharyngeal plexus and glosso-pharyngeal. Constrictor Inferior. — Origin, from the sides of the thy- roid and cricoid cartilages; insertion, into the fibrous raphe of the pharynx; action, contracts the pharyngeal canal; nerves, external laryngeal, glosso-pharyngeal, pharyngeal plexus. Stylo-phartngeus. — Origin, from base of the styloid proc- ess; insertion, into the constrictor muscles, palato-pharyngeus, and posterior border of the thyroid cartilage ; nerves, pharyngeal plexus and glosso-pharyngeal. Levator Palati. — Origin, cartilaginous portion of Eusta- chian tube and apex of the petrous portion of the temporal bone; insertion, into the back part of the soft palate, blending with its fellow of the opposite side; action, elevates the soft palate ; nerve, descending palatine from Meckel's ganglion, from the facial. Tensor Palati, or Cieoumelexus. — Origin, from spine of the sphenoid, vaginal portion of temporal bone, cartilage of Eustachian tube, and scaphoid fossa at base of internal pterygoid plate ; insertion, into the palate-bone and the soft palate ; action, renders tense the palate ; nerve, a branch from the otic ganglion. AzYGOS UvuL^, OR LEVATOR. — Origin, from aponeurosis of soft palate and posterior nasal spine; insertion, into the uvula; action, raises the palate; nerves, descending palatine branches and Meckel's ganglion, from the facial. Palato-glossus (Constrictor Isthmi Fauoium). — Ori- gin, from soft palate on either side of the uvula ; insertion, into the dorsum and side of the tongue, blending with the fibres of the styloglossus — this muscle forms the anterior pillar of the fauces ; action, constricts the fauces; nerves, palatine branches of Meck- el's ganglion. 140 HUMAN ANATOMY. Palato-phartngeus. — Origin, by two portions, from soft palate; insertion, into posterior border of thyroid cartilage and pharynx; this muscle forms the posterior pillar of the fauces; action, closes the posterior nares ; nerves, palatine branches from Meckel's ganglion. VERTEBRAL REGION. Eeotus Capitis Anticus Major. — Origin, by four tendons from transverse processes of the third, fourth, fifth, and sixth cervical vertebrse; insertion, basilar process of occipital bone; action, flexes the head; nerves, suboccipital and deep internal branches of cervical plexus. Rectus Capitis Anticus Minor. — Origin, from root of transverse process and, anterior part of the lateral mass of the atlas; insertion, basilar process of occipital, behind the former; action, flexes the head ; nerves, suboccipital and deep branches of the cervical plexus. Eeotus Lateralis. — Origin, superior surface of the trans- verse process of the atlas ; insertion, inferior surface of the jugu- lar process of the occipital; action, draws the head laterally; nerves, suboccipital and deep internal branches of the cervical plexus. Longus Colli. — Origin, from three portions, superior ob- lique portion from anterior tubercles of the transverse processes of the third, fourth, and fifth cervical; insertion, tubercle on the anterior arch of atlas; inferior oblique portion, origin, from anterior surface of the bodies of the first two or three dorsal ver- tebra; insertion, anterior tubercles of the transverse processes of the fifth and sixth cervical; vertical portion, origin, from the anterior surface of the bodies of the lower three cervical and upper three dorsal bodies of the second, third, and fourth cervical vertebrae; action, rotates and flexes the cervical portion of the vertebrse; nerves, anterior branches of the lower cervical nerve. Scalenus Anticus. — Origin, from the tubercle of the first rib ; insertion, into the anterior tubercles of the transverse proc- esses of the third, fourth, fifth, and sixth cervical vertebra; action, flexes and rotates the vertebral column; nerves, anterior branches of the lower cervical nerve. Scalenus Medius. — Origin, upper surface of the first rib, behind the groove for the subclavian artery ; insertion, into the transverse processes of the lower six cervical vertebrae; action, rotates and flexes the vertebral column; nerves, the anterior branches of the lower cervical nerve : the posterior thoracic, long thoracic nerve, or external respiratory nerve of Bell, has its THE MUSCULAR SYSTEM. 141 origin in the substance of this muscle, by union of two roots from fifth and sixth cervical nerves. Scalenus Posticus. — Origin, from the outer surface of the second rib, behind the serratus magnus ; insertion, into the pos- terior tubercles of the transverse processes of the lower two or three cervical vertebrse; action, flexes and rotates the spine; nerves, anterior branches of the lower cervical nerves. Of these muscles the scalenus medius is the longest and largest, and the scalenus posticus the smallest. Muscles of the Larynx and Epiglottis {vide Larynx) . Muscles oe the Trunk. muscles op the back. First Layer. Trapezius. — Origin, from inner third of superior curved line of the occipital bone, the ligamentum nuehfe, the spinous processes of the seventh cervical, and all the dorsal vertebrae; insertion, into the outer third of the posterior border of the clavicle, the inner margin of the acromion process, and the crest of the spine of the scapula ; action, with head fixed elevates point of shoulder, with shoulders fixed together draws head backward, or singly draws head to corresponding side; adducts, rotates inward, and depresses humerus ; nerves, cervical plexus and spinal accessory. Ligamentum NucHiE. — Origin, from external occipital protuberance ; insertion, spinous processes of, the cervical verte- brse, from the second to the seventh. This ligament is rudi- mental in man, and in the lower animals sustains the head. Latissimus Doesi. — Origin, by an aponeurosis from the spinous processes of the six lower dorsal, the lumbar and sacral vertebrse, the supraspinou.s ligament, the crest of the ilium, and the three or four lower ribs ; insertion, into the inner lip of the bicipital groove of the humerus, in front of the teres major, and a little above the pectoralis major; the tendon of this muscle twists completely on itself, so that the superior fibres become the inferior; action, draws the arm backward and downward, or, fixing the arm, raises the lower ribs and draws the trunk forward ; nerve, subscapular. Second Layer. Levator Anguli Scapulje. — Origin, from three to five tendons from the posterior tubercles of the transverse processes of the three or five upper cervical vertebrae ; insertion, into pos- terior border of the scapula, at the root of the spine; action. 142 HUMAN ANATOMY. elevates the angle of the scapula; nerve, interior division of the third and fourth cervical nerves. Ehomboideus Mtso^.— Origin, from spinous processes of seventh cervical and first dorsal vertebrae and the- ligamentum nuchffi ; insertion, into root of the spine of the scapula ; action, draws the inferior angle upward and backward; nerves, branches from the fifth cervical nerve. FiQ. 94. Muscles of the back: 1, trapezius; 2, 4, latissimus dorsl; 10, leva- tor anguli scapulsB; 11, rhomboideus minor; 12, rbomboideus major; 13, 14, splenius capitis et colli; 16, vertebral aponeurosis; 16, serratus posticus inferior. Ehomboideus Major. — Origin, from supraspinous liga- ment and spinous processes of four or five upper dorsal vertebrse ; insertion, by a tendinous arch attached above near the spine and below to the inferior angle of the scapula; action, draws the THE MUSCULAR SYSTEM. 143 inferior angle upward and backward; nerves ^ branches of the fifth cervical. Third Layer. Seeratus Posticus SuPEEiOR.-^Ong'iw, from spinous proc- esses of the two or three upper dorsal and last cervical vertebras, and from the ligamentum nuchse; insertion, into upper borders of the second to the fifth ribs inclusive ; action, assists in respira- tion; nerves, external branches of the posterior division of the cervical. Seeeatus Posticus Infeeioe. — Origin, from spinous proc- esses and interspinous ligaments of two or three upper lumbar and two lower dorsal vertebra; insertion, into lower borders of the four lower ribs, external to their angles ; action, elevates the ribs; assists in respiration; nerves, external branches of the posterior divisions of the lower dorsal. Splenius. — Origin, from spinous processes of last cervical and six upper dorsal vertebrae, the lower half of the ligamentum nuchse, and the supraspinous ligament ; insertion, by two heads — splenius capitis into the occipital bone, ]ust below the superior curved line and the mastoid process of the temporal bone ; sple- nius colli into posterior tubercles of the transverse processes of the three or four upper cervical vertebrae; action, separately, rotates the head and draws it to the other side; together, draw the head backward; nerves, external branches of the posterior divisions of the cervical. Fourth Layer. Ekectoe Spin^. — Origin, from sacro-iliac groove and from a broad tendon attached internally to the spinous processes of the lumbar and two or three lower dorsal vertebrae and supra- spinous ligament; externally, the crest of the ilium and the posterior part of the sacrum; insertion, by two parts — 1, sacro- lumbalis (ilio-costalis), inserted into bhe angles of the six or seven lower ribs; this muscle has two accessory portions, (a) musculus aecessorius ad sacro-lumbalem : origin, from angle of six lower ribs ; insertion, angles of six upper ribs ; (h) cervicalis ascendens : origin, angles of the four or five upper ribs ; inser- tion, into the posterior tubercles of the fourth to the sixth cer- vical vertebrae, inclusive; 2, longissimus dorsi; insertion, into the transverse processes of all the dorsal vertebrae, and from the seventh to the eleventh ribs, inclusive, between their angles and tubercles ; action, bends the trunk backward and erects the spine ; nerves, external branches of the posterior divisions of the lumbar and dorsal. 144 HUMAN ANATOMY. Thansversalis Colli (or Ceevicis). — Origin, transverse processes of six upper dorsal vertebra; insertion, into tlie pos- terior tubercules of the transverse processes of the second to the sixth cervical vertebrse inclusive; nerves, external branches of the posterior divisions of the cervical. Trachelo-mastoid. — Origin, from articular processes of three or four lower cervical, and from the transverse processes of the third to the sixth dorsal vertebrse; insertion, into the pos- terior margin of the mastoid process, below the sterno-mastoid and the splenius; action, steadies the head; nerves, external branches of the posterior divisions of the cervical. Spinalis Dorsi. — Origin, spinous processes of last two dor- sal and first two lumbar vertebrae ; insertion, into spinous proc- esses of the dorsal vertebrse, blending with the semi-spinalis dorsi ; action, erects the spinal column ; nerves, external branches of the posterior divisions of the cervical. Spinalis Ceevicis. — Origin, from the spinous processes of the fifth to the seventh cervical vertebrse; insertion, into the spinous process of the axis; action, steadies the neck; nerves, same as above; this muscle is absent in 20 per cent, of the subjects. CoirPLEXus OR Semispixalis Capitis. — Origin, by seven tendons from transverse processes of the upper three dorsal and seventh cervical and articular processes of the fourth, fifth, and sixth cervical; insertion, .into the occipital bone, between the curved lines ; action, separately, rotates and draws the head to one side ; together, draw the head directly backward ; nerves, sub- occipital, great occipital, and internal branches of the posterior divisions of the cervical. Fifth Layer. Sejii-spinales Dorsi. — Origin, from transverse processes from the fifth to eleventh dorsal vertebrse; insertion, into the spinous processes of the lower two cervical and upper four dorsal vertebrse; action, erects the spinal column; nerves, internal branches of the posterior divisions of the cervical. Semi-spinales Ceevicis. — Origin, from transverse processes of lower four cervical and upper four dorsal vertebrse ; insertion , into spinous processes of the second to the fifth cervical verte- brse; action, erects the spinal column; nerves, same as above. MuLTTFiDUS SPIN.E. — Origin, from the transverse processes of the dorsal region, the articular processes in the cervical and lumbar region, the posterior superior of the ilium, posterior sacro-iliac ligaments, and from the aponeurotic arch of the erec- tor spinas; insertion, each fasciculus is attached to the lamina' THE MUSCXJLAR SYSTEM. 145 and spinous process of the vertebrae above; action, preserves the erect condition of the spine; nerves, internal branches of the posterior divisions of the cervical, dorsal, lumbar, and sacral nerves. EoTATOHES Spin^. — Eleven on either side. Origin, from upper part of transverse process; insertion, into outer surface and lower border of the laminae of the verteljrae above, from the first and second dorsal to the eleventh and twelfth; action, ro- tates the spinal column; nerves, anterior branches of the pos- terior divisions of the dorsal. Sdpraspinales. — Origin and insertion, the spinous proc- esses in the cervical region of the vertebrae; action, extends cervical spine; nerves, internal branches of the posterior divi- sions of the cervical. Interspinales. — Consist of muscular bands in pairs be- tween the spinous processes of the adjoining vertebras : six pairs in the cervical region, three pairs in the dorsal, four or five in the lumbar; action, extend the spine; nerves, internal branches of the posterior divisions of the cervical, dorsal, and lumbar. Extensor Cocotgis. — Origin, from the first piece of the coccyx, or last bone of the sacrum ; insertion, into the lower ex- tremity of the coccyx; action, rudimental in man. Inteeteansversales. — Are small, muscular bands between the transverse processes: in the cervical region seven pairs, in the dorsal twelve pairs, in the lumbar region four pairs ; action, flex the spine laterally ; nerves, internal branches of the posterior division of the cervical, dorsal, and lumbar. Eeotus Capitis Posticus Major. — Origin, from the spi- nous process of the axis; insertion, into inferior curved line of the occipital bone; action, rotates the atlas and the cranium; nerve, the suboccipital. Eectus Capitis Posticus Minor. — Origin, from the tu- bercle of the posterior arch of the atlas; insertion, below the inferior curved line of the occipital bone; action, draws the head backward; nerve, suboccipital. Obliquus Capitis Inferior. — Origin, spinous process of the axis; insertion, lower back portion of the transverse process of the atlas; action, rotates the atlas and the cranium; nerves, suboccipital and great occipital. Obliquus Capitis Superior. — From upper surface of the transverse process of the atlas; insertion, between the curved lines of the occipital bone to the outer side of the complexus; action, rotates the atlas ; nerves, suboccipital and great occipital. 146 HUilAN ANATOMY. MUSCLES OF THE ABDOillNAL-THOKAOIC REGION. Obliquus Exteenus. — Origin, lower borders of the eight lower ribs; insertion, the lowermost miiscular fibres, into the anterior half of the iliac crest; the other muscular fibres, by a broad aponeurosis, which joins the anterior half of the aponeu- rosis of the internal oblique to form the anterior walls of the sheath of the rectus, above into the ensiform cartilage, below into the symphysis pubis. In the median line it blends with its fellow of the opposite side to form the linea alba. A slit in its lower portion above the pubic spine is called the external abdominal ring. The lower thickened portion of the aponeu- rosis stretching between the anterior superior iliac spine and the pubic spine is called Poupart's ligament. A reflection from it to the ilio-pectineal line is called Gimbernat's ligament ; action, flexes the pelvis on the thorax, or vice versa, and compresses the viscera; nerves, lower intercostal, ilio-hypogastrie, and ilio-in- guinal. Obliqdus Inteenus. — Origin, from the outer half of Pou- part's ligament, from the anterior two-thirds of the crest of the ilium, and the posterior lamellae of the lumbar fascia ; insertion, above to the lower four costal cartilages, below, conjointly with the tendon of the transversalis, into the os pubis and linea ilio- pectinea, to form the conjoined tendon, and into the median line (linea alba) by an aponeurosis extending from the sternum and seventh and eighth costal cartilages to the pubis. This apo- neurosis at its lower fourth consists of two united laminas passing in front of the rectus muscle, but in its upper three-fourths it divides, one lamina passing in front of the rectus and joining the aponeurosis of the external oblique, the other passing behind and joining the aponeurosis of the transversalis; action, same as the externus ; nerves, same as externus. Teansveesalis. — Origin, from outer third of Poupart's ligament and anterior three-fourths of the crest of the ilium, from the inner surface of the cartilages of the six lower ribs, and from the spinous and transverse processes of the lumbar verte- bras; insertion, by the conjoined tendon into the linea ilio-pee- tinea and crest of the os pubis; action and nerves, same as the externus. Eectus Abdominis. — Origin, by two tendons — the outer from the crest of the pubis, the inner interlacing with its fellow of the opposite side; insertion, into the cartilages of the fifth, sixth, and seventh ribs; action, depresses the thorax, flexes the vertebral column, and, acting from above, flexes- the pelvis upon the vertebral column. THE MUSCtlLAR SYSTEM. 147 Ptraiiidalis. — Origin, from the os pubis and anterior pubic ligament; insertion, into the linea alba, midway between the OS pubis and umbilicus. QuADKATUS Ldmboeum. — Origin, by two portions — ^pos- terior portion, from crest of the ilium and ilio-lumbar ligament; Fig. 95. Muscles of abdomen: 12, external oblique; 16, rectus abdominis; internal oblique; 17, pyramidalis; 19, quadratus lumborum. IS. insertion, into lower border of the last rib and transverse proc- esses of the three lower lumbar vertebrae; origin, anterior por- tion, from upper border of the transverse processes of the lumbar vertebrae, from the third to the fifth ; insertion, into one-half the lower margin of the last rib and apices of upper four lumbar vertebrse; action, draws down and fixes the last rib and assists in inspiration and expiration. 148 HUMAN ANATOMY. MUSCLES 01' THE THORAX. Intercostales Bxterni. — Eleven pairs on either side. Origin, from the outer border of the groove on the lower border of each rib, from the cartilage to the tubercle; insertion, into upper border of the rib below; action, raises the ribs; nerve, intercostal. Intercostales Interni. — Origin, inner lip of the groove on the lower border of each rib ; insertion, into the upper border of the rib below; action, pulls the ribs upward; nerve, inter- costal. Diaphragm: 1, 2, 3, central oordilorm tendon; 4, middle leaflet; 5, ligamentura arcuatum externum; 6, ligamentum arcuatum in- ternum; 8, right crua; 10, left crus; 11, aortic opening; 12, oesopha- geal opening; 13, opening for vena cava; 14, psoas magnus; 15, quadra- tus lumborum. Ikfracostales (Subcostales). — Origin, inner surface of rib ; insertion, into the inner surface, from the first to the third rib below; action, inspiratory muscles; nerve, intercostal. 1 Triangularis Sterni. — Origin, lower part of the back of the sternum and back of ensiform cartilage and inner surface of sternal end of the costal cartilages of the three or four lower ribs; insertion, into the border and inner surfaces of the costal cartilages, from the second to the sixth rib inclusive (it is con- tinuous below with transversalis abdominis muscle) ; action, draws down the costal cartilages; nerve, intercostal. THE MUSCULAR SYSTEM. 149 Levatores Costaeum. — Twelve on either side. Origin, from transverse processes of seventh cervical and eleven upper dorsal vertebrae; insertion, into upper surface of the rib below, between the angle and tubercle; action, raises the ribs; nerve, the intercostal. DIAPHRAGMATIC EE6I0N. Diaphragm. — Origin, inner surface of ensiform cartilage, cartilages and bony portions of siz or seven lower ribs in front, and from two aponeurotic arches, the ligamentum arcuatum ex- ternum and internum and the lumbar vertebras behind; inser- tion, into the circumference of the central or cordiform tendon; action, the principal muscle of inspiration and expulsion; nerves, the phrenic, and phrenic plexus of the sympathetic. THE openings IN THE DIAPHRAGM. The aortic opening is placed posteriorly between the two crura, in front of the spine. It transmits the aorta, thoracic duct, and vena azygos major, and sometimes the left sympa- thetic nerve. The oesophageal opening is in front of the decussation of the crura, a little to the left and in front of the aortic opening. It transmits the oesophagus and pneumogastric nerves. The foramen quadratum, or opening for the vena cava, is placed at the highest part of the central tendon, a little to the right. It transmits the vena cava and maintains its patency. The crus on each side transmits the sympathetic and greater and lesser splanchnic nerves, and in addition the left transmits the vena azygos minor. JICSCLES OF THE PERIXEUM (vide PERIXEUM). Muscles of the Upper Extremity, muscles of the shoulder. Pectoralis Major. — Origin, from the sternal half of the clavicle ( clavicular portion) and anterior surface of the sternum and costal cartilages from the second to the sixth or seventh rib (sterno-costal portion) ; insertion, into the anterior bicipital ridge of the humerus ; action, draws the arm across the chest ; nerves, external and internal anterior thoracic. Pectokalis JUixoR. — Origin, upper and outer surface of the third to the fifth rib, inclusive, and the aponeurosis of the inter- costal muscles; insertion, into anterior border of the coracoid ]3rocess of the scapula; action draws the scapula inward and downward ; nerve, the anterior thoracic. 150 HUMAN ANATOMY. SuBCLAVius. — Origin, from cartilage of the first rib ; inser- tion, under surface of the clavicle, about its middle third ; action, depresses the shoulder, and draws clavicle forward and down- ward; nerve, branch from the union of the fifth and sixth cervical. Sberatus Magnus. — Origin, by nine muscular portions, from the outer surface of the eight upper ribs (two divisions being from the second rib) and from the upper intercostal apo- neurosis ; insertion, by three divisions — ^upper portion into supe- rior angle of the scapula, middle portion into posterior portion of the scapula between the inferior and superior angles, lower portion into the inferior angle of the scapula; action, raises the vertebral border and carries the scapula forward; nerve, the pos- terior thoracic. Deltoid. — Origin, from the outer third of the clavicle and from the outer part of the acromion process and the lower bor- der of the scapular spine; insertion, into the outer side of the shaft of the humerus, about its middle into the deltoid tubercle ; action, raises the arm; nerve, the circumflex. SuBSOAPULARis. — Origin, from the internal two-thirds of the subscapular fossa; insertion, into the lesser tuberosity of the humerus; action, rotates the head of the humerus inward; nerves, upper and lower subscapular. SuPEASPiNATtrs. — Origin, from inner two-thirds of supra- spinous fossa; insertion, into the uppermost facet of the great tuberosity of the humerus; action, assists in raising the arm and fixing the head of the humerus; nerve, the suprascapular. Infeaspinatus. — Origin, from the inner two-thirds ojf the infraspinous fossa; insertion, into the middle facet of the great tuberosity; action, rotates the head of the humerus outward; nerve, supra-scapular. Teees Mince. — Origin, posterior surface of the upper two- thirds of the axillary border of the scapula; insertion, into the lowest facet of the great tuberosity; action, rotates the head of the humerus outward; nerve, the circumflex. Teees Major. — Origin, posterior surface of the inferior angle of the scapula; insertion, into the posterior bicipital ridge of the humerus ; action, draws the humerus backward and down- ward, assisting the latissimus dorsi ; nerve, the lower subscapular. HUMERAL REGION. CoRAco-BRACHiALis. — Origin, from apex of the coracoid process of the scapula; insertion, into the inner side of the middle of the shaft of the humerus, between the origin of the THE MUSCU1.AR SYSTEM. 151 brachialis anticus and short humeral head of the triceps; action, draws the huinerus inward and forward; nerves, the musculo- cutaneous and a branch of the musculo-spiral. Biceps (Flexor Cubiti). — Origin, by two heads; short head from the apex of the coracoid process, along with the coraco- brachialis; the long (glenoid) head from the upper margin of the glenoid cavity of the scapula; insertion, into the posterior Muscles of slioulder and arm; 4, subscapularis; 5, teres major; 6, coraco-bracliialis ; 7, biceps. Triceps muscle; 1, external head; 2, scapular head; 3, internal head; 4, insertion. part of the tuberosity of the radius ; action, flexes the forearm ; nerve, the musculo-cutaneous. Brachialis Anticus. — Origin, inner and outer surfaces of the shaft of the humerus, embracing the insertion of the deltoid ; insertion, into the anterior surface of the coronoid process of the uhia; action, flexes the forearm; nerve, the musculo- cutaneous. Triceps (Extensor Cubiti). — Origin, by three heads — middle, or scapular head, below the glenoid cavity of the scapula ; external head, from the posterior aspect of the shaft of the 152 HUMAN ANATOMY. humerus, between the upper part of the musculo-spiral groove and the insertion of the teres minor, and from the external intermuscular septum; the internal head, from the posterior aspect of the shaft of the humerus, below the insertion of the teres major, and below the groove for the musculo-spiral nerve; insertion, by a common head, into the posterior part of the under surface of the olecranon process of the ulna; action, extends the forearm; nerve, the musculo-spiral. SuBANCONEUS. — Origin, from the posterior surface of the humerus, above the olecranon fossa; insertion, into the posterior ligament of the elbow-joint; action, draws up the posterior liga- ment of the elbow-Joint during extension of the forearm; nerve, the musculo-spiral. MUSCLES OF THE FOEEAEM. Anterior Superficial Layer. Peonatoe Eadii Teees. — Origin, by two heads — ^the smaller from the coronoid process of the ulna, the larger from the humerus, above the internal condyle, and from the common tendon of this group of muscles, and intermuscular septum; in- sertion, into the outer aspect of the shaft of the radius; action, pronates the hand ; nerve, the median. Plexoe Caepi Eadialis. — Origin, by the common ten- don from the inner condyle, and from the intermuscular septum between it and the pronator teres ; insertion, into the base of the metacarpal bone of the index finger; action, flexes the wrist; nerve, the median. Palmaeis Longus. — Origin, from the inner condyle of the humerus by the common tendon and the intermuscular septum and the deep fascia ; insertion, into the annular ligament, spread- ing out in the palmar fascia; action, renders tense the palmar fascia; nerve, the median. Plexor Caepi Ulnaeis. — Origin, by two heads — one by the common tendon from the inner condyle of the humerus, the other from the inner margin of the olecranon, the inter- muscular septum, between the ulna and the flexor sublimis digi- torum, and from the upper two-thirds of the posterior aspect of the ulna; insertion, pisiform bone; action, flexes the wrist; nerve, the ulnar. Flexor Sublimis Digitoeum (Perfoeatus). — Origin, by three heads — one from the coronoid process of the ulna, above the pronator radii teres, another from the common tendon from the internal condyle of the humerus, and the third from the THE MXrSCULAE SYSTEM. 153 oblique line of the radius, from the insertion of the pronator radii teres, to the tubercle ; each tendon divides at the base of the first phalanges to allow the passage of the tendon of the flexor profundus digitorum ; insertion, into the lateral margins of the second phalanges by four tendons ; action, flexes the second phalanges ; nerve, median. Anterior Deep Layer. Plexok Profundus Digitorum (Peeforans). — Origin, from the inner side of the coronoid process, and from the upper two-thirds of the front and inner aspect of the shaft of the ulna, between the brachialis anticus above and the pronator quadratus below ; insertion, by four tendons into the bases of the last phalanges, perforating the tendons of the flexor sublimis ; action, flexes the phalanges ; nerves, the anterior interosseous and the ulnar. Flexor Longus Pollicis. — Origin, from the upper two-thirds of the shaft of the radius and from the interosseous membrane ; insertion, into the base of the last phalanx of the thumb ; action, flexes the phalanges ; nerve, the ante- rior interosseous. Pronator Quadratus. — Origin, anterior border of the ulna and from the oblique line of the lower fourth of the anterior aspect of the ulna ; insertion, into the lower fourth of the front and outer border of the shaft of the radius ; action, pronates the hand ; nerve, anterior in erosseous. Muscles of forearm: 1, biceps; 2, brachialis anticus; 3, triceps; i, supinator longus; 5, ex- tensor carpi radialis lon- gior; 6, extensor carpi radialis brevier; 8, ex- tensor communis digi- torum; 12, extensor carpi ulnaris; 13, U, extensors of thumb. Radial Region. Supinator Longus. — Origin, irora the upper two-thirds of the external condyloid ridge of the humerus and the external intermuscular septum ; insertion, into the base of the styloid process of the radius ; action, supinates the hand ; nerve, musculo-spiraL 164 HUMAN ANATOMY. Extensor Carpi Radialis Longior. — Origin^ from the lower third of the external condyloid ridge of the humerus, from the common tendon of the extensor muscles of the forearm from the external condyle, and the intermuscular septum; insertion, into the radial side of the base of the metacarpal bone of the in- dex finger; action, extends the wrist; nerve, musculo-spiral. Extensor Carpi Eadialis Brevior. — Origin, from the common tendon, from the external condyle of the humerus, the external lateral ligament, and the intermuscular septum; inser- tion, into the base of the metacarpal bone of the middle finger on its radial side; action, extends the wrist; nerve, posterior interosseous. Posterior Superficial Layer. Extensor Communis Digitorum. — Origin, from the ex- ternal condyle of the humerus, the deep fascia, and the inter- muscular septa; insertion, by four tendons into the second and third phalanges of all the fingers (at the first phalanx they receive the insertion of the lumbricales and interossei) ; action, extends the fingers; nerve, posterior interosseous. Extensor Minimi Digiti. — Origin, from the common ten- don from the external condyle and the intermuscular septum; insertion, into the second and third phalanges of the little finger ; action, extends the little finger; nerve, posterior interosseous. Extensor Carpi TJlnaris. — Origin, from the middle third of the posterior border of the ulna, from the common tendon from the external condyle of the humerus, and from the fascia of the forearm; insertion, into the base of the metacarpal bone of the little finger, on the ulnar side; action, extends the wrist; nerve, posterior interosseous. Anconeus. — Origin, from the outer condyle of- the hu- merus ; insertion, into the upper fourth of the posterior aspect of the shaft of the ulna and the side of the olecranon; action, extends the forearm; nerve, musculo-spiral. Posterior Deep Layer. Supinator Brevis. — Origin, from the external lateral liga- ment of the elbow-joint, from the external condyle of the hu- merus, from the ulna below the lesser sigmoid cavity, and from the orbicular ligament of the radius; insertion, into the neck, bicipital tuberosity and oblique line of radius ; the posterior interosseous nerve pierces this muscle; action, supinates the hand; nerve, posterior interosseous. THE iirSCULAR SYSTEII. 155 ExTENTSOR Ossis Metacaepi Pollicis. — Origin, middle third of posterior surface of radius, posterior surface of louver three-fourths of shaft of ulna, and interosseous ligament; iiiser- tian, into Ijase of metacarpal of thumb; action, extends meta- carpal of thumb ; net ve, posterior interosseous. ExTEXHOR LoxGUS PoLLicis. — Origin, from the interosseous membrane and from the posterior aspect of the shaft of the ulna; insertion, into the base of the last or terminal phalanx of the thumb ; action, extends the thumb ; nerve, posterior inter- usseous. EXTEXSOR Bre\-is Polmcis. — Origin, from the posterior aspect of the shaft of the radius and from the interosseous mem- brane ; insertion, into the base of the first phalanx of the thumb ; action, extends the thumb ; nerve, posterior interosseous. ExTEXsoR IxDicis. — Origin, from the posterior aspect of the ulna and from the interosseous membrane; insertion, into the second and third phalanges of the index finger, along with the tendon of the extensor communis; action, extends the index finger; nerve, posterior interosseous. MUSCLES or THE HAXD. These are divided into three groups — the radial region, the ulnar region, and the palmar region; Radial Region. Abductor Pollicis. — Oiigin, from the annular ligament and ridge of the trapezium; insertion, into base of the first phalanx of the thumb on its radial side; action, abducts the thumb from the median line ; nerve, median. OproxEXs Pollicis. — Origin, from annular ligament and palmar surface of the trapezium ; insertion, into metacarpal bone of the thumb throughout the whole length of its radial side; action, flexes the first metacarpal bone; nerve, median. Flexor Breyis Pollicis. — This muscle consists of two por- tions, divided bj- tendon of fiexor longus pollicis; on-igin, super- ficial portion from outer two-thirds of annular ligament and trapezium, deeper portion (by some called adductor obliquus pollicis) from sheath of flexor carpi radialis, the trapezoid, os magnum, and base of first, second, and third metacarpal bones; insertion, into either side of base of first phalanx of thumb, the inner portion joining the adductor and the outer portion the abductor; each tendon has a sesamoid bone developed in it; action, adducts thumb toward median line; nerves, ulnar and median. 156 HUMAN ANATOMY. Adductor Pollicis (Adductor Transversus Pollicis) . — Origin, from lower two-thirds palmar surface of middle meta- carpal bone; insertion, into ulnar side of base of first thumb phalanx and internal sesamoid bone; action, adducts thumb; nerve, ulnar. Ulnar Region. Palmaris Brevis. — Origin, from palmar fascia and annu- lar ligament; insertion, into skin of palm of hand; action, wrinkles skin of hand; nerve, ulnar. Abductor Minimi Digiti. — Origin, from pisiform bone and from tendon of flexor carpi ulnaris; insertion, into base of first phalanx of little finger on its ulnar side; action, abducts little finger from median line ; nerve, ulnar. Flexor Brevis Minimi Digiti. — Origin, from annular ligament and tip of unciform process of unciform bone; inser- tion, into the first phalanx of little finger; action, flexes little finger; nerve, ulnar. Opponens Minimi Digiti. — Origin, from annular ligament and from unciform process of unciform bone; insertion, into ulnar border of whole length of metacarpal bone of little finger; action, flexes little finger; nerve, ulnar. Palmar Region. LuMBRiCALES. — Origin, by four fleshy tendons from the tendons of the deep flexors — the first and second from the palmar surface and radial side of the tendons of the index and middle fingers, the third from the adjoining sides of the tendons of the middle and ring fingers, and the fourth from the adjoining sides of the tendons of the ring and little fingers ; insertion, on dorsal aspect of each finger into the expansion of the extensor communis digitorum ; action, abduct the fingers to either side of the median line; reereeSj ulnar and median. Interossei muscles consist of two groups, the dorsal and palmar. Dorsal Interossei. — Origin, from two heads from the adjacent sides of the metacarpal bones, four in number; inser- tion, the first into the radial side of the base of the first phalanx of the index finger, the second into the radial side of the middle finger, the third into the ulnar side of the middle finger, and the fourth into the ulnar side of the ring finger, the middle finger having two, one on either side; action, abduct the fingers from the median line; nerve, ulnar. THE MUSCtJIiAE SYSTEM. 157 Palmar Inteeossei. — Three in number. Origin, the first from the entire length of the metacarpal bone of the index finger on its ulnar side, the second from the ring finger on its radial side, and the third from the little finger on its radial side; insertion^ into the base of the first phalanx and into the ex- pansion of the extensor communis tendon of the same finger from which they arise; action, adduct the fingers toward the median line; nerve, ulaar. Muscles of the Lower Extremity, iliac region. Psoas Magnus. — Origin, from the sides and anterior sur- faces of the transverse processes and bodies of the last dorsal and all the lumbar vertebrae and the intervertebral substances between them ; insertion, into the lesser trochanter of the femur, uniting with the tendon of the iliacus ; this muscle is in relation behind with the capsular ligament of the hip, being separated from it by a synovial bursa; action, flexes and rotates the femur inward, and also flexes the trunk and pelvis on the thigh; nerves, an- terior branches of the lumbar nerves. Psoas Parvus. — Origin, from the lateral surfaces of the bodies of the last dorsal and first lumbar vertebrae and from the intervertebral substances between them; insertion, into the ilio- pectineal eminence, joining the iliac fascia; action, assists the psoas magnus and renders tense the iliac fascia; nerves, anterior branches of the lumbar. Iliacus. — Origin, from the base of the sacrum and the ilio- lumbar ligament behind, from the iliac fossa and inner margin of the crest of the ilium and the anterior superior and anterior inferior spinous processes of the ilium; insertion, into the oblique or intertrochanteric line of the femur to the outer side of the insertion of the psoas; action, flexes and rotates the femur inward and flexes the trunk and pelvis on the thigh; nerves, anterior crural and the anterior branches of the lumbar. femoral and GLUTEAL REGION. Tensor VAGiN-a; Femoris. — Origin, from the anterior supe- rior spinous process between the sartorius and the gluteus medius, and from the fore part of the outer lip of the crest of the ilium ; insertion, into the fascia lata, about the upper fourth of the outer side of the thigh; action, renders tense the fascia lata; nerve, superior gluteal. 158 HLTMAN ANATOMY. Sartoeius. — Origin, from the anterior superior spinous process of the ilium and the upper part of the notch below ; in- sertion, into the inner and upper part of the shaft of the tibia; action, flexes the leg upon the thigh and the thigh upon the pelvis ; nerve, branches of the anterior crural. Quadriceps Extensor. — This extensive muscle covers the front and sides of the femur and consists of four portions — the rectus femoris, the vastus extemus, the vastus internus, and the crureus. Rectus Femoeis. — Origin, by two heads — one, the long head, from the groove above the brim of the acetabulum; the other, the short head, from the anterior inferior spinous process of the ilium ; insertion, by the common tendon into the patella ; action, extends the leg upon the thigh; nerves, branches of the anterior crural. Vastus Bxtbenus. — Forms the greater part of the quadri- ceps extensor. Origin, by an extensive aponeurosis extending from the tubercle of the femur along the anterior border of the great trochanter and the whole length of the outer lip of the linea aspera; insertion, into the outer portion of the patella, joining the common tendon; action, extends the leg on the thigh ; nerves, branches of the anterior crural. Vastus Inteenus and Ceueeus. — Origin, by a tendinous aponeurosis extending from the neck of the femur throughout the whole length of the inner lip of the linea aspera ; insertion, into the common tendon, together with the rectus femoris and the vastus externus ; action, extends the leg on the thigh ; nerves, branches of the anterior crural. SuBCEUEEUS. — Origin, from the anterior aspect of the lower part of the shaft of the femur; insertion, into the synovial sac behind the patella. Gracilis. — Origin, from the inner margin of the ramus of the ischium and the pubes; insertion, into the inner aspect of tne shaft of the tibia, below the tuberosity; action, flexes the leg and draws it inward, assisting the sartorius ; nerve, the obtu- rator. Pectineus. — Origin, from the tendinous prolongation of Gimbernat's ligament and the linea ilio-pectinea ; insertion, into the rough line leading to the linea aspera from the lesser tro- chanter; action, adducts the thigh; nerves, obturator, the ac- cessory obturator, and branches of the anterior crural. Adductor Longus. — Origin, from the front of the os pubis below the crest; insertion, into the middle third of the linea aspera, between the adductor ma gnus and the vastus internus; action, adducts the thigh; nerve, obturator. THE MUSCULAK SYSTEM. 169 Adductoe Beevis. — Origin, from the outer aspect of the descending ramus of the pubes, between obturator extemus and the gracilis; insertion, into the upper part of the linea aspera; action, adducts the thigh ; nerves, branches of the obturator. Adductor Magnus. — Origin, from the luberosity of the ischium, the as- ■ — cending ramus of the ischium, and from the descending ramus of the pubes ; insertion, into the rough line leading to the linea aspera from the great trochanter to the inner side of the gluteus maximus and into the adductor tubercle above the inner con- dyle of the femur (z)tie page 64) ; action, adducts the thigh ; nerves, the obtura- tor and a branch from the great sciatic. Gluteus Maximus. — Origin, from the superior curved line of the ilium, the posterior aspect of the last segment of the sacrum, the border of the coccyx, and the surface of the great sacro- sciatic and posterior sacro-iliac liga- ments ; insertion, into the rough line leading to the linea aspera from the great trochanter, between the adductor magnus and the vastus externus, and into the fascia lata ; action, it is a ten- sor of the fascia lata, and an external rotator and extensor of the thigh ; nerves, the inferior gluteal and a branch from the sacral plexus. Gluteus Medius. — Origin, from the outer lip of the crest and the outer aspect of the ilium, between the middle and superior curved lines, and from the gluteal aponeurosis ; insertion, into the oblique line on the outer surface of the great trochanter ; action, rotates the thigh outward ; nerve, from the superior gluteal. Gluteus Minimus. — Origin, from the border of the great sacro-sciatic notch and between the inferior and middle curved lines of the outer aspect of the ilium ; insertion, into a depression on the front border of the great trochanter; action, rotates the thigh inward ; nerve, superior gluteal. FIG. 100. Anterior femoral re- gion: 4, tensor vaginas femoris; 6, sartorius; 6, rectus; 7, vastus exterr nus; 8, vastus internus; 10, iliacus; 11, psoas; 12, pectlneus; 13, adductor longus; 14, adductor mag- nus; 15, gracilis. 160 HUMAN ANATOMY. PTRiroKMis. — Origin, from the anterior surface of the sa- crum, between the first to the fourth anterior sacral foramen, and from the margin of the great sacro-sciatic foramen, and from the anterior aspect of the great sacro-sciatic ligament ; insertion, into the upper border of the great tro- chanter, with the tendon of the obtu- rator internus ; action, rotates, femur outward ; nerve, obturator. Obturator Internus. - — Origin, from the inner side of the obturator foramen and the internal surface of obturator membrane and the anterior and external wall of the pelvis it passes out of the pelvis through the lesser sacro-sciatic notch and receives the ten- dons of the gemelli muscles ; insertion, into the upper border of the great tro- chanter in front of the pyriformis ; action, rotates the thigh outward ; nerves, branches from the sacral plexus. Gemelli consist of two muscles — gemellus superior and gemellus in- ferior. Gemellus Superior. — Origin, from the outer aspect of the spine of the ischium ; insertion, into the tendon of the obturator internus. Gemellus Inferior. — Origin, from the outer border of tuberosity of the ischium ; insertion, into the lower portion of the tendon of the obturator internus ; action, rotates the thigh out- ward ; nerves, branches of the sacral plexus. QuADRATUS Femoris. — Origin, from the external border of the tuber- osity of the ischium ; insertion, into the upper part of the linea quadrati, on the posterior aspect of the trochanter major; action, rotates the thigh outward; nerves, branches of the sacral plexus. Obturator Externus. — Origin, from the inner two-thirds of the anterior aspect of the obturator membrane and the tendi- nous arch covering the canal for the obturator vessels and nerves, and the descending ramus of the pubis and the ascending ramus Fig. 101. Posterior femoral re- gion: 1, gluteus medlus; 2, gluteus maximus; 3, vastus externus; 4, bi- ceps; 6, semi-tendinosls; 7, semi-membranosls; 8, gracilis. THE MUSCULAR SYSTEM. 161 of the ischium; insertion, into the digital fossa of the femur; action, rotates the thigh outward; nerve, the obturator. BiGBTS.— Origin, arises by two heads — the long head, from the lower and inner facet of the tuberosity of the ischium by a common tendon to the long head of the semi-tendiaosus ; the short head, from the outer lip of the liaea aspera, between the vastus extemus and the adductor magnus; insertion, outer side of head of fibula, by two portions, on either side of the external lateral ligament of the knee-joint, sending a band forward to the outer tuberosity of the tibia. This tendon forms the outer ham- string; action, flexes the leg upon the thigh; nerve, the great sciatic. Semi-tendinosus. — Origin, from a common tendon, to- gether with the long head of the biceps, from the tuberosity of the ischium; insertion, into the inner and upper part of the shaft of the tibia; action, rotates the leg inward; nerve, the great sciatic. Semi-membeanosus. — Origin, from the outer and upper facet of the tuberosity of the ischium to the outer side and above the origin of the semi-membranosus and the biceps; insertion, into the posterior inner aspect of the inner tuberosity of the tibia, under the internal lateral ligament by three portions — the internal portion into the inner side of the internal tuberosity, the posterior portion into the posterior part of the outer condyle of the femur, forming the principal part~Tof the posterior liga- ment of the knee-joint, and middle portion into the posterior aspect of the inner tuberosity; action, flexes the leg upon the thigh; nerve, the great sciatic. These two tendons (semi-membranosus and semi-tendino- sus), together with the gracilis and sartorius, form the inner hamstring. MUSCLES OP THE LEG. Anterior Set. Tibialis Anticus. — Origin, from the upper and outer two- thirds of the shaft of the tibia, the interosseous membrane, and the intermuscular septum between it and the extensor longus digitorum; insertion, into the lower and inner aspect of the internal cuneiform bone and base of the first metatarsal bone; action, flexes the -foot; nerve, the anterior tibial. Extensor Peoprius Pollicis. — Origin, from the anterior aspect of the fibula and the interosseous membrane and the intermuscular septum between it and the extensor longus digi- torum; insertion, into the base of the last phalanx of the great toe; action, extends the great toe; nerve, the anterior tibial. 162 HUMAN ANATOMY. Extensor Longus Digitorum. — Origin, from the upper two-thirds of the anterior aspect of the shaft of the fibula and the outer tuberosity of the tibia, from the deep fascia of the inter- osseous membrane and the intermuscular septum, between it and the perinei on the outer and the tibialis anticus on the inner side; insertion, by three tendons into the bases of the first and I FlQ. 102. Anterior muscles ol leg: 3, tibialis Bntlcus; 4, extensor longus digitorum; B, extensor proprius pollicis; 6, pe- roneus tertius; 7, peroneus longus; 8, peroneus brevis. Fig. 103. Posterior muscles of leg: 4, pop- liteus; 5, gastrocnemius: 6, tendo Acliillis; 8, tendons of peroneus longus and brevis; 9, tibialis posticus and fiexors. second phalanges of the four lesser toes, the innermost tendon dividing into two; each tendon (except the fourth) is joined opposite the metatarso-phalangeal joint by the tendon of the extensor brevis digitorum, and receives an expansion from the lumbricales and interossei; at the first interphalangeal articula- tion the tendons divide into three slips — ^the middle one for inser- THE MUSCULAE SYSTEM. 163 tion into the second phalanges, and the other two tmiting to be inserted into the bases of the third ; action, extends the phalanges and, continuing, flexes the foot upon the leg; nervBj the anterior tibial. Peeoneus Teetius. — Origin, from the lower front part of the fibula on its outer side, from the interosseous membrane, and the intermuscular septum between it and the peroneus brevis; insertion, into the metatarsal bone of the little toe ; action, flexes the tarsus upon the leg; nerve, the anterior tibial. Posterior Superficial Set. Gasteocnemius. — Origin, from the upper and back part of the external and internal condyles of the femur on either side, above the origin of the popliteus, and from the supracondyloid ridges; insertion, by joining with the tendon of the soleus to form the tendo Achillis; action, extends the foot; nerve, from the internal popliteal. Soleus. — Origin, from the oblique line of the tibia, from the middle third of the internal border, and from the posterior surface of the head of the fibula; insertion, by joining with the tendon of the gastrocnemius to form the tendo Achillis ; action, extends the foot; nerve, the internal popliteal. Tendo Achillis. — Origin, from the union of the gastroc- nemius and soleus; is the largest and strongest tendon in the body; it is inserted into the inferior surface of the posterior tuberosity of the os calcis, having a synovial bursa between it and the bone. Plantakis. — Origin, from the posterior ligament of the knee-joint and the lower portion of the outer division of the linea aspera ; insertion, into the posterior part of the os calcis to the inner side of the tendo Achillis; action, the rudiment of a muscle intended to render tense the plantar fascia; nerve, the internal popliteal. Posterior Deep Set. Popliteus. — Origin, from the outer side of the external con- dyle, from the posterior ligament of the knee-joint; insertion, above the oblique line on the posterior aspect of the shaft of the tibia; action, assists in flexing the leg; nerve, the internal pop- liteal. Flexoe Longus Polliois. — Origin, from the lower two- thirds of the internal surface of the fibula, from the lower part of the interosseous membrane, fascia covering tibialis posticus. 164 HUMAN ANATOMY. and from the intermuscular septum; it passes behind the in- ternal malleolus; insertion, into the base of the last phalanx of the great toe; nerve, posterior tibial. Flexor Longus Digitoeum (Peefobans). — Origin, from the posterior aspect of the tibia, below the oblique line ; it passes behind the internal malleolus; insertion, into the bases of the last phalanges of the four lesser toes, passing through the division in the tendons of the flexor brevis digitorum ; action, flexes the phalanges and, continuing, extends the foot on the leg; nerve, the posterior tibial. Tibialis Posticus. — Origin, from the posterior aspect of the shaft of the tibia, and from the upper two-thirds of the shaft of the fibula, and from the whole length of the interosseous mem- brane and the intermuscular septa on either side of it; it passes behind the inner malleolus; insertion, into the internal cunei- form bone and the tuberosity of the scaphoid; it contains a sesamoid bone in its tendon ; action, extends the tarsus upon the leg; nerve, the posterior tibial. Fibular Region. Pbeoneus Longus. — Origin, from the upper two-thirds of the shaft of the fibula, the head of the fibula, the intermuscular septa, and the deep fascia; it passes behind the outer malleolus along with the peroneus brevis; insertion, into the base of the metatarsal bone of the great toe; this tendon changes its course twice, first at the external malleolus, second at the cuboid bone, and usually has a sesamoid bone developed in its tendon ; action, extends and everts the foot upon the leg; nerve, musculo-cutane- ous branch of the external popliteal. Peroneus Beevis. — Origin, from the outer and middle third of the fibula and the intermuscular septa on eitier side; insertion, into the upper surface of the base of the metatarsal bone of the little toe; action, extends the foot upon the leg; nerve, musculo-cutaneous branch of the external popliteal. muscles of the foot. Dorsal Region. ExTENSOE Brevis Digitoeum. — Origin, from the external calcaneo-astragaloid ligament, from the annular ligament, and from the outer side of the os calcis ; insertion, by four tendons — the first one into the first phalanx of the great toe, the other three into the long extensor tendons of the second, third, and fourth THE MUSCULAR SYSTEM. 165 toes on their outer sides ; action, extends the phalanges of the four inner toes and the first phalanx of the great toe; nerve, the an- terior tibial. Plantar Region. The muscles of this region are divided into four sets. First Set. Abductor Hallucis. — Origin, from the internal annular ligament, from the inner tubercle of the os calcis, from the inter- muscular septum on its outer side ; plantar fascia ; insertion, into the base of the first phalanx of the great toe on its inner side; action, abducts the great toe; nerve, branch of the internal plantar. Flexor Brevis Digitorum (Perfgratus) . — Origin, from the inner tubercle of the os calcis, from the plantar fascia and the intermuscular septa between it and the muscles on either side; insertion, into the second phalanges by a process on either side, allowing the passage of the tendon of the flexor longus digitorum; action, flexes first lesser toes; nerve, from the in- ternal plantar. Abductor Minimi Digiti. — Origin, from the plantar fascia, from the os calcis in front of the tubercle, and from the inter- muscular septum on its inner side; insertion, into the first phalanx of the little toe on its outer side; action, abducts little toe; nerve, external plantar. Second Set. Flexor Accessorius. — Origin, by two heads, one from the OS calcis in front of the outer tubercle, the other from the inner surface of the os calcis and the calcaneo-scaphoid ligament; in- sertion, by a common tendon into the tendon of the flexor longus digitorum; action, accessory to the long flexor; nerve, external plantar. LuMBRiCALES. — FouT Small muscles. Origin, from the ten- dons of the flexor longus digitorum, after their division, each one arising from two tendons; insertion, into the base of the first phalanx of the same toe and the tendinous expansion of the extensor longus digitorum; nerves, the external plantar nerve supplies the two external, the internal plantar nerve the two in- ternal, muscles. IQQ HUMAN ANATOMY. Third Set. Flexor Beevis Hallucis. — Origin, from the external cune- iform bone, and the internal border of the cuboid bone, and the tendinous expansion of the tibialis posticus ; insertion, into the first phalanx of the great toe on either side; action, flexes first phalanx of great toe ; nerves, the internal plantar and some- times a branch from the external plantar. Adduotoe Hallucis. — Origin, from the bases of the second, third, and fourth metatarsal bones and from the sheath of the tendon of the peroneus longus; insertion, into the base of the first phalanx of the great toe on the outer side ; nerve, external plantar. Flexoe Brbvis Minimi Digiti. — Origin, from the sheath of the peroneus longus and the base of the metatarsal bone of the little toe J insertion, into the outer side of the base of the little toe; nerve, external plantar. Teansveesus Pedis. — Origin, from the inferior surface of the head of the fifth metatarsal bone, from the transverse meta- tarsal ligament ; insertion, into the first phalanx of the great toe, on its outer side; action, adducts the great toe; nerve, the ex- ternal plantar. Fourth Set. Dorsal Inteeossei. — Four in number. Origin, by two heads from the adjoining bases of the metatarsal bones; inser- tion, into the bases of the first phalanges of the second, third, and fourth toes, the second having two, one on either side, and the third and fourth on their outer side; action, abduct the toes from the middle line of the second toe; nerve, external plantar. Plantae Inteeossei. — Three in number. Origin, from the bases of inner side of the third, fourth, and fifth metatarsal bones ; insertion, into the bases of the first phalanges on the same side ; action, adduct the toes toward the middle line of the sec- ond toe; nerve, the external plantar. The Fascias. The fascias are strong, fibrous (fibro-areolar) investments protecting and binding together the muscles, and in places form- ing ligaments and intermuscular septa. They consist for the most part of two layers, a superficial and a deep. Fascias of the Head and Face. — The superficial fascia of the head and face is everywhere intimately connected to the skin, except over the temporal region, where it forms a distinct lamina, inclosing the superficial temporal vessels and auricular muscles. THE MUSCULAR SYSTEM. 167 The deep temporal fascia is a dense, fibrous membrane, at- tached to the margins of the temporal fossa and zygoma, and from which in part the temporal muscle arises. Fascias of the Neck. — The superficial fascia is continuous below with that covering the pectoral muscles and deltoid, and blends above with the superficial facial muscles and fascia. The deep fascia adheres below to the clavicle and sternum, and is continuous with the thoracic fascia. As it ascends the neck it ensheathes the great vessels of the neck, sends processes to the larynx, trachea, pharynx, and oesophagus, and above en- sheathes the submaxillary and parotid glands, and forms the stylo-maxillary ligament. Fascias of the Teunk. — The superficial fascia may con- sist of two or more layers in places. It is continuous above with the fascias of the neck and upper extremity, and below with the thigh. Over the pectoral region the mammary glands are inclosed between its layers; below the level of the umbilicus it again divides into two layers, the superficial of which is con- tinuous with the superficial layer of the superficial fascia in the thigh, while the deep layer is attached to the crest of the ilium and Poupart's ligament. The deep fascia forms intermuscular septa and aponeuroses. The lumbar fascia divides into three layers, inclosing the quadratus lumborum, multifldus spinse, and erector spinas mus- cles, and giving attachment to the internal oblique and trans- versalis muscles. Its anterior and middle layers are attached to the transverse processes, and its posterior layer to the spinous processes. Above its anterior layer is attached to the lower bor- der of the last rib, forming the ligamentum arcuatum externum. (The ligamentum arcuatum internum extends from the body of the first or second lumbar vertebra to the transverse process, inclosing the psoas magnus muscle.) Abdominal and Pelvic Fascia. — The transversalis fascia is a thin aponeurosis between the peritoneum and transversalis muscle. It is attached below to the pubes, pectineal line, Pou- part's ligament, and the femoral vessels, and becomes continuous with the iliac and pelvic fascia. The pelvic fascia lines the pelvic cavity throughout, attached to the symphysis pubis, mar- gin of obturator foramen, and the sacriun, and becomes con- tinuous with the iliac and transversalis fascias. Below it forms the arcus tendince, for the origin of muscles and the division into the recto-vesical and obturator fascias (vide Perineum) . The iliac fascia is a thin, fibrous membrane investing the psoas and iliacus muscles. Above it is attached to the ligamen- tum arcuatum internum, laterally with the bodies of the verte- 168 HUMAN ANATOMY. brae and sacrum, and below it is attached to Poupart's ligament, the femoral vessels, pectineal eminence, and the capsule of the hip-joint. Upper Exteemity. — The superficial fascia is a thin mem- brane, inclosing the parts throughout, and adherent to the deep fascia beneath. The deep fascia is very dense and strong. It gives oS numerous intermuscular septa, and is attached to the olecranon and back part of ulna, and becomes continuous below with the anterior and posterior annular ligaments of the wrist-joint, both of which it forms. The anterior annular ligament is attached to the ridge of the trapezium, the tuberosity of the scaphoid, the unciform proc- ess of the unciform, and the pisiform bone, forming a strong, fibrous arch, under which the flexor tendons of the fingers pass. The posterior annular ligament is attached to the radius, ulna, cuneiform, and pisiform bones, and the palmar fascia, forming a fibrous arch for the passage of the extensors of the fingers. The palmar fascia is an exceedingly dense, triangular mass of adipose and fibrous tissue, attached behind to the anterior annular ligament, and in front divides into four fasciculi, each of which subdivides into two, inserted into the lateral ligaments of the metacarpo-phalangeal articulations. These digitations allow the passage of the fiexor digitorum tendons, the digital vessels, and nerves. It is attached above to the skin, and later- ally gives off vertical septa, which separate the middle from the lateral groups of palmar muscles. FASOIAS OF THE LOWER EXTREMITY. The superficial fascia resembles that in other localities, except that, in the front of the thigh, it consists of two or more layers between which are found the superficial vessels and nerves and lymphatic glands. The under layer is attached to the mar- gins of the saphenous opening, where it is perforated by numer- ous lymphatic vessels and blood-vessels; hence its name, cribri- form fascia. The deep fascia of the thigh, or fascia lata, is a dense fibrous aponeurosis attached to the pubes, Poupart's ligament, crest of the ilium, sacrum, and coccyx, inclosing the gluteus maximus and tensor vaginse femoris between its layers, attached to the linea aspera, and below to all the bony points about the knee-joint, being attached to the head of the fibula and tuber- osities of the tibia by the ilio-tibial band. THE MUSCULAR SYSTEM. 169 The saphenovs opening is a large opening in the fascia lata at its upper and inner part closed by the cribriform fascia. Through it pass the internal or long saphenous vein, the super- ficial epigastric and external pudic arteries, and lymphatics. It is formed by the iliac and pubic portions of the fascia lata. The iliac portion becomes continuous at the pectineal line with Gimbemat's ligament, and its free border forms the falciform harder {vide Femoral Hernia). The pubic portion is continuous behind the femoral vessels with the femoral sheath and the iliac and psoas fascia. At the lower border of the saphenous opening it is continuous with the iliac portion of the fascia lata. The deep fascia of the leg is attached above to the bony points about the knee, continuous with the fascia lata, investe the leg completely, except the inner surface of the tibia, sends in a deep transverse fascia between the superficial and deep muscles on the posterior aspect of the leg, and below becomes continuous with the annular ligament of the ankle-joint. The annular ligament of the ankle-joint consists of three portions — ^the internal, external, and anterior. It is attached to the bony points about the ankle-joint, the external and in- ternal malleoli, surfaces of the os calcis, and allows the passage of the muscles to the foot. The dorsal aponeurosis of the foot covers the back of the foot, uniting with the plantar fascia at the sides, and terminat- ing anteriorly at the heads of the metatarsal bones. The plantar aponeurosis consists of three portions, a cen- tral and two lateral portions. The middle commences at the inner tubercle of the os calcis and proceeds, gradually becoming broader, to the heads of the metatarsal bones, where it divides into four branches, each of which nearly surrounds the corresponding flexor tendon to which it gives passage, and is inserted into the edges of the dorsal expansion of the first phalanges. The inner portion arises from the internal annular ligament, covers the abductor pollicis muscle, and joins the dorsal aponeu- rosis internally and the internal septum externally. The outer portion, much stronger, arises from the os calcis, covers the abductor minimi digiti muscle, and joins the external plantar septum internally and dorsal aponeurosis externally, and is firmly attached to the base of the fifth metatarsal bone. Numerous septa pass from the upper surface of the plantar fascia between the plantar muscles and tendons to be inserted into the metatarsal and tarsal bones. THE HEART AND VASCULAR SYSTEM. The pericardium is a conical, membranous, closed sac, surrounding the heart and the roots of the great vessels. It occupies the' greater part of the middle mediastinum, its apes upward, its base downward and attached to the central tendon of the diaphragm. It is a fibro-serous membrane, consisting of an outer fibrous coat and an inner serous coat. The latter is composed of a visceral layer reflected over the heart and vessels, and a parietal layer lining the inner surface of the fibrous sac. It secretes a thin, serous fluid. The fibrous coat becomes continuous above with the deep layer of the cervical fascia, being prolonged 'upward on the outer surfaces of all the great vessels, except, the inferior vena cava. The Heart. The heart is a hollow, muscular organ, placed obliquely within the chest, and inclosed within the pericardium. Its base is directed upward and backward, opposite the fifth to eighth dorsal vertebrae ; its apex downward to the left, opposite the interspace between the fifth and sixth costal cartilages. Its weight is about ten to twelve ounces in male, eight to ten in female; length five inches, breadth three and a half inches, thickness two and a half inches. It is divided into four parts : — Eight auricle. Left auricle, Right ventricle, Left ventricle, an auricle and ventricle on each side. This division is indicated on both surfaces of the organ by two grooves crossing each other and named, from theii- position, the longitudinal the inter- ventricular, the transverse the atericulo-ventricular grooves. Eight Auricle. — The right auricle, consists of two por- tions — a sinus or atrium and an appendix awriculcB, which pro- jects to the right side of the origin of the aorta. The walls at the fore and outer part and in the auricular appendage are thick- ened by musculi pectinati. The muscular tissue of the auricles is connected with that of the ventricles by a small band of fibres, the auriculo-ventriaular fasciculus, or '^bundle of His." This smooth ridge is known inside the auricle as the crista terminaUs of His, and outside the groove is called' the sulcus terminalis of His. Its interior presents the following parts : — Openings of the superior and inferior venw cava; neither of which are protected by valves ; (170) THE HEART AND VASCULAR SYSTEM. 171 Opening of coronary sinus, protected by a valve of two un- ■equal segments — ^the coronary valve ; Foramina Thebesii, minute foramina returning the blood from the heart-muscle; Eustachian valve, the remains of a foetal structure, extend- ing from the right of the orifice of the inferior vena cava to the outer border of the oval foramen; Fossa ovalis, the obliterated foramen ovale of foetal life; Annulus ovalis, the oval margin of the preceding structure ; Tubercle of Lower, a small rudimental projection on the right wall, directing the blood toward the auriculo-ventricular ■opening : FIG. 104. Right side of heart: a, apex; b, right ventricle; d, pulmonary artery; e, t, chordae tendinae; p, aorta; 3, superior vena cava; 5, Interior vena cava. Musculi pectinati, elevated muscular columns before re- ferred to; Auriculo-ventricular opening, an oval aperture about one inch in diameter, surrounded by a fibrous ring, and protected by the tricuspid valve. The Left Aueicle. — The left auricle is smaller, and its walls somewhat thicker, than the right, and consists of a cuboidal sinus and an elongated appendix auriculae. Its interior surface is smooth, except the auricular appendage, which is provided with musculi pectinati, and presents the following openings : — Pulmonary veins, four in nupiber, two on either side; they are without valves; 172 HUMAN ANATOMY. Auriculo-ventricular opening^ an oval aperture, rather smaller than the right, and protected by the bicuspid or mitral valve. The Eight Venteicle. — The right ventricle is triangular, with thick walls (one-quarter inch), and occupies the anterior part of the organ. Its capacity is about three fluid ounces. It presents the following: — Infundibvlum, or conus arteriosus, a conical pouch, from which the pulmonary artery arises; GolumncB carnew, muscular columns projecting from the inner surface, three or four of which (musculi papillares) give attachment to the chordae tendinese; ChordcB tendinem, delicate tendinous cords, connecting the margins and central portions of the tricuspid valve with the cojumnse carnese; Tricuspid valve, consists of three triangular duplications of endocardium, strengthened by fibrous tissue; they are attached by their bases to the auriculo-ventricular orifice, and their free margins give attachment to the chordae tendinese; Semilunar valves, consist of three semicircular folds, guard- ing the orifice of the pulmonary artery; their free margins are thickened by nodules, the corpora Arantiij Opening of the pulmonary artery, at the apex of the conus arteriosus, is circular in outline and • protected by the semi- lunar valves; Sinuses of Valsalva, three pouches, situated one behind each semilunar valve. The Left Ventricle. — The left ventricle is longer, more conical, and its walls three times as thick as the right. It forms the apex and most of the posterior portion of the heart. It presents the following parts in its interior : — ColumncB carnece, and ChordcB tendinew, much the same as the right; Auriculo-ventricular opening, a little smaller than the corre- sponding orifice on the right side, similarly formed, and pro- tected by the mitral valve; The mitral valve consists of two unequal segments, attached the same as the tricuspid, and also affording attachment to the chordae tendinese; Aortic opening, a circular opening to the right of the auriculo-ventricular opening ; The semi-lunar valves, three in number, surround the aorta, and are la:rger and stronger than those on the right side; Sinus aortici (sinuses of Valsalva) are depressions situated behind each valve. THE HEART AND VASCULAR SYSTEM. 173 Stedctdke of the Heart. — The heart is composed of finely striated muscular walls, with external serous covering from the pericardium and internal serous lining the endocardium^ the latter continuous with that of blood-vessels. The endocardium is a thin, translucent membrane, consisting of endothelium and a fine basement membrane, beneath which is a fibro-elastic layer. The doublings of these layers constitute the valves. The tricuspid and mitral valves, as well as the aortic and pulmonary, are surrounded by a fibro-elastic ring, which furnishes a "punctum fixum" for the various muscles of the heart. The muscular structure consists of an intricate interlace- ment of fibrous bands. Of these there are two groups — ^those of the auricles and those of the ventricles. The former consist of a superficial transverse set and the internal or deep set, of which there are the looped and the circular. The fibres of the ventricles consist of seven layers in a gen- eral oblique and circular manner, and terminating in a whorl or vortex at the apex, some of the fibres terminating in the columnse earner, musculi pectinati, while others ascend, form- ing in their course a twisted loop like 8. The arteries of the heart are the right and left coronary. The veins are the an- terior or great, middle or posterior cardiac, the left auricular, the right auricular, and venae Thebesii. The lymphatics end in thoracic and right lymphatic ducts. The nerves are derived from the cardiac plexus of pneumogastric, spinal, and great sympathetic. The Vasculae System. The vascular system consists of four sets of vessels — arteries, capillaries, veins, and lymphatics. Anastomoses are common. The vessels that convey blood to and from the tissues of the body generally constitute the general system. Those that carry blood to and from the lungs form the pulmonary system. The vessels passing to the liver form the portal system. The aetekies, for the most part, are composed of three coats : — Tunica adventitial or external coat, consists of fibrous tissue, thinnest on the largest trunks, and disappears in those which merge into capillaries. The tunica media, or middle coat, is thickest in the large trunks, and consists mainly of elastic tissue, together with some unstriated muscle and some connective tissue; in smallest ar- teries it consists alone of muscular tissue. The elastic tissue 174 HUMAN ANATOMY. in layers forms nets, constituting so-called ■perforated or fenes- trated membranes. The tunica intima, or internal coat, thinnest, most trans- parent, and elastic, is composed of lining endothelium, basement membrane, and layers of elastic tissue. It becomes continuous with capillaries. The -walls of larger arteries are supplied by blood-vessels from neighboring arteries. Nutrient arteries form a net-work of capillaries, and returning veins empty into con- tiguous veins. The nerves are both medullated and non- meduUated and chiefly pass to the muscle-fibres. Arteries dilate and contract with the action of the heart, con- stituting the pulse. The Veins. — Larger, more numerous, and more capacious than the arteries. With one exception (the portal vein) they convey blood to the heart. The large veins emptying into the heart are: — Four pulmonary; Superior and inferior venae cavae; Coronary veins from walls of the heart through the coronary sinus. The larger arteries have each one companion vein, the medium arteries two. In some positions they form venous plex- uses, corresponding to arterial anastomoses. Many large veins have valves, usually in pairs, attached by convex border. There are, however, no valves in the pulmonary, superior and inferior cavae, azygos, portal vein and branches, hepatic, renal, uterine, and spinal, and most of those of head or neck. In certain membranes and organs channels exist, lined by internal coat of blood-vessel, termed venous sinuses, as iu the dura mater, bones, and uterus. The coats of the veins are similar to those of the arteries, and differ chiefly in the weakness of the middle or muscular coat, which allows them to remain open. The external fibrous coat has also longitudinal, unstriated muscular tissue in veins of abdominal cavity : — The middle J unstriated muscular and fibrous tissue; The internal^ no fenestrated membrane. The walls have nutritive vessels, and are well supplied with non-medullated and medullated nerves, which pass to the muscu- lar coat. Both arteries and veins are insensitive in health. The capillaries communicate with the terminations of the arteries and veins, and do not communicate with tissue proper, which is supplied by imbibition. They vary in size in different THE HfiART AXD VASCULAR SYSTEil. 175 organs from ^/gooo to ^Aooo inch in diameter, but all permit the passage of blood-corpuscles. Their form is dependent on the form of tissue supplied. CoMPOSiTiox OF THE Blood. — Blood within the vessels is a perfect fluid, of alkaline reaction, saline taste, average specific gravity 1060, and feeble, peculiar odor. Within the arterial vessels it is bright-red color, becoming in the veins (particularly the venae portse and pulmonary arteries) a deep maroon or red- dish-black color. It consists about one-twelfth or one-fourteenth of the body-weight, and consists of the liquor sanguinis and the cor- puscles, red, white, etc. The red corpuscles or erythrocytes are elastic, of specific gravity 1088, nonnucleated, and average about one three- thousandth line in diameter. A cubic millimetre of blood con- tains, in the male, 5,000,000 red blood-cells, and, in the female, 4,500,000. On evaporation they become stellate, and swell up with water. They vary much in size and shape in different animals, and contain haemoglobin, which carries oxygen to the tissues and COg to the lungs. The colorless or leucocytes are free nucleated protoplasmic masses, capable of amoeboid movement. They contain fat- corpuscles, myosin, cholesterin, protagon, glycogen, and nuclein. There is about 1 white corpuscle to every 600 red corpuscles, or in 1 cubic millimetre of blood there are present about from 5,000 to 10,000 white corpuscles, average 7,500. According to Bhrlich and others, by means of a "tri-acid stain," fine varieties of colorless cells may be distinguished as follows : — 1. Small lymphocytes, 2. Large lymphocytes or mononuclear leucocytes, 3. Transitional leucocytes, 4. Polymorphonuclear leucocytes, and 5. Eosinophiles. The liquor sanguinis is a pale, amber-colored fluid, holding the corpuscles in suspension, and very prone to coagulate. The coagulation of, the blood results in the formation of a clot, or crassamentum, and the serum, in the following manner : — Fluid blood . Corpuscles. \ "'^""^ corpuscles. ■] '^ ( Red corpuscles. mi Plasma or 1 ,,., . \ *^''°*- Liquor ^'^""^ J Sanguinis J Serum. Serum. 176 HUMAN ANATOMY. Fig. 105. Diagram of the (cetal circulation: 1, umbtUoal oorfl, with veins and arteries; 2, placenta; 3, divisions of umbilical vein; 4, hepatic branches; 6, ductus venosus; 6, Inferior vena cava; 7, portal vein; 8, right auricle; 9, left auricle; 10, left ventricle; 11, arch of aorta; 14, superior vena cava; 15, right ventricle; 16, pulmonary artery; 17, ductus arteriosus; 18, descending aorta; 19, umbilical arteries. THE HEART AND VASCULAR SYSTEM. I77 THE CIKCULATION OF THE BLOOD. Fcetal Circulation. — The fresh arterial blood returning from the placenta through the umbilical vein enters the foetus at the umbilicus, passes along the suspensory ligament to the under surface of the liver, where a portion passes directly into the inferior vena cava by the junction of the ductus venosus with the left hepatic vein, a portion enters the liver and reaches the inferior vena cava through the hepatic veins, and the larger por- tion mixes with the portal venous blood before reaching the vena cava through the hepatic veins. It ascends in the inferior vena cava along with the venous blood from the trunk and lower extremities to enter the right auricle of the heart (Fig. 105). Here it is directed by the Eustachian valve through the foramen ovale into the left auricle, into the left ventricle, and so through the aorta, chiefly to the head and upper extremities, a small por- tion only entering the descending aorta. It is returned by the superior vena cava to the right auricle, passes over the Eustachian valve to the right ventricle, and so through the pulmonary artery toward the lungs, but (the lungs being almost impervious) most of it passes through the ductus arteriosus into the descending aorta (mixing with the small quantity from the left ventricle before mentioned) to supply the lower extremities, abdominal and pelvic viscera, and as venous blood to return through the umbilical arteries to the placenta. Circulation After Birth and in Adult. — The dark venous blood from the entire body is received through the superior and inferior venffi cavje and coronary sinus into the right auricle, from whence it passes into the right ventricle, to be sent through the pulmonary artery into the lungs. Here it becomes oxy- genated, and as bright arterial blood it returns to tlie left auricle by the pulmonary veins, and passes into the left ventricle, from whence it is distributed through the aorta and its branches to the entire body. AOKTA. This is the main trunk of the systemic arteries. Commenc- ing at the upper part of the left ventricle, it ascends for a short distance, arches backward over the right pulmonary artery, the root of the left lung, to the left side of the body of the fourth dorsal vertebra, from where it passes downward through the dia- phragm and becomes the abdominal aorta. It then descends to the lower border of the fourth lumbar vertebra, where it termi- nates by dividing into the left and right common iliac arteries. 178 HUMAN ANATOMY. In its course it is divided into the arch, the thoracic aorta, and the abdominal aorta, the arch being subdivided into the ascend- ing, transverse, and descending portion. Relations. — The ascending portion of the arch is in relation on the right side with the superior cava and right auricle; on the left side with the pulmonary artery; in front, with the pulmonary artery, the pericardium, the right appendix auriculae, and the thymus gland; and liehind, with the root of the right lung and the right pulmonary vessels. The transverse portion of the arch is in relation in front with the left lung and pleura, the left pnemnogastric and phrenic nerves, the superficial cardiac nerves, the left superior intercostal vein, and the thymus gland; behind, with the trachea, the oesophagus, the thoracic duct, the deep cardiac plexus, and the left recurrent nerve; above, with the arteria innominata, the left innominate vein, the left subclavian, and the left carotid; and below, with the left bronchus, the bifurcation of the pulmonary artery, the left recurrent nerve, and the remains of the ductus arteriosus. The descending portion of the arch is in relation in front with the root of the left lung and the pleura; behind, with the left side of the body of the fifth dorsal vertebra; on the right side with the thoracic duct and cesophagus; and on the left side with the pleura. The branches of the aorta are, from the arch: — ■ Two coronary, Left common carotid. Innominate, Left subclavian. The coeonaet aeteribs supply the heart and are two in number — the right and the left. The right arises from the aorta, above the right semi-lunar valve, and passes downward in a groove between the right auricle and ventricle on its posterior surface, dividing into two branches, one of which anastomoses with the left coronary. The left coronary arises above the left semi-lunar valve and descends in the anterior interventricular groove, where it divides into two branches, one of which anas- tomoses with the right coronary. Arteeia Innominata. — This is the largest branch from the arch of the aorta. It ascends to the upper border of the right sterno-clavicular articulation, where it divides into the right common carotid and the right subclavian arteries. It is a short vessel, from one and a half to two inches long. Relations. — It is in relation in front with the sternum, the sterno- hyoid and sterno-thyroid muscles, the left innominate and right inferior thyroid veins, the thymus gland, and the cardiac branch from the right pneumogastrio ; behind, with the trachea; on the right side, with the pleura and right vena innominata and right pneumogastrio nerve; on the left side, with the left carotid and the thymus gland. The Common Carotid Arteries. — The left common ca- rotid, arising directly from the arch of the aorta, is longer and ascends more obliquely and is more deeply placed than the right. THE HEART AXD VASCULAR SYSTEM. 179 From the stemo-clavicular articulation to their division at the upper border of the thyroid cartilage the common carotid on either side pursues the same course. At the lower part of the neck the two vessels are separated by a very short interval, which is occupied by the trachea, but at the upper part they diverge widely, the larynx, pharynx, thyroid body, and several muscles being interposed between them. It ascends the neck in a direc- tion indicated by a line drawn from the stemo-clavicular articula- tion to midway between the mastoid process and the angle of the lower jaw. Relations. — It is in relation in front with the sterno-mastoid, sterno-thyroid, sterno-hyoid, and omo-hyoid muscles, the anterior jugular and the superior and middle thyroid veins, the sterno-hyoid artery, and the descendens and communicans noni nerves; behind it rests upon the longua colli and rectus capitis anticus major muscles and is in relation with the sympathetic nerve, the recurrent laryngeal nerve, and the inferior thyroid artery; on the outer side, with the pneumogastric and the internal jugular vein; to the inner side, the larynx, trachea, pharynx, thyroid gland, inferior thyroid artery, and the recurrent larygeal nerve. THE EXTERNAL CAROTID AETEET. From its commencement at the superior border of the thy- roid cartilage it passes upward and forward and then backward, beneath the anterior margin of the sterno-mastoid muscle, in a direction indicated by an imaginary line drawn from the supe- rior border of the thyroid cartilage to a point midway between the meatus and the neck of the condyle of the lower Jaw. It ascends through the space known as the superior triangle of the necJc. Relations. — It is in relation in front with the parotid gland and the facial nerve and temporo-maxilJary passing through it, covered by the skin, superficial fascia, platysma, and deep fascia, and Is crossed by the digastric and stylo-hyoid muscles, hypoglossal nerve, lingual and facial veins; behind it is separated from the internal carotid by the parotid gland, stylo-glossus and stylo-pharyngeus muscles, and the glosso- pharyngeal nerve, and at its lower part the superior laryngeal nerve ascends behind it; to its inner side it is in relation with the pharynx, hyoid bone, superior laryngeal nerve, and the ramus of the lower jaw, from which it is separated by the parotid gland. The branches of the external carotid are eight : — 1. Superior thyroid, 5. Posterior auricular, 2. Lingual, 6. Ascending pharyngeal, 3. Facial, 7. Temporal, 4. Occipital, 8. Internal maxillary. 180 HUMAN ANATOMY. 1. Sdpeeior thyeoid arises below the great cornu of the hyoid bone, passes beneath the sterno-hyoid, omo-hyoid, and sterno-thyroid muscles to the upper part of the thyroid gland, where its terminal branches anastomose with its fellow of the opposite side. Its branches are : — a. Hyoid, to the lower border of the hyoid bone; 6. Superficial descending branch, crosses the common carotid artery to supply the skin and sterno-mastoid muscle; c. Superior laryngeal, passes beneath the thyro-hyoid muscle with the superior laryngeal nerve; (2, Crico-thyroid, crosses the crico-thyroid membrane. 2. The LINGUAL ARTERY arises from the anterior part of the external carotid, between the superior thyroid and the facial. It passes inward and upward to the greater cornu of the hyoid bone, and, descending, reaches the under surface of the tongue and runs along its under surface as far as its tip, where it termi- nates as the ranine. In its course it passes through a triangle bounded on two sides below by the two bellies of the digastric, and above by the hypoglossal nerve, passing under the hyoglossus muscle. Its branches are: — Hyoid, to the hyoid bone; Dorsalis linguae, supplies the epiglottis, soft palate, tonsil, and the mucous membrane of tongue ; Sublingual, supplies mylo-hyoid and genio-hyoglossus muscles and sublingual gland; Eanine, the terminal branch, ends at the tip of the tongue. 3. The FACIAL ARTERY arises from the anterior portion of the external carotid artery above the lingual. It ascends for- ward through the submaxillary gland, crosses the lower jaw at the anterior border of the masseter muscle, and crosses the cheek to the angle of the mouth and the side of the nose, where it terminates as the angular artery. Its branches are in the neck : — Inferior or ascending palatine, supplies the soft palate, palatine glands, tonsil, and Eustachian tube; Tonsillar, supplies the root of the tongue and tonsil; Submaxillary, supplies the submaxillary gland and neighboring lymphatics, muscles, and skin; Submental, supplies the chin, lower lip, and neighboring struc- tures ; Muscular, to the buccinator, masseter, and internal pterygoid mus- cles. THE HEART AND VASCULAR SYSTEM. In the face : — 181 Muscular, to the same muscles given above; Inferior labial, to the skin of the lower lip; Inferior coronary, to the mucous membrane, muscles, and glands of the lower lip; Superior coronary, to the structures of the upper lip and nasal septum; Fig. 106. Arteries of face and head: 1, common carotid; 2, Internal carotid; 3, external carotid; 4, occipital; 6, superior tliyrold; 7, lingual; 9, facial; 10, temporal: 11, submental; 12, transverse facial; 13, in- ferior labial; 15, 17, inferior and superior coronary; 19, lateral nasal; 21, angular. Lateralis naai, to the skin of the nostrils, anastomoses with the nasal branch of ophthalmic; Angular, supplies lachrymal sac and skin. All of these branches anastomose freely with the neighboring arteries. 182 HUMAN ANATOMY. 4. The OCCIPITAL artekt arises from the posterior portion of the external carotid, about the lower border of the digastric muscle. It passes beneath the stylo-hyoid muscle, the digastric muscle, and part of the parotid gland, and, ascending, grooves the internal surface of the mastoid portion of the temporal bone and distributes itself over the occiput as high as the vertex. Its branches are : — Muscular, to splenius, digastric, stylo-hyoid, etc.; Sterno-mastoid, crosses the hypoglossal to the sterno-mastoid muscle ; Auricular, is distributed to the back part of the concha; Meningeal, passes through the foramen lacerum posterius to the dura mater; Arteria princeps cervicis. Its superficial branch anastomoses with the superficial cervical, and its deep branch with the deep cervical branch from the superior intercostal. Cranial branches are distributed to the scalp over the occiput. 5. The POSTERiOE auriculae artery arises from the pos- terior portion of the external carotid, on a level with the apex of the styloid process. It ascends beneath the parotid gland to a point between the mastoid process and the cartilage of the ear, where it divides into two branches, one going to the ear and the other to the occiput. Its branches are: — Stylo-mastoid, supplies the mastoid cells, tympanum, and semi- circular canals, entering the cranium through the stylo-mastoid foramen; Auricular, supplies the cartilage of the ear; Muscular, supply the sterno-mastoid, digastric, and stylo-hyoid muscles ; Glandular, to parotid gland. 6. The ASCENDING PHARYNGEAL artery arises about the com- mencement of the external carotid artery and ascends to the base of the skull upon the rectus capitis anticus major. Its branches are: — Pharyngeal, supplies the three constrictors of the pharynx and the stylo-pharyngeus muscles; External, to the neighboring muscles, glands, and hypoglossal and pneumogastric nerves; Meningeal, enter the skull through the posterior and middle lacer- ated foramina and the anterior condyloid foramen. 7. The TEMPORAL artery is the smaller of the two terminal branches. It arises in the parotid gland, crosses the root of the zygoma, ascends forward a couple of inches, and divides into THE HEART AND VASCULAR SYSTEM. 183 the anterior temporal and the posterior temporal. Its branches are: — skin; Transverse facial, supplies the masseter muscle, parotid gland, and Middle temporal, crosses the face to supply the temporal muscle and fascia; Anterior auricular, supplies the lobule, external meatus, and front part of pinna. 8. The INTERNAL MAXILLARY is the larger of the two termi- nal branches of the external carotid. It arises in the parotid gland about the level of the lower part of the lobe of the ear. Fig. 107. Internal maxillary artery; A, third portion; B, second portion; C, first portion. passing close to the inner side of the neck of the condyle of the lower jaw to be distributed to the deep structures of the face. Its course is divided into three portions : the maxillary portion, passing inward and forward between the internal lateral liga- ment and the ramus of the jaw; the pterygoid portion, passing upward and forward upon the external pterygoid muscle; and the third part, or spheno-maxillary , enters the spheno-maxillary fossa. Its branches are, from the maxillary portion : — Tympanic (anterior), supplies the tympanum through the Gla- serian fissure, forming a capillary plexus on the tympanic membrane; Middle meningeal, enters the cranium through the foramen spinosum, supplies the dura mater, and gives oS a petrosal branch to facial nerve in hiatus Fallopii; 184 HU:\rA^' ANATOMY. Small meningeal, or meningea parva, enters foramen oA'.nle to supply dura mater and Gasserian ganglion; Inferior dental, accompanies the dental nerve through the inferior dental canal to the mental foramen, supplying the jawbone, teeth, chin, and mouth. The pterygoid portion : — Deep temporal, supply the temporal muscle; Pterygoid, to pterygoid muscles; Masseteric, to the masseteric muscle; Buccal, to the buccinator muscle. Splieno-maxillary portion : — Alveolar or posterior dental branch, supplies the upper molar and bicuspid teeth, antrum, and gums; its principal branch is the superior dental; Infraorbital, passes through infraorbital canal to face; Posterior or descending palatine, descends posterior palatine canal and runs forward to foramen of Stenson; Vidian, accompanies the Vidian nerve; Pterygo-palatine, supplies Eustachian tube and pharynx; Nasal, or spheno-palatine, descends through spheno-palatine fora- men to antrum, sphenoidal, and ethmoidal cells. THE UTTEENAL CAEOTID AETERT arises about the upper border of the thyroid cartilage from the bifurcation of the common carotid. It ascends in front of the transverse processes of the cervical vertebrae to the carotid fora- men in the petrous portion of the temporal bone to supply the anterior part of the brain, the eye- with its appendages, and send branches to the nose and forehead. It gives no branches until it passes through the carotid canal, vs^here, after piercing the dura mater at the anterior clinoid process, it divides into its terminal branches — ^the anterior and middle cerebral. Relations. — ^The internal carotid artery in the neck is in relation in front with the parotid gland, the glosso-pharyngeal nerve, the stylo- glossus and stylo-pharyngeal muscles, and is covered by the skin, super- ficial and deep fascia; behind it rests upon the rectus capitis anticus major and has resting upon it the superior laryngeal nerve and the sympathetic; internally it is in relation with the pharynx, tonsil, superior laryngeal nerve, and ascending pharyngeal artery; externally it is in relation with the internal jugular vein and the pneumogastric nerve. Its branches are, from the petrous portion: — Tympanic (internal or deep), supplies tympanum. From the cavernous portion :— Arteriw receptaculi, supply the Gasserian ganglion and pituitary body; POSTEROMEDIAN fiANC.LION FOSI-KHIOR TEMI-ORAI, ANTEKIOR Tt:Mr'OR,^L- SUPERFICrAL TEMWJRAI, ANTERIOR AURICULAR MIDDLE TEMITJRAL ORBITAL TRANSVERSE FACIAL ANTERIOR CHOROID TRANSVERSE PUS.TERIOR AURICULAR INTERNAL MAXILLARV ARTERIA PKINCEPS CERVIC OCCIPITAL SUP OR. PRINCEPS CERVICIS- DEEP BR, PBINCEPS rERVICIS' ASCENDING CERVIf Al DEEP CERVICAL anterioh communicating anterior cerebral^ .middle cerebral - ophthalbiic . temi'oral . POST communicatinc; ■POST CEREBRAL - 3UPERI0D CEREBELLAR, SUP STERNO-MASTOID. BASILAR- A.NT. INFERIOR CEREBELLAR - L, INT. CAROTID . FACIAL _ R CORONARY fNF VENA CAVA THOE-ACJC AOKTA Fro. 108. The arteries of the head and neck. THE HEART AND VASCULAR SYSTEM. 185 Anterior meningeal, a branch from the preceding vessel to the dura. Ophthalmic arises about the position of the anterior elinoid process and passes through the optic foramen to the outer side of the optic nerve to reach the orbit. At the inner angle of the eye it divides into its two termiQal branches — the frontal and the nasal. It supplies the muscles and the globe of the eye, and its branches may be divided into two groups — ^the orbital and the ocular. Orbital group: — Lachrymal, supplies the lachrymal gland and gives off a malar branch which passes through the malar bone to the temporal fossa; Supraorbital, passes out through the supraorbital foramen to supply the surrounding structures; Posterior ethmoidal, descends through the posterior ethmoidal foramen to the cells and adjacent parts; Anterior ethmoidal, through the anterior ethmoidal foramen to the nose; Palpebral, supply the eyelids; Frontal, supplies the skin and muscles of the forehead. Nasal, supplies the lachrymal sac and bridge of the nose. Ocular group: — Muscular, supply the muscles of the eye; Anterior ciliary, supply the iris, piercing the sclerotic near the cornea ; Short ciliary, supply the choroid and ciliary processes, piercing the sclerotic near the optic nerve; Long ciliary, enter with the short ciliary, but run forward to the iris; Arteria centralis retinae, supplies the optic nerve and the retina. From the cerebral portion: — Anterior cerebral, supplies the anterior part of the cerebrum, the optic and olfactory nerves, and communicates with its fellow by the anterior communicating; Middle cerebral, or Sylvian artery, ascends in the Sylvian fissure, and supplies the middle lobes of the cerebrum; Posterior communicating, runs backward to join the posterior cerebral ; Anterior choroid, supplies the choroid plexus. THE CIKCLE OF WILLIS is an anastomotic circle formed at the base of the brain about the pituitary body, between the branches of the internal carotid and the vertebral arteries, for the purpose of equalizing the cere- bral circulation. The anterior cerebrals, from the internal ca- rotids, pass forward and are united in front by a short trunk, the 186 HUMAN ANATOMY. anterior comnmnicating. The posterior comnmnicating imites the internal carotid with the posterior cerebral, the latter being the terminal branches of the basilar, which in its turn is formed by the union of the two vertebrals. THE SUBCLAVIAN ARTEET on the right side arises from the arteria innominata, behind the right sterno-clavicular articulation, and on the left side directly from the arch of the aorta. They pass outward to the outer margin of the first rib, where they become the axillary, being divided in their course by the scalenus anticus muscle into three portions — the first portion to the inner side of the muscle, second portion behind it, and the third portion between its outer margin and the lower border of the first rib. The vessels differ only in their first portions, the left ascending more vertically. Relation. — ^The first portion of the right subclavian is in relation in front to the internal jugular and vertebral veins, the sternohyoid and sterno-thyroid and the clavicular portion of the sterno-mastoid muscles, and the pneumogaatric, phrenic, and cardiac nerves; behind it is in relation with the longus colli muscle, the transverse process of the first dorsal or seventh cervical vertebrae, the sympathetic nerve, and the recurrent laryngeal nerve beneath with the pleura. The first portion of the left subclavian artery is in relation in front with the left internal jugular and innominate veins, the left carotid artery, the sterno-thyroid, sterno-hyoid, and sterno-mastoid muscles, the pleura and left lung, and the pneumogastric, phrenic, and cardiac nerves; behind, with the vertebral column and longus colli muscle, the oesophagus, and thoracic duct, and the inferior cervical ganglion of the sympathetic; on the inner side with the cesophagus, thoracic duct, and trachea, and on the outer side with the pleura. The second portion of the subclavian artery on either side is in relation with the subclavian vein, the scalenus anticus muscle, and the phrenic nerve; behind with the middle scalenus muscle and pleura; above by brachial plexus, and below by the pleura. The third portion of the subclavian artery on either side is in relation in front with the external jugular vein and its branches, the clavicle, subclavius muscle and suprascapular artery, the descending branches of the cervical plexus and the cervical fascia; behind with the scalenus raedius, above with the omo-hyoid muscle and the brachial plexus, and below with the first rib. Its branches are: — 1. Vertebral, 5. Transverse cervical, or 2. Thyroid axis, transversalis colli, 3. Inferior thyroid, 6. Internal mammary, 4. Suprascapiilar, 7. Superior intercostsd. These branches are all given off from the first portion, ex- cept the superior intercostal, which arises on the right side, from the second portion. mil. *tl, li.VLllAR PlOrTAUS Fi(.i. 109. The Jirtories a]' the uppi^r oxirciniiy. THE HEART AND VASCULAR SYSTEM. 187 1. The vertebral artery passes upward, enters the foramen in the transverse process of the sixth cervical vertebra, and ascends through the foramina until it reaches the atlas, when it runs backward, pierces the posterior occipito-atloid ligament and dura mater to enter the skull through the foramen magnum. After entering the cranium it passes forward to the lower border of the pons Varolii, where it unites with its fellow of the oppo- site side to form the basilar artery. Its branches are — Cervical branches : — Lateral spinal, supply the spinal cord and its membranes, entering the spinal canal through the intervertebral foramina; Muscular, supply the deep muscles of the neck. Cranial branches: — Posterior meningeal, supply the falx cerebelli; Anterior spinal, supplies the cord; uniting with branches from the inferior thyroid, intercostals, and lateral sacral, descend to the cauda equina ; Posterior spinal, descends in the same manner as the anterior; Posterior inferior cerebellar, supplies the inferior surface of the cerebellum and the choroid plexus of the fourth ventricle. The branches of the basilar artery are : — Transverse, to the pons Varolii and inferior surface of the cere- bellum ; Anterior inferior cerebellar, a branch of the transverse; Superior cerebellar, supplies pia mater, pineal gland, and velum interpositum ; Posterior cerebral, to posterior lobes of cerebrum and choroid plexus. 2. The thyroid axis consists of three branches — ^the inferior thyroid, the suprascapular, and the transversalis colli. 3. The inferior thyroid artery is distributed to the thyroid gland, its branches being : — Laryngeal, to muscles and mucous membrane of larynx; Tracheal, to the trachea, anastomosing with the bronchial arteries; OSsophageal, to oesophagus; Ascending cervical, supplies the spinal cord and membranes, ver- tebrae, and muscles of neighborhood; Muscular, to inferior constrictor of pharynx, scalenus anticus, and other muscles. 4. The suprascapular artery (transversalis humeri) passes beneath the posterior belly of the omo-hyoid to the upper border of the scapula, passing over the transverse ligament of the scapula to the supraspinous fossa, where it is distributed to the supra- spinatus, sterno-mastoid, and other muscles, and by means of the 188 HUMAN ANATOMY. supra-acroinial branch to the skin over the acromion, anastomos- ing with the posterior circumflex and acromial thoracic arteries. In about 50 per cent, of bodies this artery will be found to arise from the third portion of the subclavian. 5. The transversalis colli passes outward to the trapezius muscle, where it divides into The (a) superficial cervical, an important branch passing beneath the trapezius to anastomose with the superficial branch of the arteria princeps cervicis; And the (t) posterior scapular, the larger passing to and along the upper and posterior border of the scapula, beneath the levator anguli scapulae, rhomboidei, latissimus dorsi and trapezius, to anas- tomose with the subscapular, suprascapular, and posterior branches of the intercostals. 6. The internal mammary arises from the subclavian, oppo- site the thyroid axis, and descends upon the internal surfaces of the costal cartilages to between the sixth intercostal space, where it divides into the musculo-phrenic and the superior epi- gastric. Its branches are : — Comes nervi phrenici (superior phrenic), supplies diaphragm, pass- ing between the pleura and pericardium ; Mediastinal, to the anterior mediastinum; Pericardiac, to upper portion of pericardium; Sternal, to posterior surface of sternum; Anterior intercostal, to fifth or sixth upper intercostal spaces; Perforating, perforate the intercostal spaces to supply the mam- mary gland; Musculo-phrenic, supplies lower part of pericardium, diaphragm, and upper part of abdominal muscles; Superior epigastric, supplies the rectus muscle, and anastomoses with the deep epigastric from the external iliac. 7. The superior intercostal artery arises from the subclavian, behind the anterior scalenus muscle, and descends behind the pleura and anastomoses with the first aortic intercostal artery. It gives off branches in the intercostal spaces to the spinal cord and its membranes and the posterior spinal muscles. Its princi- pal branch is the deep cervical (profunda cervicis). THE AXILLARY ARTERY is the continuation of the subclavian, extending outward from the lower edge of the first rib to the lower margin of latissimus dorsi and teres major muscles, where it becomes the brachial. It is divided by the pcctoralis minor into three parts — the first portion above, the second portion behind, and tlie third portion THE HEART AND VASCULAR- SYSTEil. 189 below that muscle. It passes throiigli the space known as the axilla {vide Jiegions). R( hit ions. — First portion: The first portion of the axillary is in relation in front with the subclaviirs and pectoralis major muscles, costo-coracoid membrane, and acromial thoracic and cephalic veins, and the external anterior thoracic nerve; the axillary vein to its inner side, the brachial plexus to its outer side, and the posterior thoracic nerve and serratus magnua behind. Second portion: The relations of the second portion are much the same as the first, with the addition of the pectoralis minor in front and the division of the brachial plexus sur- rounding the vessel. Third portion: The third portion is in relation in front with the inner head of the median nerve and the pectoralis major muscle ; to the inner side with the axillary vein, internal cutaneous and ulnar nerves; to the outer side, the median and museulo-cutaneoua nerves and coraco-brachialis muscle; and, behind, the circumflex and musculo-spiral nerves and the subscapularis, latissimus dorsi, and teres major muscles. Its branches are: — Superior thoracic, supplies the pectoralis major and minor muscles ; Acromial thoracic, gives ofE three branches — acromial, tho- racic, and descending, supplying the pectoralis major, minor, serratus, and deltoid ; Long thoracic follows the lover border of the pectoralis minor to supply the serratus, pectoralis major and minor, and mammary glands ; Alar thoracic supplies the axillary glands; Subscapular, arises from the axillary artery about the level of the lower border of the subscapularis muscle. It passes backward and downward to the inferior dorsum of the scapula. Its branches anastomose with the posterior scapular and the suprascapular. They are: — Subscapular, supplies the subscapularis muscle; Dorsalis scapulae, is distributed to the infraspinous fossa, anasto- mosing with the posterior scapular and suprascapular arteries; Median branch, descends the axillary border of the scapula to supply teres muscles. Posterior circumflex, encircles the neck of the humerus to supply the shoulder- joint and the deltoid muscle; Anterior circumflex, descends beneath the short head of biceps and coraco-brachialis muscles to supply the shoulder-joint. THE BEAOHIAL ARTERY commences at the lower border of the latissimus dorsi and teres major muscles and descends on the front and inner aspect of the arm to half an inch below the middle of a line drawn through 190 HUMAN ANATOMY. the condyles of the humerus, where it terminates by dividing into the radial and ulnar artery. Relations. — ^It is covered in front by the skin, superficial, and deep fascia, and is crossed by the median nerve; behind it rests upon the triceps, brachialis anticus, and coraco-brachialis muscles ; And is in relation with the musculo-spiral nerve and the superior profunda artery; to the outer side above it is in relation with the biceps and coraco-brachialis muscles and the median nerve; and on the inner side with the internal cutaneous and ulnar nerve, and the basilic vein and median nerve below. At the bend of the elbow it is crossed by the bicipital fascia. Its branches are : — Superior profunda, accompanies the musculo-spiral nerve to supply the triceps, deltoid, and coraco-brachialis muscles; Nutrient artery, supplies the shaft of the humerus ; Inferior profunda, accompanies the ulnar nerve to supply the triceps muscle and structures about the internal condyle ; Anastomotica magna, forms an anastomosis on the lower posterior aspect of the arm, with branches from the inferior profunda and recurrent ulnar vessels. It supplies the triceps; Muscular, supply the muscles on the anterior aspect of the arm. THE EADIAL AKTERT descends from the bifurcation of the brachial below the bend of the elbow to the wrist, where it winds around the carpus beneath the extensor tendons of the thumb, passes between the two heads of the first dorsal interosseous muscle and into the palm of the hand, where it forms the deep palmar arch, anasto- mosing with the deep branch of the ulnar artery. Its branches are, in the forearm : — Radial recurrent, supplies the brachialis anticus and supinator longus and brevis; Muscular, supply the radial muscles; Superficialis volse, supplies the muscles of the thumb and anasto- moses with the ulnar to form the superficial palmar arch; Anterior carpal, supplies the wrist-joint. In the wrist: — Posterior carpal, supplies the wrist-joint, forms the posterioT carpal arch, and gives off the dorsal interosseous arteries; Metacarpal, is called the first dorsal interosseous; it supplies the index and middle finger; Dorsales pollicis, supply the back of the thumb; Dorsalis indicis, supplies the outer and dorsal side of the index finger. THE HEART AND VASCULAR SYSTEM. 191 In the hand : — Princeps poUicis, supplies the sides of the palmar aspect of the thumb; Eadialis indicis, supplies the radial side of index finger; Perforating, anastomose with dorsal interosseous arteries; Interosseous, anastomose with digital branches of superficial palmar arch. THE ULNAE AETEEY descends from the bifurcation of the brachial below the bend of the elbow to the ulnar border of the wrist, passing beneath all the superficial flexors excepting the flexor carpi ulnaris, crosses over the annular ligament at the radial side of pisiform bone, and enters the palm to form the superficial palmar arch, anastomosing with the superficialis volaa from the radial. Its branches are, in the forearm : — Anterior ulnar recurrent, supplies the pronator radii teres and brachialis anticus muscles; Posterior ulnar recurrent, supplies the joint and neighboring muscles, and anastomoses freely; Interosseous, passes backward to the upper border of the inter- osseous membrane, where it divides into Anterior branch, descends on the front of the interosseous membrane ; Posterior, descends on the posterior aspect of the foramen and gives off a recurrent interosseous branch; Muscular, to muscles on ulnar side of forearm. In the wrist: — Anterior carpal, supplies the front of wrist and anastomoses with the carpal branches of radial artery; Posterior carpal, passes beneath the tendon of flexor carpi ulnaris and forms posterior carpal branch. In the hand: — Deep, or communicating branch, passes between the flexor and abductor minimi digiti to form part of deep palmar arch; Digital, supply the ulnar side of the little finger and adjoining sides of the little, ring, middle, and Index fingers. THE DESCENDING AORTA. The thoracic aorta extends from the lower border of the fifth dorsal vertebra on the left side to about the last dorsal vertebra, where by passing through the aortic opening in the diaphragm it becomes the abdominal aorta. Relations. — It is in relation in front with the left bronchus, oesoph- agus, pericardium, and left pulmonary artery; behind it rests upon the 192 HUMAN ANATOMY. vertebral column and is in relation with the vena azygos minor ; on the right side it is in relation with the thoracic duct, the oesophagus at its upper part, and the vena azygos major; on the left side with the left lung, the pleura, and the oesophagus below. The branches of the thoracic aorta are : — • Pericardiac, supply the pericardium; Bronchial, two or three in number, supply the bronchi ; CEsophageal, supply the oesophagus; Posterior mediastinal, supply the neighboring pleura and lymphatic glands; Intercostal, ten on each side, supply the intercostal spaces. They divide into — anterior branch, to intercostal and pectoral muscles; posterior branch, to spinal column and dorsal muscles. THE ABDOMINAL AORTA descends from the last dorsal vertebra at the aortic opening of the diaphragm, to opposite the fourth lumbar vertebra, a little to the left of the median line, where it divides into the two common iliac arteries. Relations. — The abdominal aorta is in relation in front with the pancreas, transverse duodenum, stomach, and lesser omentum, the mes- entery, splenic vein, left renal vein, coeliae, solar, and aortic plexuses; behind it is in relation with the vertebral column, the thoracic duct, the left lumbar veins, and the receptaculum chyli; on the right side with the inferior vena cava, vena azygos, thoracic duct, the right crus of tne diaphragm above, and the right semilunar ganglion; on the left side with the left semilunar ganglion and the sympathetic nerve. The branches of the abdominal aorta are: — Phrenic, two in number, supply the diaphragm; Cceliac axis, divides into three large branches, the gastric, hepatic, and splenic; 1. Oastric, or coronaria ventriculi, is the smallest, and sup- plies the lesser curvature of the stomach ; 2. Hepatic, enters the transverse fissure of the liver, and divides into two branches to supply the lobes of the liver. It gives ofE : — Pyloric, to lesser curvature of stomach; Gastro-duodenalis, which descends behind the pylorus, and divides into Pancreatico-duodenalis superior branch to supply the pancreas and duodenum; Gastro-epiploica dextra, along the greater curvature of the stomach from right to left. Cystic, supplies the gall-bladder. THE HEART AND VASCULAR SYSTEM. 193 3. Splenic, the largest of the three, supplies the spleen and gives off the (a) Pancreatic — ^to the pancreas; (h) Gastro-epiploica sinistra, runs from left to right, along the greater curvature, to join the gastro-epiploica dextra; (C) Gastric (vasa brevia), supply the fundus of the stomach. Superior mesenteric, supplies the small intestine (except first part of the duodenum) and the large intestine as far as the splenic flexure, and part of the descending colon. It gives off : — Fig, 110. Abdominal aorta: 1, phrenic; 2, coelfac axis; 8, superior mesen- teric; 4, suprarenal; 5, renal; 6, spermatic; 7, inferior meseuteric; 8, lumbar; 9, sacra media. - (a) Inferior pancreatico-duodenal, supplies duodenum and pan- creas ; (h) Vasa intestini tenuis, supply nearly the whole length of the small intestine (ileum and jejunum) ; (c) Ileo-colic, descends to the right iliac fossa, between the layers of the mesentery, and divides into two branches; fdj Colica dextra, passes to the middle of the ascending colon, to divide into two branches; (ej Colica media, ascends in the layers of the transverse meso- colon, to divide into two branches. 13 iy4 ' HUMAN ANATOMY. Suprarenal, supply the suprarenal bodies ; Benal, supply the kidneys, a large trunk on each side spring- ing directly from the aorta; Spermatic, descend to supply the testicles, and in their pas- sage form one of the constituents of the spermatic cord ; Inferior mesenteric, gives off : — (a) Coliea sinistra., supplies the descending colon; (l>) Sigmoid, supplies the sigmoid flexure of the colon; (g) Superior hsemorrhoidal, the continuation of the inferior mes- enteric, supplies the mucous a,nd muscular coats of the rectum on its posterior aspect. Lumiar branches, four or five on each side, pass backward and outward and divide into the Dorsal branches, supply the spinal cord, muscles, and skin of back; Spinal branches, divide within the spinal canal, ascend and descend, and form an arterial net-work throughout the whole length of the spinal canal ; Abdominal branches, supply the abdominal walls; Middle sacral, is the continuation of the abdominal aorta, descending in the middle of the sacrum to the upper part of the coccyx. THE COMMON ILIAC AKTEKIES descend from the bifurcation of the abdominal aorta, opposite the body of the fourth lumbar vertebra, to a point midway be- tween the last lumbar vertebra and the sacrum, where they divide into the external and internal iliac arteries. The righb common iliac is a little larger than the left, and passes more obliquely outward. Relations. — The right common iliac is in relation in front with the small intestine, the ureter, the peritoneum, and the sympathetic nerves; behind, with the left and right common iliac veins, and to its outer side it rests upon the psoas muscle, and is in relation with the vena cava and the right common iliac vein. The left common iliac is in relation in front with the ureter, the peritoneum, the superior hsemorrhoidal artery, and the sympathetic nerves; behind with the left common iliac vein, to its outer side the psoas muscle, and to its inner side the left common iliac vein. The common iliac arteries give off a few unimportant branches to the ureters, peritoneum, etc. THE INTERNAL ILIAC ARTERY is about an inch and a half in length, extending from the bifurca- tion of the common iliac, about the sacro-lumbar junction, to R. SUP. SUPRARENAL CVSTIC, IL MID, SUPRARKNAL, INTERNAL PUDIC BCLATIC MIDDLE \ tSlCAL DEEP EPIGASTRIC Fig. 111. The arteries of the trunk. THE HEART AOT) VASCULAR SYSTEM. 195 the upper border of the great sacro-sciatic foramen, where it divides iato the anterior and posterior trunks. The branches from the anterior trunk are : — Superior vesical ^ is the pervious remains of the foetal hypo- gastric artery. It supplies the apex and body of bladder, ureter, and vas deferens; Middle vesical, supplies the base of the bladder and part of vesiculae seminalesj it may be given off from the superior vesical ; Inferior vesical, supplies the prostate gland, base of bladder, and vesiculse seminales ; Middle hwmorrhoidal, supplies part of the rectum; OMurator, passes along the side of the pelvis to the obturator foramen, through which it passes. It gives off within the pelvis the iliac, vesical, and pubic branch ; Internal pudic, supplies the external organs of generation. It passes out of the pelvis through the great sacro-sciatic foramen, winds aroimd the ischial spine, and re-enters the pelvis through the lesser sacro-sciatic foramen. It gives off the following branches in the perineum : — (a) Inferior JiamorrTwidal, supplies the muscles and skin of the anus; (bj Siiperflcial perineal, supplies the erector penis and accelerator urinse muscles and the scrotum; (c) Transverse perineal, crosses the perineum on the transversus muscle, which, together with the structures between the bulb and anus, it supplies; (d) Bulbo-urethral, supplies the bulb of the spongy body and Cow- per's gland; (e) Cavernous, supplies the corpus cavernosum, and is one of the terminal branches; (f) Dorsal artery of penis, or clitoris, runs forward on the dorsum and supplies the glans and prepuce. The distribution of the internal pudic artery is much the same in the female as in the male, supplying analogous struc- tures — ^the superficial artery to the labia, the artery of the bulb to the bulb of the vagina, the artery of the corpus cavernosum and dorsal artery to the clitoris. Sciatic artery supplies the muscles at the back of the pelvis, coccygeus, pyriformis, and levator ani, and passes out of the great sacro-sciatic foramen to follow the course of the sciatic nerve. Its branches external to the pelvis are : — Coccygeal, supplies back part of coccyx; Inferior gluteal, to the gluteus maximus muscle; Comes nervi ischiadici, rims in the substance of the great sciatic nerve j 196 HUMAN ANATOMY. Muscular, to the back part of the hip; Articular, to supply the capsule of the hip-joint. Uterine, is distributed to body of uterus, giving branches to the ureter and bladder ; Vaginal, supplies the mucous membrane of the vagina, giv- ing branches to the rectum and neck of the bladder. It is analo- gous to the male inferior vesical. From the posterior trunk Ilio-lumbar gives off two branches: — Lumbar, supplies quadratus lumborum and psoas muscles; Iliac, supplies iliacus, gluteal, and abdominal muscles. Lateral sacral, are two in number — the superior and inferior — supplying the contents of the sacral canal ; Gluteal, gives off a superficial and deep branch to supply the gluteus maximus, medius, and minimus muscles. THE EXTERNAL ILIAO AETEEY extends from the bifurcation of the common iliac to Poupart's ligament, under which it passes to become the femoral. Relations. — It is in relation in front with the peritoneum and intestines, circumflex iliac vein, spermatic vessels, genito-crural nerve, and the lymphatic vessels and nerves; behind it rests upon the psoas muscle, and is in relation with the external Uiac vein, which, together with the vas deferens, is also in relation to it on its inner side, beneath Poupart's ligament. On its outer side it is in relation with the psoas magnus and iliac fascia. Its branches are : — The deep epigastric, arises above Poupart's ligament and ascends obliquely inward between the transversalis fascia and peritoneum to the rectus muscle, in which it ascends to anasto- mose with the termination of the internal mammary artery. It gives off : — Oremasterio branch, to supply the cremaster muscle; PuMe, crossing Poupart's ligament to descend to the inner side of the femoral ring, and Muscular, to supply abdominal muscles and the peritoneum. The deep circumflex iliac ascends outwardly behind Pou- part's ligament to the crest of the ilium, where it gives off Muscular branches, to supply the abdominal muscles. UJO-LL'MBAfl K EXTERNAL [LUt OUTER SIDE OF GREAT TOi; Fig. 112. The arteries of the lower txtremity. THE HEART AND VASCULAR SYSTEM. 197 THE FEMORAL AETEET extends from Poupart's ligament down the inner side of the thigh to the opening in the adductor magnus (Hunter's canal), where it becomes the popliteal. Its course corresponds to a line drawn from a point midway between the anterior superior spine of the ilium and the symphysis pubis to the inner tuberosity of the internal condyle. In the upper part of its course it is super- ficial, where it passes through Scarpa's triangle, but in the lower part it passes backward and becomes very deep. Where it passes under Poupart's ligament the femoral vein is to the inner side, and the anterior crural nerve to the outer side; thus, from within outward, V. A. N. Relations. — It is in relation in front with the Bartorius, the long saphenous nerve, a branch of the anterior crural nerve, and is covered by the fascia lata; behind it is in relation with the psoas magnus, adductor longus, adductor magnus and pectineus, and the profunda vein, and about the middle with the femoral vein; at its inner side it is in relation with the sartorius and adductor longus muscles, and at its upper part with femoral vein; on the outer side, with the vastus internus, and at its lower part, the femoral vein. The branches of the femoral artery are : — Superficial epigastric, descends through the saphenous open- ing in the fascia lata, and ascends in abdomen, supplying the skin, inguinal glands, and superficial fascia; Superficial circumfiex iliac, passes outward to the skin of the groin and over crest of ilium; Superficial external pudic, supplies the skin on the lower part of the abdomen ; Deep external pudic, supplies the scrotum in the male, the labia pudendi in the female; Profunda femoris, arises about two inches below Poupart's ligament, and passes beneath the adductor longtis, giving off the following: — External circumflex, gives o£E ascending, transverse, and descending branches, supplying the muscles on the front of the thigh as low as the knee; Internal circumflex, passes internally, supplying the adduc- tor muscles and the hip- joint; Three perforating, pierce the adductor magnus and brevis, and supply the flexor muscles of the thigh, and give off the medullary nutrient artery. The vessel terminates by a fourth perforating. Muscular, vary in number, and supply the sartorius and vastus internus; 198 HUMAN ANATOMY. Anastomotica magna, gives off two branches — superficial, accompanying the long saphenous nerve, and deep branch, to the inner side of knee, and supplies knee-joint. THE POPLITEAL ARTEET descends from the opening in the adductor magnus to the lower border of the popliteus muscle, passing behind the knee-joint, where it divides into the anterior and posterior tibial arteries. Its branches are : — MtLscular, gives ofE two principal branches: — Superior, to flexors of thigh and vastus externus muscles; Inferior, or sural, to heads of -gastrocnemius and plantaris. Cutaneous, supplies the skin of the calf; Superior external articular, supplies knee-joint and vastus externus ; Superior internal articular, supplies knee-joint and vastus internus ; Azygos articular supplies the synovial membrane and liga- ments of joints; Inferior external articular, supplies knee-joint and fibular head; Inferior internal articular, supplies knee-joint and head of tibia. THE ANTEEIOH TIBIAL AETEET descends from the bifurcation of the popliteal at the lower border of the popliteus muscle; passes over the upper border of the interosseous membrane, between the two heads of the tibialis posticus, and descends on the anterior part of the interosseous membrane and lower part of the tibia to the ankle-joint, where it terminates as the dorsalis pedis. In the upper third of its course it lies between the tibialis anticus and extensor longus digitoruni, resting upon the interosseous membrane, in the middle third, between the tibialis anticus and extensor proprius pollicis, and at the lower third it becomes more superficial, and lies between the extensor proprius and extensor longus digitorum tendons. Its branches are: — Recurrent tibial, ascends to supply the front and sides of the knee-joint and anastomose with the anastomotica magna and popliteal ; Muscular, supplying the muscles and skin of the neighbor- ing parts; Internal malleolar, arises two inches above the ankle-joint, and supplies it and the structures on its inner side; THE HEART AND VASCULAR SYSTEM. 199 External malleolar, supplies the outer side of the ankle. The doesalis pedis artery extends from the bend of the ankle, where it is continuous with the anterior tibial, to the first interosseous space, where it divides into the dorsalis hal- lucis and the communicating. Its branches are:— Tarsal, supplies the tarsal articulations and the extensor brevis digitorum muscle; Metatarsal, passes to the outer side of the foot, and gives off three interosseous branches to supply the adjacent sides of the toes; Dorsalis pollicis, or hallucis, passes along the outer border of the great toe, supplying it; Communicating, descends between the two heads of the first dorsal interosseous to complete, with the external plantar, the plantar arch; Interosseous, branches of the tarsal branch, three in number. THE POSTEHIOE TIBIAL AETEEY begins at the lower border of the popliteus muscle and descends along the posterior and tibial side of the leg to the space between the heel and the inner ankle, where it passes beneath the abduc- tor pollicis and divides into the internal and external plantar arteries. Relations.— In the upper part of its course it lies deeply, being covered by the gastrocnemius and soleus muscle, but in the lower part it becomes more superiieial, being covered only by the skin and the fascia. Its branches are: — Peroneal, descends the inner border of the fibula, supplying the muscles and skin of that region and the back of the ankle ; Anterior peroneal, a branch of the peroneal, pierces the lower part of the interosseous membrane to supply the dorsum and outer side of the tarsus; Muscular, supplies the posterior muscles; Nutrient, supplies the tibia, being the largest nutrient artery in the body; Communicating, passes to the peroneal ; Internal calcanean, supplies the inner side of the sole and heel. The inteenal plantae aeteet, the smaller of the two, passes along the inner side of the foot between the abductor pollicis and the flexor brevis digitorum to the inner border of the great toe, anastomosing with its digital branch. 200 HUMAN ANATOMY. The- EXTERNAL PLANTAR ARTERY passes across the foot to the base of the fifth metatarsal bone, where it turns inward and crosses the foot to the first interosseous space, where it anasto- moses with the communicating branch of the dorsalis pedis to form the plantar arch. Its branches are : — Posterior perforating, are three branches which pass between the heads of the dorsal interossei muscles ; Digital branches — these are four branches which supply the adjacent sides of the three outer toes, and the outer sides of the second and little toes. TABLE OE THE ARTETIIAL SYSTEM. Arch of aorta, Right coronary, Left coronary, Innominate, Aorta. i < LRig ht common f External carotid, carotid, \ Internal carotid, gilt subclavian. T ti i.j f External carotid. Left common carotid, ■< t <. i i-j ' (. Internal carotid. _Left subclavian. External carotid. Superior thyroid, Lingual, Occipital, Facial, ■ Hyoid, Superficial descending branch, Superior laryngeal, Crico-thyroid. Hyoid, Dorsalis lingute. Sublingual, Ranine. In the neck. {Inferior or ascending palatine, tonsillar, submaxillary, sub- mental, muscular. On the f Muscular, inferior labial, inferior face I coronary, superior coronary, ' L lateralis nasi, angular. Muscular, Sterno-mastoid, Auricular, Meningeal, Arteria princeps cervieia, Cranial brandies. THE HEART AXD ^'ASCULAR SYSTEJI. 201 External carotid coutinued, Posterior auricular, Stj-lo-mastoid, Auricular, Muscular, Glandular. pliaryngefl, } ^'^aryngeal, external, meningeal. Temporal, Internal maxillary. Transverse facial. Middle temporal. Auricular, Anterior temporal. Posterior temporal. Maxillary portion. Pterygoid portion. Spheno- maxillary portion, Tympanic (anterior ) , Deep auricular, iliddle meningeal, Small meningeal, Inferior dental. Deep temporal. Pterygoid, Masseteric, Buccal. Alveolar, Infraorbital, Post., or ascending palatine. Vidian, Pterygo-palatine, Nasal or spheno-palatal. portion! } Tympanic (internal, or deep). Arterise receptaculi. Anterior meningeal, Cavern- ous portion. Cerebral portion. . Ophthalmic, Anterior cerebral, Middle cerebral, Anterior choroid, Orbital group. Ocular L group. Lachrymal, Supraorbital, Post, ethmoidal. Ant. ethmoidal. Palpebral, Frontal, Nasal. Muscular, Ant. ciliary. Short ciliary. Long ciliary, Arteria centralis retinae. Posterior communicating. 202 HUMAN ANATOMY. Verte- bral, Cervical Branches, I * > Lateral spinal. Muscular. Cranial Branches, , * V Posterior meningeal, Anterior spinal, . Posterior spinal. Basilar, - Transverse, Anterior inferior cerebellar, Sup. cerebellar. Post, cerebral. Thyroid axis. Inferior thyroid. Superior scapular, (Transversalis humeri), Transversalis colli. Laryngeal, Tracheal, QSsophageal, Ascend, cervical. Muscular. Muscular, Supra-acromial. Superfic. cervical, Post, scapular. Internal mam- mary. Comes nervi phrenici (superior phrenic). Mediastinal, Pericardiac, Sternal, Anterior intercostal. Perforating, Musculo-pnrenie, Superior epigastric. Superior T inter- ?■ Deep cervical branch (profunda cervicis), costal, J Axillary, Superior thoracic, Acromio-thoracic, Thoraeica longa Thoracica alaris, Subscapular, Anterior circumflex, Posterior circumflex. f Dorsal is scapuh-e, \ iledlau branch. Brachial, Superior profunda. Nutrient artery, Inferior profunda, Anastomotica magna. Muscular, Radialj Ulnar. THE HEART AND VASCULAR SYSTEM. 203 Radial, In forearm. In the wrist. In the hand, Radial recurrent. Muscular, Superficialis volse. Anterior carpal. Posterior carpal. Metacarpal, Dorsalis poUicia, Dorsalis indicis. Princeps poUicis, Radialis indicis. Perforating, Interosseons. Ulnar, Thoracic aorta. tn the forearm. In the wrist, , In the hand. ' Anterior ulnar recurrent, Posterior ulnar recurrent. Interosseous, Muscular. f Anterior carpal, I Posterior carpal. f Deep, or communicating branch, \ Digital. Descending Aorta. Pericardiac, Bronchial, Posterior mediastinal. Intercostal. Abdominal aorta. Two phrenic Cceliac axis. Gastric. Hepatic, - Splenic, Pyloric, Gastro- duodenalis. Cystic, Pancreatico- duodenalis superior, Gastro- epiploica dextra. { Pancreatic, Gastro-epiploica sinistra. Gastric (vasa brevia). Superior mesenteric, Suprarenal, Renal, Panoreatico-duodenalis inferior, Vasa intestinalis tenuis, Ileo-colio, Colica dextra, . Colica media. 204 HUMAN ANATOMY. Abdominal aorta, -<, con tin ae i, Spermatic, Inferior mesenteric. Colica sinistra. Sigmoid, Superior haemorrhoidal. Lumbar (four or five branches). Dorsal branch, Spinal branch. Abdominal branches. Middle sacral, ^ Common ilii. Branches to ureter, peritoneum, etc. (- Anterior trunk, o Internal iliac. s External iliac, Posterior trunk. Superior vesical, Middle vesical, Inferior vesical. Middle hcemorrhoidal. Deep epigastric, ie, I Deep circumflex iliac. Within r Iliac, Vesical, pelvis. Pubic. Obturator, . ■ External Without and pelvis. inteinal branches. ■ Inferior haer noiThnidal, Superficial f evineal, Internal Transverse lerineal. pudic, Bulbo-ureth Cavernous, ■al, . Dorsal art'y of the penis. Sciatic, In the female. Coccygeal, Inferior gluteal. Conies nervi ischiadicis. Muscular, Articular. f Uterine, \ Vagir aginal. Ilio-lurabar, Lateral sacral. Gluteal. Cremasteric, Pubic, Muscular. Muscular branches. THE HEART AND VASCULAK SYSTEM. 205 Femoral (continuation). Popliteal, Anterior tibial. Posterior tibial. Plantar arch (from external plantar) , Superficial epigastric, Superficial circumflex iliac. Superficial external pudie. Deep external pudic. Profunda, External circumflex, Internal circumflex. Three perforating. Muscular, Anastomotica magna, Popliteal (continuation). f Superior, \ Inferior, Muscular, Cutaneous, Superior external articular, Superior internal articular, Azygos articular, Inferior external articular. Inferior internal articular, Anterior tibial 1 BUurcation. Posterior tibial, j Recurrent tibial. Muscular, Internal malleolar. External malleolar. Dorsalis pedis (continuation). Tarsal, \ Three Metatarsal, / interosseous. Dorsalis poUicis or hallucia. Communicating, . Interosseous. Anterior peroneal. Peroneal, Muscular, Nutrient, Communicating, Internal caleanean, LExtriltur, } Bifurcation. (Three posterior perforating. Four digital. Pulmonary Artery. — The pulmonary artery carries venous blood from the right ventricle to the lungs. It is about two inches in length, passes upward and baclrward to the left side to the under surface of the transverse portion of the arch of the aorta, where it divides into the right and left pulmonary arteries. It is attached to the under portion of the arch by a fibrous cord, the remains of the ductus arteriosus of foetal life. The right 206 HUMAN ANATOMY. pulmonary artery is larger and longer than the left, and passes behind the ascending aorta and superior vena cava to the root of the right lung, where it divides into two branches. The left pulmonary artery passes in front of the descending aorta and left bronchus to the root of the left lung, where it divides into two branches. THE VENOUS SYSTEM. The venous system, like the arterial, consists of two distinct sets, the systemic and pulmonary. It is composed of seven (7) main trunks and their branches: — Systemic. Pulmonary. 1. Coronary vein; Four pulmonary veins. 2. Superior vena cava; 3. Inferior vena cava. The systemic veins return the venous blood from the body, head, and extremities to the right auricle.. The portal vein, with its branches and capillaries, is an appendage to the systemic set, collecting the venous blood from the organs of digestion and carrying it to the liver, where it breaks into capillaries, and finally reaches the inferior vena cava by means of the hepatic veins. The pulmonary veins are peculiar in carrying arterial blood from the lungs to the left auricle. Systemic Veins. — The coronary sinus returns all the blood from the substance of the heart, except that returned directly from the the walls of the right auricle by the vence Thebesii. It is a dilatation of the great cardiac vein, about one inch in length, situated in the posterior part of the left auriculo-ventricular groove. It opens into the right auricle, its orifice being pro- tected by the coronary valves, and receives the following : — Great cardiac, or coronary vein; Posterior cardiac; Anterior cardiac; Small cardiac; Middle cardiac; Oblique vein. The superior vena cava is a short trunk formed by the union of the right and left innominate veins. It receives the vena azygos major, has no valves, and is smaller in size than the aorta. It ends in the right auricle, receiving the blood from the whole upper half of the body and the right lymphatic and thoracic ducts. The left innominate passes to join the superior cava in front of the great arteries of the arch. The inferior thyroid, the internal mammary, and vertebral veins follow closely the courses of the corresponding arteries, and terminate in the innominate vein. THE HEART AND VASCULAR SYSTEM. 207 The internal jugular vein is formed by the lateral and the superior petrosal sinuses, descends at the outer side of the com- mon carotid behind the anterior border of the stemo-mastoid muscle, and joins the subclavian vein to form the innominate. At its junction with the subclavian the left internal jugular vein receives the thoracic duct and the right internal jugular vein the right lymphatic duct. The Sinuses of the Dura Mater. — These are venous channels analogous to veins between the layers of the dura mater. They are destitute of valves, follow no vessels, and their inner coat is continuous with the lining membrane of the veins. They are fifteen in number, divided into two sets, those at the back and upper part of the skull five ia number : — Superior longitudinal, Lateral sinuses (2), Inferior longitudinal. Occipital sinus. Straight sinus. And those of the base five also : — Cavernous (2), Superior petrosal (2), Circular, Transverse (anterior occipital. Inferior petrosal (2), Leidy). The superior longitudinal sinus arises at the foramen csecum, passes backward along the margin of the falx cerebri to the crucial ridge of the occipital bone, where it terminates in the torcular herophili (confluence of the sinuses). The inferior longitudinal sinus passes in the free margin of the falx cerebri. The straight sinus passes backward from the junction of the tentorium with the falx cerebri to enter the confluence of the sinuses. The lateral sinuses pass horizontally outward from the con- fluence of the sinuses, torcular Herophili, along the temporal bone to the jugular foramen, where they terminate in the internal jugular vein. The occipital sinus, the smallest of all the sinuses, com- mences at the margin of the foramen magnum on either side and passes backward to the confluence of the sinuses. The cavernous sinuses pass from the sphenoidal fissure along either side of the sella turcica to the apex of the petrous portion of the temporal bone, where they join the petrosal sinuses. They are crossed by fibrous bands or offsets of the dura mater, and inclose the pathetic, motor oeuli, abducens, and ophthalmic nerves, ' and the internal carotid artery, from which they are separated by the lining membrane. 208 HUMAN ANATOMY. The circular sinus is. formed by two small vessels passing in front of and behind the pituitary body, and connecting the cavernous sinuses. The inferior petrosal sinus on each side runs in a groove between the petrous portion of the temporal bone and the basilar portion of the occipital, connecting the cavernous sinuses with the lateral sinuses. The superior petrosal sinus on each side passes along the upper border of the petrous portion of the temporal bone, con- necting the cavernous with the lateral sinuses above. The transverse sinus is a small, straight sinus, connecting the inferior petrosal and cavernous sinuses. Cehebral Veins. — The cerebral veins consist of two sets — the superficial, on the surface, and the deep, within the substance. They include: — Superficial — Superior cerebral veins. Inferior cerebral veins. Deep — Ventricular veins, or venae Veni corporis striati, Galeni, Choroid vein. The cerebellar veins consist of the superior, inferior, and lateral. DiPLOio AND Meningeal Veins. — The diploic veins, five in number, — frontal, anterior temporal, posterior temporal, occipital, — communicate with the sinuses of the brain and with the veins of the dura mater, the scalp, and orbit. The meningeal veins follow the course of the corresponding arteries, two accompanying each vessel through its course, and open into the sinuses of the dura mater. The great meningeal veins terminate either in the cavernous sinuses or by emptying into the internal maxillary vein. The ophthalmic vein commences at the internal canthus of the eye in an anastomosis with the facial, passes backward along the inner part of the orbit, through the sphenoidal fissure, to empty into the cavernous sinuses. The external jugular vein, smaller than the internal jugular, is formed by the union of the posterior auricular with the tem- poro-maxillary veins; or it may be formed by union with the facial. It descends beneath the platysma muscle from the angle of the jaw to the middle of the clavicle, where it terminates by emptying into the subclavian. It receives the following veins : — Anterior jugular, Suprascapular, Posterior jugular, Transverse cervical. THE HEART AND VASCULAR SYSTEM. 209 The facial vsin commences as the angular at the internal canthus of the eye, where it anastomoses with the ophthalmic vein, and receives the frontal vein. About the angle of the jaw it usually ends in the internal jugular, but sometimes empties into the external jugular or unites with the temporo-maxillary, to enter the external jugular. It receives — • Supraorbital and superior palpebral, Labial, Nasal veins. Submental, Inferior palpebral, Submaxillary, Buccal and masseteric. Palatine. The temporal vein is formed by the anterior temporal, which anastomoses with the frontal, and the posterior temporal, which anastomoses with the occipital. It penetrates the parotid gland and forms the temporo-maxillary vein by uniting with the in- ternal maxillary. It receives Articular veins. Transverse facial. Anterior auricular. Parotid. Middle temporal. The internal maxillary vein follows the course of the corre- sponding artery, and receives veins corresponding to the branches of that vessel. Some of these branches form the pterygoid plexus. It passes backward and unites with the temporal vein to form the temporo-maxillary. The temporo-maxillary vein, formed by the junction of the internal maxillary in part or whole with the temporal, passes through the parotid gland and receives the posterior auricular, to form the external jugular veia. The occipital veins follow the course of the artery, and terminate, usually, in the internal jugular; occasionally, in the external jugular. The mastoid vein, passing through a foramen in the mastoid portion, connects it with the lateral sinus. Veins op the Tongue and Throat. — The dorsal lingual, formed by branches from the tonsils, epiglottis, and tongue, fol- low the course of the lingual nerve to empty into the facial, either jugular, or the pharyngeal. The ranine pursues the course of the hypoglossal nerve, and terminates either in the facial or in one of the jugulars. The pharyngeal, commencing in the pharyngeal plexus, re- ceives branches from the brain, and terminates about on a level with the hyoid bone in the internal jugular. The superior thyroid conveys the blood from the larynx, trachea, and thyroid gland to the internal jugular. Veins of the Uppee Extremity. — Superficial and deep. 210 HUMAN ANATOMY. The subclavian vein, the continuation of the axillary, unites with the internal jugular to form the innominate. In its pas- sage over the first rib it is separated from the artery by the scalenus anticus muscle. Deep Set. — The deep veins accompany the corresponding arteries and form the venae comites, one on either side, except the axillary, which has a single vein. The valves are more numer- ous in the deep set. About the middle of the arm one of the brachial veins receives the basilic vein, one of the larger veins of the superficial set. The two brachial veins unite with the basilic to form the axillary vein, which in turn becomes the subclavian vein, receiving in its course beneath the clavicle the cephalic vein, a branch of the superficial set. Superficial Set. — The anterior ulnar vein commences on the inner surface of the hand and wrist, and terminates by uniting with the posterior ulnar or the median vein. The posterior ulnar vein commences on the inner back portion of the hand, anasto- mosing with the radial cutaneous vein, and ascends to the bend of the elbow, where it becomes the basilic vein ; The basilic vein, from its formation at the bend of the elbow, ascends the inner side of the biceps, pierces the fascia, to join the brachial vein. The radial cutaneous commences on the radial, dorsal aspect of the hand, and at the bend of the elbow unites with the median cephalic, to become the cephalic. The cephalic vein ascends along the outer border of the biceps, and above in the groove between the deltoid and pectoralis major, and passes beneath the clavicle, to end in the axillary. The median vein receives the blood from the palmar surface of th.e hand and the front of the forearm, and usually divides into two branches — the median cephalic, passing outward to join the cephalic, and the median basilic, passing inward to join the basilic. The median basilic vein, the larger, is the one usually selected for plebotomy. Veins of the Trunk. — The vena azygos major commences in the abdomen, opposite the first or second lumbar vertebra, as a continuation upward of the right ascending lumbar vein. It communicates with the right renal and the inferior vena cava. Through the lumbar veins it establishes communication with the right common iliac vein. It passes through the aortic opening of the diaphragm, or through an aperture in the right crus, ascends on the dorsal vertebrse, arches over the root of the right lung, and empties into the superior vena cava. It is in relation on its left side with the aorta, thoracic duct, and oesophagus. It receives THE HEART AND VASCULAR SYSTEM. 211 Vena azygos minor, Right superior intercostal Vena hemi-azygos accessoria, vein, Lower end of left superior Right subcostal vein, intercostal vein. Oesophageal, Lower 8 or 9 intercostal Right bronchial. veins. The hemi-azygos veins are two in number : — • The vena azygos minor arises on the left side, similarly to the azygos major, and passes through the left crus of the dia- phragm and about the eighth dorsal vertebra crosses beneath the aorta to join the vena azygos major; The vena hemi-azygos accessoria communicates with the upper left intercostal vein, and terminates either in the vena azygos minor or the azygos major. The inferior cava formed by the junction of the common iliac veins, at the sides of the fourth lumbar vertebra, ascends on the right of the aorta, grooves the posterior border of the liver, and passes through the quadrate foramen in the central tendon of the diaphragm, and ends in the right auricle. It receives Middle sacral, Renal veins — the left, the Lumbar, longer, crosses in front Spermatic (from spermatic of the aorta, plexus). Suprarenal, Ovarian (from ovarian Phrenic, plexus), Hepatic (2 or 3). The portal vein, about three inches in length, is formed by the inferior and superior mesenteric, the gastric, splenic, and pancreatic veins. It ascends in the right border of the lesser omentum to the transverse fissure of the liver, where it divides into the right and left. Its blood is distributed through the liver, mixing with the arterial blood from the hepatic artery, to be returned to the inferior cava by the hepatic veins. It receives The superior mesenteric. Cystic, Splenic, Inferior mesentery, Coronary, Right gastro-epiploie. The portal vein and its branches are destitute of valves. Veins of the Veetebeal Column. — 1. The dorsi-spinal veins commence in an intricate net-work surrounding the verte- bral arches and their processes. They communicate with the in- tercostal, vertebral, intraspinal plexus, lumbar, sacral, and the superficial veins of the back. 2. The intraspinal or meningo-rachidian veins form an in- tricate plexus between the vertebra and dura mater within the spinal canal. They consist of four longitudinal veins — two in 212 HUMAN ANATOMY. front and two behind. The anterior longitudinal spinal veins, two in number, extend along the sides of the vertebral bodies and opposite the bodies communicate by transverse trunlcs, which, in their passage beneath the vertebral ligament, receive the di- ploic veins, or vence basis vertebrce, from the interior of the body. The posterior longitudinal veins, also two in number, smaller than the anterior, extend down the vertebral arches and are con- nected by transverse branches opposite the latter. They receive the veins from the spinal cord and its mem- branes. 3. The venoe basis vertebrce return the blood from the bodies of the vertebrte into the anterior intraspinal plexuses. The vence medulli spinalis are the essential veins of the cord, situated between the arachnoid and pia mater. Common Iliac Veins. Formed by the internal and external iliac veins uniting opposite the sacro-iliac articulation, pass beneath the right com- mon iliac artery to a point a little to the left of the body of the last lumbar vertebra, where they unite to form the inferior vena cava. The right is shorter and more vertical in its course, and both are without valves. The internal iliac vein corresponds to the distribution of the corresponding artery. It receives the following venae comites : — Gluteal, Internal pudic, Sciatic, Lateral sacral, and Obturator, Middle sacral; and the following plexuses : — Vesico-prostatio, 1 . „„,„. Uterine and 1 .„ , ,_ HiEmorrhoidal, /"i«'^^«' Vaginal, ) 'n female. The lateral and middle sacral form a small plexus — the plexus sacralis. The- veins of the rectum, bladder, and generative organs anastomose freely and form three plexuses : — ; 1. Hcemorrhoidal plexus encircles the lower part of the rec- tum, communicates with the sacral and prostatic plexuses, and veins from it join the inferior mesenteric, internal iliac, and pudic veins. 2. Vesica-prostatic plexus surrounds the membranous por- tion of the urethra, neck of bladder, prostate body, and seminal vesicles. It communicates behind with the hemorrhoidal. THE HEART AND VASCULAK SYSTEM. 213 3. Utero-vaginal plexuses. — These correspond in the female with the prostatic, and communicate with the vesical and hsemor- rhoidal plexuses, the ovarian, pudic, and through the uterine veins join the internal iliac veins. During pregnancy these veins or plexuses become greatly dis- tended and enlarged, forming the uterine sinuses, but retain a straight course. The pudic vein follows the same course as the artery, com- municates with the prostatic and hsemorrhoidal plexuses, and terminates in the internal iliac vein. In the female it originates from the clitoris and perineum, and communicates with the vaginal plexus. The dorsal vein of the penis returns the blood from the body of that organ. It commences as two venje comites of the dorsal artery, which unite on the dorsal surface of the root of the penis, perforates the triangular ligament, and again divides to termi- nate in the prostatic plexus. The veins of the corpus cavernosum emerge at the lower groove, and turn round their outer side to join the dorsal vein. The dorsal vein of the clitoris has a corresponding origin and course, and empties into the vaginal plexus. The EXTERNAL ILIAC VEIN, the Continuation of the femoral, lies internal to the artery beneath Poupart's ligament, and joins the iaternal iliac opposite the sacro-iliac symphysis, to form the common iliac vein. It runs along the brim of the pelvis. It receives at its commencement the venae comites of the epigastric artery and the circumflex iliac. Lower Extremity. — The deep veins of the lower extremity pursue the exact course of the corresponding arteries, anastomos- ing across the vessels they accompany, except the femoral, which has but one vein. The popliteal vein, formed by the junction of the anterior and posterior tibial veins, ascends to the lower margin of Hunter's canal, where it becomes the femoral. It receives the articular veins, sural veins, and the external saphenous. At its commencement it lies internal and superficial, at the middle of the space directly behind, and, in the upper part of its course, external to the artery. The femoral vein, at first behind the artery, inclines to the inner side as it ascends, and at Poupart's ligament becomes the external iliac. It receives the profunda femoris, the internal saphenous vein, and numerous muscular veins. The superficial veins of the lower extremity consist of two principal trunks: the external short saphenous, or vein, com- mences on the dorsum of the foot, passes behind the outer mal- 214 HUMAN ANATOMY. leolus, ascends the leg, and pierces the deep fascia in the popliteal space, to terminate in the popliteal vein. The long saphenous vein, and the larger, arises from the dorsTim and inner side of the foot, ascends the front and inner side of the leg and thigh, 'and passes through the saphenous opening to Join the femoral vein. The Pulmonary Veins. — The pulmonary veins are four short, venous trunks, two from the base of each lung passing to the left auricle, returning arterial blood. They differ from other veins in the following respects : — • 1. They are without valves; 2. They carry arterial blood; 3. They accompany the arteries singly; 4. They are a little larger only than their arteries. The right are longer than the left, and pass from the root of the lung, on a lower level than the artery, behind the aorta, superior cava, and right auricle, to enter the left auricle. The Lympliatic System. The lymphatic system includes the lymphatic vessels and glands and the lacteals, and forms an important accessory to the blood-vascular system, collecting the transuded, unappropriated fluids of the body and the nutritive material derived from the food and conveying it into the venous system. It consists of two main trunks : — (a) Thoracic duct; (bj Eight lymphatic duct; and five smaller trunks : — 1. Jugular lymphatic trunk; 2. Subclavian lymphatic trunk; 3. Broncho-mediaatinal lymphatic trunk; 4. Lumbar lymphatic trunk; 5. Intestinal lymphatic trunk. Lymphatics have been found in nearly every organ and tex- ture in the body except the brain, the spinal cord, cartilage, tendon, eyeball, placenta, umbilical cord, membranes of the ovum, hair, cutis, and the labyrinth of the ear. They appear to originate as fine capillary nets interwoven among the blood- vessels and proper elements of the tissues, or, more minutely, in the lymph, perivascular, and perineural spaces. In the villi they commence as closed, club-like tubes. The lymphatic capillaries are somewhat larger than the vas- cular capillaries and destitute of valves. Their main trunlcs pass through lymphatic glands lying in their course, before doing THE HEART AND VASCULAR SYSTEM. 215 which, however, they divide into afferent vessels, which, on emerg- ing, unite into a smaller number of larger vessels, the efferent. In structure, they are composed of three coats — ^the external fibro-areolar, middle muscular, and internal or endothelial and elastic. The lymphatic glands are generally situated in the course of the blood-vessels, lymphatic vessels, or lacteal vessels, being ac- cumulated together in certain localities, as the neck, abdomen, axilla, etc. The lymphatic glands and vessels are named from the regions they occupy or the vessels they accompany, and consist usually of a superficial and deep set. Thus, we have cervical, axillary, mediastinal, lumbar, inguinal, etc., and the lymphatic vessels corresponding. The lacteals, or chyliferous vessels, are the lymphatic vessels of the small intestine, and differ only from the others in carrying chyle during digestion from the intestines to the thoracic duct. The thoeacic duct is formed by the junction of the two lumbar lymphatic trunks with the intestinal lymphatic trunk, in front of the second lumbar vertebra, between the aorta and in- ferior vena cava, as the receptaculum chyli, or cistern of Pecquet. This receptacle is about one to two inches long and a quarter of an inch wide. From its origin the thoracic duct ascends through the abdomen, passes through the aortic orifice in the diaphragm, and ascends behind the oesophagus between the aorta and azygos vein to the fourth dorsal vertebra, where it passes to the left be- neath the aorta, and ascends between the oesophagus and the left subclavian artery to the last cervical vertebra, where it arches forward, outward, and downward to enter the junction of the sub- clavian and the left internal jugular vein at its posterior aspect. Its orifice is protected by a pair of valves. It receives all the lymphatic vessels below the diaphragm, those of the left side of the head, neck, and left upper extremity. The right lymphatic duct is about half an inch long and one-twelfth inch wide. It empties in a corresponding manner on the right side to the thoracic duct. It receives all the lym- phatics of the right side of the thorax, neck, head, and right upper extremity. Lymphatics op the Head and Neck. — The substance of the brain is probably destitute of lymphatics, but they are very numerous in the pia mater and choroid plexuses of the lateral ventricles, and pursue the same course as the principal veins, to emerge at the base through the various foramina, to terminate in the deep cervical glands. The occipital lymphatic vessels termi- nate in the posterior auricular and occipital glands. The tem- 216 HUMAN ANATOMY. poral lymphatic vessels terminate in the anterior or auricular glands. The lymphatics of the face are superficial and deep. The superficial lymphatics are numerous, and terminate in the submaxillary glands, six or more in number; the deep lym- phatics accompany the branches of the internal maxillary artery, and terminate in the deep cervical and deep parotid glands about the ramus of the jaw. Lymphatics of the Upper Extremity. — Lymphatics of the upper extremity are composed of two sets, the superficial and deep. The superficial lymphatic glands are few in number, one or two only being situated at the internal condyle of the humerus. The deep lymphatic glands lie along the course of the vessels, and communicate with the axillary glands. All of these glands unite in the deep axillary glands, about eight to ten in number, which communicate with the deep cervical glands, and through them empty into the subclavian lymphatic trunk, to end finally in the thoracic or right lymphatic duct. The superficial and deep ves- sels of the thorax, the former in the skin, the latter from the mammary glands, pectoral and other muscles, for the most part, pass to the axilla, a few only terminating in the glands below the clavicle. The Cavity op the Thorax. — The intercostal lymphatic vessels, derived from the side of the abdomen and thorax, pleurte, diaphragm, spinal canal, muscles of the back, etc., follow the course of the veins, traverse fifteen to twenty intercostal glands near the heads of the ribs, and terminate in the thoracic duct. The posterior mediastinal glands are between the intercostal glands, and communicate with them. They receive vessels from the pericardium, oesophagus, and diaphragm. Some of the effer- ent vessels end in the bronchial glands, others in the thoracic duct. The anterior mediastinal lymphatic vessels are derived from the anterior wall of the abdomen and thorax, the dia- phragm, pericardium, upper surface of the liver, heart, and thy- mus gland. They traverse about eighteen to twenty anterior mediastinal glands, situated in the course of the internal mam- mary vein, pericardium, and great vessels of the heart, and terminate in thoracic and right lymphatic ducts. The pulmo- nary lymphatic vessels consist of a superficial and deep set, trav- ersing in the last part of their course the pulmonary glands. The bronchial glands are twenty or more glands at the bifurcation of the trachea and root of the lungs, and receive the lymphatic vessels of the lungs and bronchi. They become pigmented, and are often the seat of disease. Their efferent vessels terminate on the right side in the right lymphatic duct, either directly or by forming the broncho-mediastinal trunk, and on the left side into the thoracic duct. THE HEART AND VASCULAR SYSTEM. 217 Lower Extremity and Pelvis. — The superficial lymphatic vessels from the back of the foot follo-w the course of the long saphenous vein, passing to the superficial inguinal glands; and those from the sole of the foot follow the short saphenous vein, joining the deep set in the popliteal space. The deep lymphatic vessels follow the deep veins, traverse two to four popliteal glands, and join the deep inguinal glands. The deep lymphatic glands in the pelvis consist of three sets : the external iliac, around the external iliac vessels ; the internal iliac, around the internal iliac vessels ; and the sacral glands, on the anterior surface, and in the mesorectal folds. Lymphatics of the Abdomen. — The lymphatic vessels of the external and internal iliac glands enter the lumbar glands, twenty-five or more in number, situated upon the vertebrae, the origin of the diaphragm, the psoas and quadratus lumborum mus- cles, and the great blood-vessels. They receive vessels from the kidneys, ureters, loins, suprarenal bodies, ovaries, and testicles, and form lumbar lymphatic trunks, which ascend to join the tho- racic duct or receptaculum chyli. The lymphatic vessels of the stomacli follow the general course of the blood-vessels. They consist of three groups : the first, along the lesser curvature, pass- ing to the glands along the pylorus ; the second, at the great end of the stomach, passing to the splenic lymphatic glands ; and the third, at the greater curvature, passing to one of the principal lacteal vessels. The lymphatics of the small intestine are called also lacteals. They pass between the layers of the mesentery ,and traverse a large number of mesenteric glands, arranged irregularly into three rows. The lymphatic vessels of the large intestine traverse the mesocolic glands, about thirty in number, and pass into the superior mesenteric glands. The vessels from the spleen and pancreas follow the course of the splenic vein, traverse a number of the glands, to end in the coeliac glands. The lymphatics of the liver consist of superficial and deep, and are very extensive, both on the upper and lower surfaces. They terminate in the right lymphatic duct, the glands of the gastro- hepatic omentum, the anterior mediastinal glands, the oesopha- geal glands, the glands of the lesser curvature, and of the thoracic duct. The deep lymphatics follow the course of the portal vein and hepatic artery and duct, and, emerging, join one of the lacteal vessels before it enters into the thoracic duct. The cceliac glands, fifteen or twenty in number, are situated behind the pan- creas and duodenum, the aorta, portal vein, cceliac, and superior mesenteric vessels. Their efferent vessels form the intestinal lymphatic trunk and empty into receptaculum chyli. THE ALIMENTARY APPARATUS. The alimentary apparatus consists of the alimentary canal and of certain accessory organs. The former is a mnscnlo-membranous canal about thirty feet in length, extending from the mouth to the anus, and comprises : first, the organs of deglutition, consisting of the mouth, pharynx, and oesophagus ; and, second, the organs of digestion, consisting of the stomach and small and large intestines. It is lined throughout by mucous membrane. The accessory organs com- prise the teeth, salivary glands, liver, and pancreas. Alimentary canal. Alimentary Apparatus. Mouth, Organs of deglutition, ] Pharynx, CEsophagus. Accessory organs, Organs of digestion. Teeth, Salivary glands. Liver, . Pancreas. Stomach, Duodenum, Small intestine, ■{ Jejunum, eum. f Dii e, \ Je; (. He tine, \ I Coecura, _ Large intestine, -j Colon, Rectum. Parotid, Submaxillary, Sublingual. The mouth is an oval cavity in which mastication takes place preparatory to deglutition. It is placed at the entrance of the alimentary canal, is bounded laterally by the alveolar processes of the upper and lower jaws and cheeks ; above by the upper teeth and hard palate; below by the tongue, the lower jaw, the mucous membrane between, and the lower teeth; in front by the lips; and behind by fauces and soft palate. It terminates posteriorly at the anterior pillars of the fauces, through the fauces into the pharynx. ('218') THE' ALIMENTARY APPARATUS. 219 It is invested throughout (except on the teeth) with highly vascular mucous membrane covered with stratified squamous epithelium containing conical papillae. It presents for examination the following parts : — The Teeth. — The teeth are firmly implanted within the alveoli of the jaws and surrounded by the gums. The gums are composed of dense fibrous tissue, covered by smooth, vascular, mucous membrane of slight sensibility. This fibrous tissue is continuous with the periosteum of the jaws, and forms about the neck of the teeth a constricted ring — ^the dental ligament. " There are four kinds of teeth — incisors, canines or cuspids, premolars or bicuspids, and molars. Man is provided with two sets of teeth, the temporary, deciduous, or milk teeth, which appear in childhood, and the permanent, which appear after the shedding of the "milk teeth and last until old age. The temporary or deciduous teeth are twenty in. number, ten in each jaw, or five ia each side of each jaw : — Two incisors, one canine, and two molars. The permanent teeth are thirty-two in number, sixteen in each jaw, or eight in each side of each jaw : — Two incisors, one canine, two bicuspids, three molars. Bach tooth consists of three parts: — Crown, or body, the enameled portion above the gum ; necTc, the constricted portion between the crown and root; root, or fang, within the alveolus, and covered with cement. Characteristics. — Incisors, or cutting, are so called from their wedge-shaped, chisel-like crown, being adapted for biting or cutting the food. The fang is long, single, conical, and com- pressed at the sides. Cuspids, or canines, have been so named from their con- spicuous character in the canine or dog tribe. The crown is large, conical, convex in front, and beveled behind. The fang is single, longest, and thickest of all the teeth. Bicuspids, or premolars. The crown has a pair of project- ing tubercles or cusps. Pang is conical, single, but deeply grooved, indicating a disposition to bifurcate. Molars, commonly known as grinders or jaw teeth. Crown, broad, quadrilateral, with four cusps in upper, five in lower molars. Lower molars have usually a pair of fangs placed lat- erally; the upper, three fangs, two external and one internal. The last or third molar has but one fang (with a tendency to divide into the same number of roots as the other molars), and is known as the dens sapientice, or "wisdom tooth/' from its late appearance. 220 HUMAN ANATOJIY. Structure. — On section'a tooth consists of two portions, the pulp cavity and the solid portion surrounding it. The solid portion consists of three structures, dentine, or ivory, which forms the principal mass of the tooth; enamel, which covers the crown, and cement, which covers the surface of the fang. Dentine, or ivory, resembles bone, but differs from it in composition and structure, consisting of twenty-eight parts ani- mal and seventy-two parts mineral matter, and being made up of minute tubuli held together by the intertubular substance. The dental tubuli are minute cylindrical canals Visoo of an inch in diameter. They pass in a spiral direction from the pulp cavity to the periphery. Enamel, the hardest and densest of all organized bodies, con- tains but 3.5 per cent, animal matter, and is composed of minute hexagonal rods ^/ggoo of an inch in diameter,' placed at right angles to the surface of the dentine. The external surface of unworn enamel can be separated as a thin, homogeneous mem- brane, Nasmyth's mernbrane. Cement, or cru^ta petrosa, is a thin layer of true bone with eanaliculi and lacunse, disposed on the surface of the fang. The pulp cavity is a cavity within the base of the crown, continuous with a canal in the centre of the fang, and open at the apex of the fang for the entrance of vessels and nerves. It is filled with dental pulp. Dental pulp consists of two kinds of cells, the fusiform and the columnar, or odontoblasts ai Waldeyer, held together by loose connective tissue. It is soft, vascular, and highly sensitive. The nerves are both medullated and non-medullated, and form a rich plexus beneath the odontoblastic layer. The terminal fibrils probably unite with these cells, but the exact distribution is still unsettled. Development — Temporary. — They are formed very early, seven to eleven weeks, in the primitive dental groove by an in- volution of the epithelium of the oral cavity covering the maxil- lary arches into the blastema or corium and connective tissue below, the former forming the enamel, the latter the cement and dentine. The enamel is formed by the enamel germ (a mass of epi- thelial cells) descending into the dental groove until it meets the papilla, a vascular growth extending upward from the connective tissue, upon which it forms a cap. A vascular membrane in- closing the enamel germ then extends itself — as the dentinal sac — ^upon the united papilla and enamel germ and cuts the latter off from its former epithelial structure. The cells become dif- ferentiated and finally calcify. THE ALIMENTARY APPARATUS. 221 The dentine is formed by tlie development of odontoblasts in the periphery of the papilla in a similar manner to the develop- ment of osteoblasts in bone. The cement is developed from the wall of the dental sac by the intramembranous process of ossification. Permanent. — The successional permanent teeth, or those re- placing the temporary, are formed in a different manner from the superadded, or three molars in each side of each jaw. The former are developed similarly to the temporary teeth, but in a secondary dental groove, from which after their forma- tion they recede behind the germs of the temporary teeth, in- closed in sacs. The molars, or superadded teeth, however, are formed by extensions backward of a portion of the enamel germ of the tooth immediately in front. e-8 C. la. C.In. L.In. C. M. M. 6-8 7-9 17-18 14-16 18-24 Fig. 113,— Temporary Teeth. Eruption, or "cutting of the teeth," takes place by the growth of the fang, the gums being absorbed by the pressure of the advancing crown. The development of the fangs of the permanent causes the absorption of the fangs of the temporary teeth through the agency of the odontoclasts, multinucleated cells corresponding to the osteoclasts of bone. The eruption of the temporary teeth takes place in months in the order shown in Fig. 51, the lower preceding by a short time the upper (according to Dr. C. N. Pierce, in "American System of Dentistry") . The order of the eruptions is : first the lower central incisors, the upper central incisors, lateral incisors, upper and lower, first molars, followed by the canines or cuspids, and ending with the second molars. 222 HUMAN ANATOMY. The eruption of permanent teeth takes place in years as in Pig. 63, the order of the eruptions being, first the "first or sixth year molar," followed by the first central incisors, four lateral incisors, four first bicuspids, four second bicuspids, the first canines or cuspids, the four second molars, and ending -with the four third molars or "wisdom teeth." About the sixth year the jaws contain the temporary teeth fully erupted, and the crowns of all the permanent teeth except- ing the four wisdom teeth, in all forty-eight. It should be noted that the first permanent or "sixth -year" molar is erupted before any of the permanent teeth, and that the second bicuspid takes the place of the second temporary molar. 7-8 8-9 12-14 10-11 1W2 6-7 12-16 C. III. L. In. Co,. Bi. Bi. M. 31. 16-20 Wi. C. In. L. In. Ca. Bi. Bi. M. M. Wi. 7-8 8-9 12-14 lO-U 11-12 6-7 12-16 16-20 Fig. 114.— Permanent Teeth. Vessels and Nerves of the Teeth. — The arteries of the upper teeth are derived from the anterior dental branches of infra- orbital and posterior or alveolar dental branches of the internal maxillary; of the lower teeth, from the inferior dental branch of the internal maxillary. The nerves are distributed to the upper teeth from the an- terior and posterior dental branches of the superior maxillary (second division of fifth cranial nerve), and to the lower teeth from the inferior maxillary (third division of fifth cranial nerve) . Soft palate is a movable fold of mucous membrane sus- pended from the posterior border of the hard palate, and inclos- ing an aponeurosis, vessels, nerves, glands, and the following THE ALIMENTAHY APPARATUS. 223 muscles on each side : tensor palati, levator palati, palatoglossus, palato-pharyngeus, and azygos uvulse, the latter uniting with its fellow to form the uvula. Hard palate is formed by the palatal process of the superior maxillary and the palate-bone, and covered by a thick, dense structure composed of mucous membrane and periosteum com- bined. It forms the roof of the mouth, and presents a median raphe and corrugated surface. Anterior pillars of the fauces are folds of mucous mem- brane arching downward and forward from the base of the uvula to the base of the tongue, and inclosing the palatoglossus muscles. Posterior pillars of the fauces are similar folds arching downward and backward from the base of the uvula to the sides of the pharynx, and inclosing the palato-pharyngeus muscles. The tonsils, or amygdalse, are small, almond-shaped, glandu- lar bodies situated on each side of the fauces between the an- terior and posterior pillars. They rest upon the superior con- strictor of the pharynx, which separates them from the ascending pharyngeal and internal carotid arteries. They are composed of numerous follicles (lined by closed capsules containing adenoid tissue), which contain a thick, grayish secretion and open on the surface of the gland by a dozen or more orifices. The arteries to the tonsil are from the tonsillar and ascend- ing palatine of the facial, dorsalis linguse from the lingual, ascending pharyngeal from external carotid, branch from small meningeal, and descending palatine branch of internal maxillary. The nerves are from glossopharyngeal and Meckel's gan- glion. The Salivary Giands. — The salivary glands communicat- ing with the mouth are three: the parotid, submaxillary, and sublingual. The parotid gland, so called from its location near the ear, is the largest, weighing from a half to one ounce. It occupies the space in front of the ear, bounded below by the angle of the jaw, and a line extended from it to the mastoid process, above by the zygoma, in front by the masseter muscle, and behind by the mastoid process, the external meatus, and the digastric muscle. The external carotid artery, the temporo-maxillary vein, the facial nerve, and the great auricular nerve pass through it. The duct of the parotid gland, Steno's or Stenson's duct, empties its secretion into the mouth. It is about two and a half inches in length, of the diameter of a crow's quill, and crosses the face upon the masseter muscle, through the substance of the buccinator muscle, in the direction of an imaginary line 224 HUMAN ANATOMY. drawn about a finger's breadth below the zygoma, from the lower part of the ear to midway between the ala of the nose and the margin of the upper lip, to open about the position of the second molar tooth of the upper jaw. Its arteries are branches of the external carotid. The veins empty into the external jugular. The lymphatics empty into the superficial and deep cervical glands, and the nerves are derived from the facial, from the superficial temporal branches of the great auricular, the auriculo-temporal, and sympathetic. The submaxillary gland occupies the submaxillary fossa on the inferior surface of the inferior maxilla, within the sub- maxillary triangle of the neck. The facial artery grooves its upper and posterior border, and it is separated behind from the parotid gland by the stylo-maxillary ligament. The duct of the submaxillary gland, or Wharton's duct, about two inches in length, passes forward between the hyoglossus, geniohyoglos- sus, and mylohyoid muscles, to open at the side of the frsnum linguse. Its arteries axe from the facial and lingual ; the veins follow the course of the corresponding arteries; the nerves are from the submaxillary ganglion, the sympathetic, and the mylohyoid branch of the inferior dental. The sublingual gland, the smallest, lies on the floor of the mouth, at the side of the frsenum linguEe, beneath the mucous membrane. Its ducts, called the ducts of Eivini, from eight to twenty, open on the mucous membrane. One of them, the longest, called the duct of Bartholin, joins Wharton's duct. Its arteries are from the submental and sublingual. The nerves are branches from the gustatory. Besides these glands, the mucous membrane of the mouth is plentifully supplied with mucous glands. The pharyns is a masculo-membranous sac, extending from the basilar process of the occipital bone above to the level of the fifth cervical vertebra or the cricoid cartilage below. It is about four and a half inches in length. It has communicating with it seven openings : — - Two posterior nares, Larynx, Two Eustachian tubes, CEsophagua, Mouth. It is composed of three coats: — 1. Fibrous coat, or pharyngeal aponeurosis, attached above to the pharyngeal spine of the basilar process of the occipital bone ; it affords attachment in the median line to the constrictor muscles of the pharynx. THE ALIMENTARY APPARATUS. 225 2. Mucous coat, continuous with that of the various open- ings. It is covered in its upper part with columnar ciliated epithelium, as low as the floor of the nares, below which it is squamous. It contains numerous racemose glands, crypts, and lymphoid structure similar to the tonsils, a mass of which, be- tween the Eustachian tubes, has been called the "pharyngeal tonsil." FI&. 115. Pharynx laid open from behind: 1, styloid process; 2, body of oc- cipital; 3, sxptum nasi; 4, middle turbinated bone; B, posterior naria; 6, inferior turbinated bone; 7, soft palate; 9, uvula; 10, tonsil; 11, back of tongue; 12, epiglottis; 13, arytasno-epiglottidean fold; 14, tip of arytsenoid cartilage; IS, oesophagus; 16, back of cricoid cartilage. 3. Muscular coat, consists of the three pharyngeal con- strictors, the palato-pharyngeus, and the stylo-pharyngeus. {Vide Muscles.) Arteries are from the inferior palatine, pharjmgeal, and thyroid arteries. Nerves are branches of the pneumogastric, glosso-pharyngeal, and sympathetic. The cesophagus, or gullet, is a musculo-membranous tube about nine inches long and less than one inch in diameter, flat- tened from before backward, and extending from the pharynx to the stomach or from the level of the fifth cervical to the ninth dorsal vertebra. 226 HUMAN ANATOMY. Relations. — In the neek the trachea is in front, the common carotids on either side, and in the chest the pericardium, the left carotid, left sub- clavian, the aorta, and the left bronchus are in front; the longus colli muscle and the intercostal vessels and the vertebral column are behind. The pleura covers it laterally, the descending portion of the arch of the aorta lying on the left and the vena azygos major on the right. Its structure consists of three coats: — Muscular coat, consisting of two layers, the longitudinal and the circular, continuous with the inferior constrictor; Fig. 116. 1, left hypochondriac region; 2, epigastric; 3, right hypochon- driac; 4, left lumbar; B, umbilical; 6, right lumbar; 7, left iliac; 8, hypogastric; 9, right iliac. Areolar or fibrous coat, connecting the two ; Mucous coat, covered with stratified pavement epithelium, and having beneath it some non-striated muscular fibres, the mus- cularis mucosae. It also contains numerous compound racemose glands, the oesophageal glands. THE ABDOMEN. The abdomen, the largest cavity in the body, is bounded in front and laterally by the abdominal muscles, the lower ribs, the ilii, and above by the diaphragm, below by the brim of the Fig, 117. The abdominal viscera. (Miklei/.) THE ALIMENTARY APPARATUS. 227 pelvis, and behind by the vertebral column, quadratns lumbomm, and psoas muscles. It is lined throughout by peritoneum, inclos- ing the greater portion of the alimentary canal. It has six openings, as follows: — Aortic opening, for the vena azygos, aorta, and thoracic duct; CEsophageal opening, for the oesophagus and pneumogastric nerves; Quadrate opening, for the vena cava inferior; Umbilicus, in front, for the foetal umbilical vessels; Femoral canal {vide femoral hernia) ; Inguinal canal, for the round ligament in the female, and the sper- matic cord in the male. Eegions op the Abdomen. — Por convenience of study, the cavity of the abdomen is divided into nine regions by four imag- inary lines, two circular lines drawn around the body, one at the lower margin of the thorax (Leidy) or on a level with the car- tilages of the ninth ribs, the second at the highest point of the crest of the ilium; and two vertical lines, drawn from the car- tilage of the eighth rib on each side through the centre of Pou- part's ligament, or from the anterior inferior spinous process of the ilium, drawn upward (Leidy). These regions are named as follows: — Right hypochondriac, Right lumbar, Right inguinal (iliac). Epigastric region, Umbilical region. Hypogastric region. Left hypochondriac. Left lumbar. Left inguinal (iliac). Right Hi/pocliondriao. Hepatic flexure of colon, right lobe of liver, gall-bladder, and upper part of right kidney. Right Lumbar. Greater part of right kidney, ascending co- lon, and portions of the small intestine. Contents of Regions. Epigastric. Left lobe of liver, lobulus Spigelii, greater part of stom- ach, duodenum, and pancreas, portions of the kidneys and su- prarenal capsules, vena cava, aorta, tho- racic duct, semilunar ganglia. VnibiUcal. Transverse portions of colon and duode- num, jejunum, and ileum, part of mesen- tery and great omen- t u m , reeeptaculum chyli, and portions of both kidneys. Left HypocJiondi-iac. Splenic flexure of colon, spleen, tail of pancreas, splenic end of stomach, and upper part of left kidney. Left Lumbar. Part of left kidney, descending colon, some convolutions of small intestine, and part of the omentum. 228 HUMAN ANATOMY. BiffJit Inguinal (Iliac). Hypogastric. Left Inguinal (Iliac). Eight ureter, CEecum, Portions of the small Left ureter, sigmoid spermatic vessels, and intestine under certain flexure of colon, and appendix cseci. circumstances, the spermatic vessels. uterus (pregnant), bladder (distended ) , sometimes the caecum, sigmoid flexure, and appendix. Peritoneum. — The peritoneum is a closed serous sac, its parietal layer lining the cavity of the abdomen, its visceral layer inclosing more or less completely all the abdominal and pelvic viscera. It is not in all cases a closed sac, for in the female it is continuous with the mucous membrane of the Fallopian tubes. It consists essentially of tvfo sacs of unequal size, the greater and lesser peritoneal sacs, united by a central constriction — ^the fora- men of Winslow: — The greater sac is located in front of the viscera, one layer lining the internal abdominal wall, the other reflected upon the viscera. Its cavity is known as the greater peritoneal cavity. The lesser sac covers the upper part of the posterior abdom- inal wall, and is reflected upon the posterior surface of the liver and stomach. Its cavity is called the lesser peritoneal cavity. It also gives off three processes, one of which is a broad, loose fold, — the great omentum, — passes downward from the greater curvature of the stomach between the two layers of the greater sac, and is reflected upon itself back to the under surface of the transverse colon. The foramen of Winslow is a narrow canal, large enough to admit one finger, between the greater and lesser peritoneal cavities, located behind the right border of the lesser omentum, and formed by the hepatic and gastric arteries, constricting the sac at this point as they ascend from the cceliac axis. It is bounded as follows: — In front, by the lesser omentum, containing the hepatic artery, portal vein, duodenum, and the ductus communis choledochus; Behind, by the right crus of the diaphragm and the inferior vena cava; Above, by the lobus Spigelii; Below, by the hepatic artery. Reflections. — The reflections of the peritoneum viewed in an antero-posterior section (the greater and lesser sacs together) may be traced as follows : From the diaphragm it is reflected to the upper surface of the liver. Enveloping this organ, it then presents a doubling or fold — the gastro-hepatic omentum — ex- tending downward from the transverse hepatic fissure to the THE ALIMENTARY APPARATUS. 229 lesser curvature of the stomach. Inclosing the stomach, it is reflected upon itself in front of the intestines as a broad apron — the great omentum — ^making a quadruple fold of peritoneum. The two layers then embrace the transverse colon, unite, and pass back to the vertebral column, forming the transverse mesocolon. From here the layers separate, the upper one ascend- FiG. 118. D, diaphragm; L, llyer; S, stomach; P, pancreas; D, duodenum; C, colon; I, small Intestine; B, bladder; R, rectum; 3, posterior surface of liver; 4, foramen of Winslow; 5, great omentum; 6, lesser omentum; 7, mesocolon; 8-9, lesser cavity of peritoneum; 10, mesentery; 11, recto-vesical told. ing in front of the pancreas to the starting point. The lower layer descends in front of the duodenum aorta, incloses the small intestine (forming the mesentery proper), is reflected upon the rectum (forming the mesorectum) and the bladder, and ascends upon the anterior abdominal wall to the starting point. In the female, from the rectum it envelops the uterus and upper part of the vagina before reaching the bladder. 230 HUMAN ANATOMY. In addition to the folds seen in the antero-posterior section, the peritoneum passes between the various organs, and also lat- erally to the sides of the abdominal and pelvic cavities. In this manner are formed the right, left, and suspensory ligaments of the liver, the suspensory ligament of the spleen, the broad liga- ment of the uterus, and the three great ligaments or omenta of the stomach — the gastro-hepatic, gastro-splenic, and gastro-colic (already described) and the mesenteries. The latter, the mesenteries, include the mesentery proper, mesocaecum; ascending, transverse, and descending mesocolon; sigmoid mesocolon, and mesorectum. The mesentery proper is a broad fold, reflected from the vertebra around the jejunum and ileum. Its base, attached ob- liquely from the left side of the second lumbar vertebra to the right iliac region, measures about six inches, while its expanded extremity is quite considerable. In the male the recto-vesical folds, one on each side, pass from the rectum to the bladder, including between them the recto-vesical pouch. In the female, however, this pouch is divided into two— the recto-uterine and vesico-uterine pouches, the recto-vesical folds being called recto-uterine and vesico- uterine folds. Folds of peritoneum (superior false ligaments of the bladder) ascend from the bladder to the umbilicus, in- closing the remains of the f cetal urachus and hypogastric arteries, and also a fold on each side follows the course of the epigastric arteries toward the umbilicus, dividing the inguinal region into the internal, middle, and external inguinal fossae. The peri- toneum at the external inguinal fossa (corresponding to the internal abdominal ring) is continuous in the male foetus with the tunica vaginalis testis, and in the female forms a blind sac about the round ligament — the canal of Nuch. Viscera Partly Invested hy Peritoneum. Vagina — upper part; Bladder — posterior wall; Duodenum — descending and transverse portions; CtECum ; Colon — ascending and descending; Rectum — middle portion. Viscera Entirely or Almost Entirely Covered. Stomach, Sigmoid flexure, Spleen, Rectum — upper part. Ileum, Liver, Jejunum, Uterus, Colon — transverse, Ovaries. THE ALIMENTAKY APPARATUS. 231 Viscera Without Peritoneal Investment, Bladder — except posterior wall; Rectum — lower third; Vagina — lower part of posterior wall; Pancreas ; T Suprarenal capsules ; > partly covered anteriorly. Kidneys. J THE STOMACH. The stomach is a musculo-membranous sac, pyriform in shape, situated below the diaphragm in an oblique position across the upper abdomen, where it is held in position by the lesser omentum and the gastro-splenic ligament. It measures about nine to twelve inches long, four to five inches in its greatest diameter, and has a capacity of from one to two quarts. It occupies the epigastric, right and left hypochondriac regions, and is ia relation above with the diaphragm and liver, below with the transverse colon, in front with the abdominal walls, and behind with the pancreas. When distended the stomach rotates on its long axis and the greater curvature points somewhat for- ward. It presents for examination a greater and lesser extrem- ity, greater and lesser curvatures, anterior and posterior surfaces, and cardiac and pyloric orifices. The greater extremity, or fundus, occupies the left hypo- chondriac region, in contact with the spleen, with which it is attached by the gastro-splenic omentum, and behind the lower rib. The lesser or pyloric end is smaller and lies in contact with the under surface of the liver and the wall of the abdomen ia the right hypogastric region. The greater curvature extends between the pyloric and oesophageal orifices, along the lower border of the stomach, and to it is attached the great omentum. The lesser curvature extends between the same two points along the superior border of the organ. The (esophageal or cardiac orifice occupies the highest part of the stomach, behind the left seventh costal cartilage, and receives the oesophagus. The pyloric orifice occupies the right extremity, and opens into the duodenum, being protected by a muscular valve, the pylorus, or pyloric valve. The anterior surface is in contact with the under surface of the left lobe of the liver, the abdominal walls, and the dia- phragm. The posterior surface is in contact with the peritoneum of the diaphragm, solar plexus, the pancreas, and the abdominal vessels. 232 HUMAN ANATOMY. The structure of the stomach consists of four coats, a serous, muscular, fibrous or areolar, and mucous. The serous coat is derived from the peritoneum, which com- pletely covers it, except along the lesser and greater curvature, where the vessels and nerves enter. The muscular coat consists of three sets of fibres : — The longitudinal, the most superficial, are continuous below with the longitudinal fibres of the small iatestine below, and the oesophagus above; The circular fibres, the second layer, are most abundant at the pyloric extremity, where they form the pyloric valve; The oblique fibres are distributed over -both surfaces, pass- ing obliquely from right to left and left to right. The areolar or submucous coat connects the muscular with the mucous layer, and is sometimes named the vascular coat. The mucous membrane is of a pale pinkish-ash color, thick- ened toward the pylorus, where it presents numerous rugse, or pleats, and at the pyloric end it helps to form the pyloric valve. It is lined throughout with columnar epithelium, and is studded with three kinds of minute tubes, the gastric follicles, and len- ticular glands. The gastric follicles consist of two kinds, the pyloric and the peptic glands, the former most abundant at the pyloric end and the latter distributed all over the surface of the stomach. The pyloric or mucous glands consist each of from two to four blind tubes opening into a common duct, and lined through- out by columnar epithelium. The peptic glands are similar in structure, but have a much shorter duct, and contain in addition peculiar large, spheroidal, granular peptic cells. The lenticular or simple solitary glands are small masses of lymphoid tissue scattered throughout the connective-tissue framework of the stomach between the gastric follicles. The arteries are derived from the gastric, pyloric, and right gastro-epiploic branches of the hepatic artery, and the left gastro- epiploic and vasa brevia branches of the splenic artery {vide Arterial System). The veins terminate in the portal, superior mesenteric, and splenic veins. The nerves are derived from the gastric plexuses (Auer- bach's and Meissner's, in the muscular and submucous coats, respectively), formed by the terminal branches of the right and left pneumogastric, and the branches of the coeliac plexus, an ofEshoot of the solar plexus of the sympathetic. THE ALIMENTARY APPARATUS. 233 THE SMALL INTESTINE. The small intestine is a convoluted tube about twenty to twenty-five feet in length, for the chylification of the food, occu- pying the lower and central portions of the abdominal and pelvic cavities, and held in position to the spinal column by the mesen- tery. It is divided into three portions — ^the duodenum, jejunum, and ileum. The duodenum, so called from being about twelve fingers' breadth in length, is about eight to tett inches in length. It consists of four portions, from the position of its course — ascend- ing, descending, transverse, and terminal ascending: — The -first or ascending portion is about two inches in length, and ascends to the neck of the gall-bladder. It is completely invested by peritoneum for about an inch. The second or descending portion, about three inches in length, descends in front of the right kidney as far as the third or fourth lumbar vertebra, and is overlapped in front by the head of the pancreas, and into its posterior aspect the duct of the pancreas and the common biliary duct open by a common orifice. The third or transverse portion is attached to the crura of the diaphragm and the vessels in front of the vertebral column, passes behind the transverse mesocolon, and has the pancreas above it, and the superior mesenteric blood-vessels cross from beneath the latter between the two or over the duodenum. The fourth or terminal ascending portion runs upward and forward to the duodeno-jejunal flexure. The arteries are derived from the inferior pancreatico- duodenal branch of the superior mesenteric and the superior pancreatico-duodenal branch of the gastro-duodenal, a branch of the hepatic. The veins terminate in the superior mesenteric and splenic veins. The nerves are from the solar plexus. The jejunum, named from jejunus, empty, includes the upper two-fifths of the small intestine, is continuous above with the duodenum and below with the ileum. It occupies chiefly the left iliac and umbilical region. The ileum, so called from its twisted condition, includes the remaining three-fifths, is continuous above with the jejunum and below with the caput caecum of the large intestine. It occu- pies the right iliac, hypogastric, and umbilical regions. The structure of the small intestine consists of four coats — the serous, muscular, fibrous or areolar, and mucous : — The serous coat is the peritoneal covering; The muscular coat consists of two sets — a, longitudinal and a circular; 234 HUMAN ANATOMY. The areolar or submucous coat connects the muscular with the mucous coat; The mucous coat is thinner and redder than that of the stomach, ana is thrown into numerous transverse folds — the valvulse conniventes— most numerous in the upper part; they diminish as it descends and finally disappear in the ileum. They increase the secreting and absorbing surface of the mucous mem- brane and retard the passage of the food. The mucous memlrane also contains the villi and four kinds of glands: — Simple follicles, or crypts of Lieberkiihn; Duodenal glands, or Brunner's glands; Solitary glands; Agminate, or Peyer's glands. The villi are minute vascular projections of the mucous membrane scattered throughout the surface of the small intes- tine. Their structure consists of a pouchlike termination of a lacteal in the centre surrounded by a minute plexus of capillary vessels inclosed in a basement membrane and covered with co- lumnar epithelium. The simple follicles^ or crypts of LieberTciihn, are scattered throughout the mucous membrane of the entire small intestine. They consist of, minute tubes of basement membrane, lined with columnar epithelium and surrounded by a capillary net-work. The duodenal or Brunner's glands are distributed to the duodenum and jejunum only. They are largest and most numer- ous in the vicinity of the pylorus. They are composed of tubular alveoli, lined by epithelium, and having a small duct opening on the mucous membrane. The solitary glands are distributed throughout the small intestine, being most numerous in the last portion of the ileum. They consist of lymph follicles, and communicate with the lac- teal system by means of lymph spaces. Peyer's glands, or Peyer's patches, consist of an aggrega- tion of the solitary glands into oval groups of twenty or thirty along the small intestine at a point opposite the attachment of the mesentery. The patches are about fifteen to thirty in number, each measuring about one-half to two inches in length and one-half inch in breadth. Their axes are parallel with the length of the intestine. In the duodenum they are few and small in size, and the mucous membrane of the valvulas conni- ventes over them is reduced in size and much distorted. The arteries are derived from the pyloric, pancreatico- duodenal, and superior mesenteric. They reach the intestines THE AllMENTAKY APPAEATUS. 235 inclosed in and along the course of the mesenteries. In the in- tervals of the coats they form three vascular nets, which supply the serous, muscular, and mucous coats. The veins accompany the arteries and join the portal vein. The lymphatics follow the course of the superior mesenteric vessels to enter the mesenteric glands. The nerves are derived from the solar plexus of the sympa- thetic, THE LARGE INTESTINE. The large intestine is about five feet long and extends from the ileum to the anus. It commences in the right iliac fossa, ascends through the right lumbar and right hypochondriac regions, passes trans- versely between the epigastric and umbilical regions to the left hypochondriac region, where it descends through the left hypo- chondriac, lumbar, and iliac regions, and through the pelvis on its posterior wall to terminate at the anus. It consists of three divisions: — 1. Csecum, 2. Colon, 3. Kectum. ' Ascending, Transverse, Descending, Sigmoid flexure. The C^cum is a blind pouch measuring about two and one- half inches in every diameter, lying free in the right iliac region. It has opening into its lower back part the appendix vermi- formis, and into its inner back part the ileum, guarded by the ileo-csecal valve. The appendix vermiformis is a long, narrow, twisted tube, the rudiment of the prolonged csecum in all marmnalia. It ter- minates in a blunt extremity. Its mueoiis membrane is contin- uous with that of the csecum, and contains many solitary glands. The ileo-cwcal valve, or valve of Bauhin, protects the opening of the ileum into the csecum, and consists of two valve-like semi- lunar folds of mucous membrane, strengthened by bands of cir- cular fibres. The upper one is attached to the junction of the ileum with the colon, the lower one to the junction of the ileum with the csecum. On each side where the folds coalesce, a ridge of mucous membrane continuous for a short distance around the canal, forms the frcena or reiinaciild of the valve. The mucous membrane on either side of the valves corresponds 236 HUMAN ANATOMY. to that of the large and small intestines respectively, being cov- ered with villi on the side toward the ileum, and being destitute of villi, and containing numerous tubular glands or crypts of Lieberkiihn, on the side toward the cascum. This difference oc- curs abruptly at the free margin of the valves. The colox consists of four portions — ascending, transverse, and descending colon, and sigmoid flexure: — The ascendincj portion begins opposite the ileo-cseeal valve, ascends through the right lumbar and hypochondriac regions, and terminates beneath the under surface of the liver at the hepatic flexure in the transverse portion. Its posterior surface is desti- tute of peritoneum, and is attached to the quadratus lumborum muscle by loose areolar tissue. The transverse portion, or transverse arch of the colon, passes from the hepatic flexure, through the adjoining portions of the epigastric and umbilical regions, to terminate at the splenic flexure in the descending portion. It is attached by transverse mesocolon, and is most movable portion of the colon. The descending portion begins at the splenic flexure, de- scends through the left lumbar and iliac regions to terminate in the sigmoid flexure. Like the ascending colon, its posterior surface is destitute of peritoneum. The sigmoid flexure (omega loop) is a narrow, twisted por- tion of the colon, occupying the left iliac fossa, between the descending portion of the colon and the rectum. It is held in position by the sigmoid mesocolon. It terminates opposite the third sacral vertebra. The rectum — the terminal portion — extends from the sig- moid flexure to the anus. It is four to six inches in length, not sacculated, but club-shaped, with its large extremity downward, and consists of two portions — superior and inferior: — The lower four inches, formerly described as the upper por- tion of the rectum, rest upon the sacral plexus of nerves and the pyriformis muscle ; The superior portion, about three and three-quarters inches, is in relation in the male with the bladder, ]irostate gland, and vesiculse seminalis; in the female with the uterus and vagina, being adherent to the latter ; The inferior portion, about one and one-quarter inches, turns backward and terminates in the anus. It is supported by the levator ani muscle, and is surrounded by the internal and external sphincters of the anus. The slruciure of the larije intestine con- sists of four coats — serous, muscular, cellular, and mucous: — The serou.t coat, derived from the peritoneum, invests lom- pk'toly the transverse portion and sigmoid portion of the colon. THE ALIMENTARY APPARATUS. 237 but only partially the others, being absent in the posterior sur- faces of the ascending and descending portions, and on the superior portion of the rectum. The inferior portion of the rectum is without peritoneal covering. The appendices epiploicw are folds of serous membrane filled with fat and attached chiefly to the transverse colon, depending from it. The muscular coat consists of longitudinal fibres arranged into three flat bands, shorter by nearly one-half than the intes- tine itself, and the circular, distributed more evenly, but accumu- lated at points, producing with the former a sacculated condition. The cellular coat connects the mucous with the muscular coat beneath. The mucous coat is smooth, destitute of villi and valvule conniventes, and thrown into numerous folds, two to four of which, situated in the rectum, have received the name of Hous- ton's valves. It contains crypts of Lieberkiihn, more numerous here than in the small intestine, and solitary glands scattered throughout, but most numerous in the appendix and csecum, where some of them are accumulated into Peyer's patches. The arteries of the csecum and colon are from the mesen- teric. The lymphatics empty into the mesenteric glands, and the nerves are derived from the mesenteric plexus of the sym- pathetic system. The arteries of the rectum are the hsemorrhoidal branches of the inferior mesenteric, internal iliac, and internal pudie. The veins form the hsemorrhoidal plexus, and empty into the inferior mesenteric and internal iliac veins. The lymphatics go to the sacral and lumbar glands, and the nerves are from the hypogastric plexus of synipathetie and con- tiguous spinal nerves. THE PANCREAS. The pancreas is an oblong compound racemose gland, about six to eight inches in length, one and one-half inches in breadth, and one-half to one inch in thickness, situated across the back part of the epigastric and left hypochondriac regions. Its weight varies from two to six ounces. It consists of a head, body, and tail: — The head, or right extremity, is received into the concavity of the duodenum. The lesser pancreas, a detached portion of the gland, lies behind it. The tail, or lesser end, terminates above the left kidney and suprarenal capsule, in contact with the spleen. 238 HUMAN ANATOMY. The lody is in relation in front with the stomach and trans- verse mesocolon ; behind it rests upon the first lumbar vertebra, having interposed the crura of the diaphragm, vena cava, left renal vein, aorta, superior mesenteric artery and vein, the in- ferior mesenteric vein, and commencement of portal vein. The splenic artery and vein are lodged in a groove on its upper border. The pancreatic duct, or canal of Wirsung, runs throughout the substance of the organ, to emerge at the head, and open into the duodenum by an orifice in common with the ductus com- munis choledochus. The lesser pancreas, when it exists, empties by the ductus pancreaticus minor. The structure resembles that of the salivary glands, but is softer and looser. Arteries are from the splenic and pancreatico-duodenal branches of the hepatic and superior mesenteric. The veins join the splenic or superior mesenteric. The nerves are from splenic plexus of the sympathetic. The lymphatics empty into the lumbar glands. THE LIVER. The liver is the largest gland in the body, measuring in its transverse diameter from ten to twelve inches, and its antero- posterior six to seven, and its thickest part about three inches, and weighing about from three to four pounds. It occupies the upper part of the abdominal cavity, and the right hypo- chondriac, epigastric, and a portion of the left hypochondriac regions. Its upper surface is convex and rests against the dia- phragm and a small portion of the abdominal parietes in front. Its lower surface is in contact with the duodenum and stomach, the right kidney and suprarenal capsules, and the hepatic flexure of the colon. It is divided by the longitudinal fissure into the right and left lobes. The liver has five fissures, five lobes, five ligaments, five sets of vessels, and is inclosed in a fibrous coat, continuous at the transverse fissure with the capsule of Glisson. It is also invested by the peritoneum, except at the attachment of the coronary ligament. Structure. — The liver is made up of lobules, which are small, granular bodies about one-fifteenth of an inch in diameter, held together by delicate connective tissue and the branches of the five sets of vessels (to be described) and nerves, the whole being inclosed in a serous and fibrous coat. Each lobule is made up of a mass of polyhedral, nucleated cells, inclosed in a capillary plexus derived from the hepatic THE ALIMENTARY APPARATUS. 239 artery and portal vein, and giving origin to the hepatic vein and biliary duct. The five fissures of the liver are all situated on its under surface. They separate the five lobes from one another. They are : — 1. Longitudinal fissure extends from before backward, from the notch in front to the posterior border. It is sometimes called the umbilical fissure, and lodges the round ligament, the remains of the foetal umbilical vein. 2. Fissure for the ductus venosus is the posterior portion of the longitudinal fissure, and lodges the remains of the ductus venosus of foetal life. Fig. 119. 1, left lobe ; 2, right lobe ; 8, quadrate lobe ; 4, caudate lobe ; 6, Spigelian lobe ; 6, hepatic artery ; 7, portal rein ; 8, fissure of ductus venosus ; 9, gall- bladder : 10, cysti..- duct; 11, hepatic duct ; 12, fissure for ^ ena cava ; IH, vena cava ; 1^ right inferior phrenic Ttin ; 15, hepatic vein ; 16, right renal vein ; 17, left rend vein, 3. Transverse fissure, or portal fissure, crosses the inferior surface of the liver transversely and joins the longitudinal. It transmits the portal vein, hepatic artery and nerves, and the hepatic duct and lymphatics. 4. Fissure for the gall-bladder is a shallow depression be- neath the right lobe, running parallel with the longitudinal fissure in front. 5. Fissure for the vena cava runs obliquely upward along the inferior surface near its posterior margin to the left side, joining the fissure for the ductus venosus behind, and separated from the transverse fissure in front by the lobulus caudatus. It lodges the inferior cava, which, withia this fissure, receives the hepatic veins. Missing Page Missing Page 242 HUMAN ANATOMY. and the cystic, about one inch in length. It empties into the descending portion of the duodenum in common with the pan- creatic duct, about three and one-half inches below the pylorus. THE DUCTLESS GLANDS. The following group includes the glands without ducts, of unknown function, which resemble each other in structure: — Spleen, Thymus, Thyroid, Suprarenal capsules. Pituitary body, Glandula coeeygea, Grlandula intercarotica. The Spleen. — The spleen is a soft, very vascular, sponge- like organ, situated deeply in the left hypochondriac region. It varies much in size and weight, measuring about five inches in length, three in width, one and one-half ia thickness, and weigh- ing between seven and ten ounces. Outer surface, smooth and convex, corresponds to the ninth, tenth, and eleventh ribs, and is adapted to the inferior surface of the diaphragm, to which it is connected by the suspensory ligament. Inner surface is concave and adapted to the cardiac end of the stomach, to which it is attached by the gastro-splenic omentum. Hilus is a vertical fissure on the concave inner surface, ad- mitting the passage of blood-vessels, lymphatics, and nerves. The structure consists of two coats — a serous and fibro-elastic coat, inclosing in its interior the spleen pulp. Serous coat, derived from the peritoneum, covers the entire organ, except at the hilus, where it forms the gastro-splenic omentum. Fibro-elastic coat, or tunica propria, surrounds the organ, and from the hilus and periphery sends numerous fibrous bands, or trabeculce, into the substance of the organ, dividing it into small trabecular spaces, or areolce. Splenic substance, or spleen pulp, is a soft, reddish-brown mass, consisting of a fine reticulum of connective-tissue cor- puscles, inclosing red and white blood-corpuscles, nucleated and non-nucleated cells, granular matter, etc. Malpighian corpuscles, or bodies, are spheroidal hyper- plasise of lymphoid tissue from the outer coat of the arterioles. They are not encapsuled, vary from one-sixtieth to one-twenty- fifth of an inch, and are visible ia the fresh specimens to the naked eye. THE ALIMENTARY APPARATUS. 243 Splenic artery, remarkable for its size and toriniosity, divides in the hilns into four or five branches, each distributed to a segment, terminating in a capillary plexus without anastomoses, or opening directly into the areolae of the splenic pulp. Splenic vein commences in the same manner as the arterioles end, and empties into the portal vein. The smaller veins anas- tomose freely. Nerves are from the splenic plexus, formed from the right pneumogastric nerve, and the left semilunar ganglion of the solar plexus. Thteoid Gland. — The thyroid is a vascular, glandlike body, situated on the sides of the upper part of the trachea, and con- sists of two lateral lobes connected by a transverse portion, the isthmus. A third lobe — ^the pyramid — sometimes arises from the left lobe or upper margin of the isthmus. Levator glandules thyroidecB are muscular bands sometimes found passing from the isthmus to the body of the hyoid bone. Structure. — ^This is similar to other glands, being made up of a capsule and radiating septa inclosing alveoli — ^the closed vesicles — each of which is lined with one layer of columnar epithelium, and contains more or less viscid, transparent fluid — the colloid substance. The closed vesicles are abundantly sup- plied with blood by meshes of capillaries, while penetrating the septa are lymphatic net-works, and lymph sinuses are found in the tissue between the vesicles and septa. Arteries are the superior thjrroid, a branch of external carotid, and inferior thyroid, a branch of the thyroid axis, and sometimes a branch from the arch of the aorta or innominate artery, the middle thyroid, or arteria thyroidea ima. All the vessels anastomose freely. Veins form plexus about the gland and give off the superior and middle thyroid to internal jugular, and inferior thyroid to innominate vein. Nerves, from middle and inferior cervical ganglia and from pneumogastric. Thymus Gland. — The thymus gland is a temporary organ of unknown function which attains iis full size at the end of two years and at puberty has almost disappeared. It occupies the upper part of the anterior mediastinum, and is in relation in front with the sternum, and below with the pericardium, aorta, left innominate, and trachea. It is a flat, triangular body, com- posed of a pair of lateral unequal lobes, about two inches in length, one and one-half inches at the widest part, and one- quarter inch thick, and weighing about one-half ounce. 244 HUMAN ANATOMY. Structure. — It is composed of a framework of fibro-connec- tive tissue, consisting of a capsule and septa, and the gland sub- stance, which is divided into lobes, these into lobules, and these into units or follicles. The follicles vary in shape, present for study a cortex and medulla, and consist of adenoid tissue. The cortical meshes of the reticulated adenoid tissue are filled with lymph-corpuscles, while in the medulla the meshes are less numerous and filled with large endothelioid plates and giant cells. The capillary blood-vessels surround the follicles. Arteries are from superior and inferior thyroid and internal mammary. Veins join thyroid and left innominate veins. Nerves are from sympathetic and pneumogastric. SupEAEENAL CAPSULES. — The Suprarenal capsules are two small, triangular bodies, situated upon the upper and front part of either kidney. They measure from one and one-quarter to two inches in length, and less in breadth, two to three lines in thickness, and weigh about two drachms. Structure. — Like the kidneys, they consist of a cortical and medullary portion, inclosed in a capsule which sends septa into the substance of the body. The cortex is composed of three zones of epithelial cells — outer, middle, and inner — the middle being the largest. The medulla consists of streaks of small, transparent cells, separated by connective tissue and capillaries. These streaks are continuous with the inner zone of the cortex. The nerve supply is rich, consisting of non-medullated fibres connected with small ganglia. Relations. — The inferior concave border rests upon the upper sur- face of the kidney. The inner border rests against the inferior vena cava on the right side, the aorta on the left, and is in relation with semi- lunar ganglion and great splanchnic nerves. The anterior surfaces touch on the right the under surface of the liver and on the left side the pancreas and spleen. The posterior surface lies upon the crus of the diaphragm, about opposite the tenth dorsal vertebra;. Arteries are suprarenal from the aorta, the renal, and phrenic arteries. Veins on the right join -^'ena cava, on left renal vein. Nerves, from renal and solar plexus. PiTUlTAET BODY (hi/pophysis cprebii) is a small, vascular mass on the inferior surface of the cerebniui, but resembling iu structure the ductless glands. THE ALIMENTARY APPARATUS. 245 Structure. — The upper lobe is part of the central nervous system. The lower and larger lobe has a fibrous capsule which sends processes into the interior, forming minute septa; these divide and reunite around spaces called alveoli. The alveoli con- tain epithelial cells of various shapes, between which are small irregular cells having flattened nuclei. The glandula cocctgba, lyiag near the tip of the coccyx and the intekcaeotica, at the angle of bifurcation of the com- mon carotid, have been recently included among the ductless glands. They were both discovered by Luschka. The frame- work of their bodies is identical with that of other glands, except that the septa contain non-striped muscular tissue. The alveoli are filled with gland substance, consisting of connected masses of epithelial cells, having in. their centre a twisted capillary blood-vessel. VOCAL AND RESPIRATORY APPARATUS. The larynx is a musculo-cartilaginous box at the top of the trachea, below the root of the tongue and the hyoid bone, and is the organ of the voice. It is composed of cartilages connected by ligaments, provided with muscles, blood-vessels, and nerves, and lined with mucous membrane. The cartilages are nine in number, three single and three pairs: — Thyroid, Cricoid, Epiglottis, Two arytenoid. Two comieula laryngis. Two cuneiform. The thyroid, the largest cartilage, consists of two quadri- lateral halves, united in front in the median line, or entering angle of the thyroid, the upper part of which is the pomum Adami. The outer surface is marked by an oblique ridge for the attachment of muscles. The inner surface is smooth and cov- ered by mucous membrane, and has in front attached the true and false vocal cords. The posterior angles are prolonged into superior and inferior horns, the superior giviag attachment to the thyro-hyoid ligament, the inferior articulating with the sides of the cricoid cartilage. The cricoid cartilage resembles a seal ring, narrow in front, the back part of the upper border articulates with the arytenoid cartilage, and on each side externally are two facets for the articulation of the inferior horns of the thyroid. The epiglottis is a spoon-shaped, fibro-cartilaginous plate, large above, its narrow inferior extremity is prolonged and attached by a band of fibro-elastic tissue of thyro-epiglottic liga- ment to the thyroid cartilage. It is also attached to the posterior surface of the hyoid bone by the hyo-epiglottic ligament. The anterior or lingual surface has three reflections of mucous mem- brane between it and the tongue, called the glosso-epiglottidean ligaments. The arytenoid cartilages each resembles the mouth of a pitcher, from which they are named. They are smaller than the other two, and are situated on the summit of the cricoid cartilage posteriorly. They are three-sided, the apex extends backward, and is surmounted by the supra-arytenoid, comieula laryngis, or cartilages of Santorini. (246^ VOCAL AND RESPIRATORY APPARATUS. 247 The posterior surface has attached to it the arytenoid mus- cle. The anterior surface has attached to it the thyro-arytenoid muscle and the false vocal cord. The internal surfaces are op- posed to each other and supplied with mucous membrane. Of the three angles at the base, the outer gives attachment to the crico-arytenoid muscle, lateral and posterior. The anterior is prolonged for attachment of the true vocal cord. The cuneiform cartilages, or cartilages of Wrisberg, are two small rod-shaped bodies extending upward from the arytenoid cartilages into the aryteno-epiglottidean fold. Fig. 121. Vertical section of larynx; 1, body of hyold bone; 2, epiglottis; S, thyro-hyoid membrane; 4, great cornu of hyoid; B, false vocal cord; 6, thyro-hyoid ligament; 7, ventricle of larynx; 8, thyro-Iiyoid membrane; 9, true vocal cord; 10, aryteno-epiglottidean fold; 11, thyroid cartilage; 12, superior cornu of thyroid; 14, arytenoid muscle; 16, arytenoid cartilage; 18, cricoid cartilage. The cartilages of the larynx, with the exception of the epiglottis, are composed of true cartilage. The LIGAMENTS of the LARYNX are divided into two sets — the extrinsic, those connecting the epiglottis and thyroid car- tilage with the hyoid bone, and the intrinsic, those which connect the various cartilages together. The extrinsic consist of three : — 1. Thyro-hyoid membrane, connecting the upper border of the thy- roid cartilage with the inner surface of the hyoid bone; 2 and 3. The two lateral thyro-hyoid ligaments, fibro-elastie cords connecting the superior horns of the thyroid cartilage with the extrem- ities of the great horns of the hyoid bone. They contain a small nodule, tke cartilago-triticea. 248 HUMAN ANATOMY. The intrinsic ligaments are sixteen in number, as follows : — Hyo -epiglottic ligament; Crico- thyroid membrane ; Two crioo-thyroid capsular ligaments; Two crico-arytenoid ligaments ; Two crico-arytenoid capsular ligaments; Two superior thyro-arytenoid ligaments (in false cords) ; Two inferior thyro-arytenoid ligaments (in true cords) ; Thyro-epiglottio ligament; Three glosso-epiglottic folds. The hyo-epiglottic ligament is a fibro-elastic band, connect- ing the anterior surface of the epiglottis with the upper border of the hyoid bone. The ligaments connecting the cricoid to the thyroid cartilage are three also — ^the crieo-thyroid ligament, cap- sular ligaments, and synovial membranes. The crico-thyroid membrane, a yellow, elastic band, connects the adjacent margins of the cricoid and thyroid cartilages, and extends from the upper border of the cricoid cartilage to the lower margin of the true vocal cords. The two crico-thyroid capsular ligaments surround the ar- ticulations between the inferior horns of the thyroid and the cricoid cartilage. The snyovial membrane lines the capsular ligaments, form- ing a true enarthrodial joint. The ligaments of the epiglottis are the thyro-epiglottic, the hyo-epiglottic, and the three glosso-epiglottic folds of mucous membrane before described. The superior aperture of the larynx is a triangular opening with the apex in front. It is bounded behind by the apices of the arytenoid cartilages and eomiculse laryngis, ia front by the epiglottis, and laterally by the aryteno-epiglottidean folds. Prom this, as its superior boundary, the cavity of the larynx extends as low as to the lower border of the cricoid cartilage. The true vocal cords and the thyro-arytenoid muscle divide it into two parts, the narrow fissure between the two cords being called the glottis, or rima glottidis. The superior or false vocal cords are two folds of mucous membrane inclosing the superior thyro-arytenoid ligaments. The inferior or true vocal cords are two folds of mucous membrane inclosing the inferior thyro-arytenoid ligaments, com- posed of elastic tissue, from the sides of the upper border of the cricoid cartilage, extending upward to the bases of the arytenoid cartilages, and lower portion of the angle of the thyroid. Their upper margins correspond to the lower edges of tie ventricles of the larynx. VOCAL AND RESPIRATORY APPARATUS. 249 ♦ The ventricle of the larynx is a deep fossa on either side of the larynx, bounded above by the false vocal cords, below by the inferior or true vocal cords, and externally by the thyro- arytenoideus muscle. The sacculus laryngis, or laryngeaV pouch, is a membranous sac lined with mucous membrane, opening into the anterior portion of the ventricle of the larynx. Its inner or laryngeal surface is covered by the aryteno-epiglottideus inferior, and the outer side by the thyro-epiglottideus and thyro-arytenoideus muscles. It is compressed by these muscles, discharging its mucous secretion upon the true vocal cords. Muscles. — The intrinsic muscles of the larynx consist of two sets, five connected with the vocal cords and rima glottidis, and three with the epiglottis. The five muscles of the vocal cords are: — Crico-ihyroid. — Origin^ from the front and sides of the cricoid cartilage; insertion j into anterior border of the inferior comua, and lower margin of the thyroid cartilage; action, elongates and renders tense the vocal cords; nerve, superior laryngeal. Crico-arytwnoidev^ Posticus. — Origin, from the sides and posterior surface of the cricoid cartilage; insertion, into the outer angle of the base of arytenoid cartilage; action, rotates the arytenoid cartilages outward, opening the glottis, and rendering tense the vocal cords ; nerve, recurrent laryngeal. Crico-arytcenoideus Lateralis. — Origin, from upper and outer side of the cricoid cartilage; insertion, in front of the preceding into the outer angle of the base arytenoid; action, rotates the arytenoids inward, closing the glottis; nerve, recur- rent laryngeal. Arytcenoideus. — Origin, from outer border and posterior sur- face of one arytenoid cartilage ; insertion, into the same part of the other — its fibres are oblique and transverse; action, by ap- proximating the arytenoids closes the back part of the glottis; nerves, superior and recurrent laryngeal. Thyro-arytcenoideus. — Origin, from the crico-thyroid mem- brane and lower half of the entering angle of the thyroid car- tilage ; insertion, into anterior surface and base of the arytenoid cartilage — it consists of inferior and superior portions, the for- mer entering into the formation of the true vocal cords ; action, mainly relaxes the true vocal cords by drawing the arytenoids forward ; nerve, recurrent laryngeal. The muscles of the epiglottis are : — Thyro-epiglottideus. — Origin, from the inner surface of thy- roid cartilage; insertion, into the margin of epiglottis and 250 HUMAN ANATOMY. aryteno-epiglottidean fold ; action^ compress the sacculus laryngis and depress the epiglottis; nerve, recurrent laryngeal. Arytceno-epiglottideus superior. — Origin, from apex _ of arytenoid; insertion, into aryteno-epiglottidean folds; action, constricts the superior laryngeal aperture; nerve, recurrent laryngeal. Arytceno-epiglottideus inferior. — Origin, from middle of in- ternal portion of the arytenoid ; insertion, into upper and inner part of epiglottis; action, compresses the sacculus laryngis; nerve, recurrent laryngeal, muscle of Hilton. The lining mucous membrane of the larynx is continuous with that of the pharynx and trachea. It forms the glosso- epiglottic and arytseno-epiglottic folds, adheres tightly to the epiglottis, vocal cords, and the interior of the cricoid cartilage, but is more loosely attached to other parts. It contains numer- ous racemose glands, particularly along the posterior margin of the arytseno-epiglottidean fold, and in front of the arytenoid car- tilages, where they are called the arytenoid glands. Its epithe- lium is of the ciliated, columnar variety below the true vocal cords, and above this point in front as high as the middle of the epiglottis. The other portions are covered by squamous epithelium. The arteries a;re the laryngeal branches from the superior and inferior thyroid, and the crico-thyroid branches of the supe- rior thyroid. The veins join the inferior, middle, and superior thyroid veins. The lymphatics enter the deep cervical glands. The nerves are the inferior or recurrent laryngeal, the supe- rior laryngeal branches of the pneumogastric, and branches from the sympathetic nerve. The superior laryngeal supplies sensation to the larynx. It descends from the inferior ganglion of the pneumogastric, behind the internal carotid at the side of the pharynx, and divides into two branches — ^the internal laryngeal pierces the thyro-hyoid membrane to supply the mucous membrane and arytenoid muscle, the exterrial laryngeal supplies the crico-thyroid muscle. The inferior or recurrent laryngeal, from its origin, winds around the subclavian on the right side and around the arch of the aorta on the left side, and ascends by the side of the trachea to the larynx, of which it is the motor nerve, supplying all the muscles of the larynx except the crico-thyroid, giving off in its course' cardiac, oesophageal, tracheal, and pharyngeal branches, and anastomosing with the superior laryngeal nerve. VOCAL AND RESPIRATORY APPARATUS. 251 TRACHEA AND BRONCHI. The trachea, or windpipe, is a membrano-cartilaginous tube, about four and a half inches in length, three-quarters to one inch in width, extending from the fifth cervical to the upper border of the fifth dorsal vertebra. It divides into two bronchi, the right, about an inch long and nearly at right angles, passing behind the right pulmonary artery on a level with the fifth dorsal vertebra; the left, narrower, about twice as long, passes beneath the arch of the aorta on a level with the fifth or sixth dorsal vertebra, behind the left pulmonary artery. The trachea and bronchi are formed of a series of cartilaginous rings, incomplete at their posterior third, connected by fibro-elastic membrane, and lined by columnar ciliated epithelium. The last ring is triangular, so as to fit the rings of the two bronchi. The mucous membrane contains numerous racemose glands, the largest on the posterior surface, called the tracheal glands. The muscular fibres consist of two layers, longitudinal, the most external and trans- verse internal, both unstriated. Relations. — The trachea has the following relations in the neck :— Laterally. Lobes of the thyroid body; Carotid arteries; Inferior thyroid arteries. Anteriorly. Skin, superficial and deep fascia; Anterior jugular veins; Left innominate vein; Inferior thyroid plexus of veins; Isthmus of thyroid body; Arteria thyroidea ima (sometimes). Posteriorly, Qj^sophagus ; Vertebral column; Recurrent and inferior laryngeal nerves. The following in the thorax: — Laterally, Pleura; Pneumogastrie nerves. 252 HUMAN ANATOMY. Anteriorly. Sternum ; Remains of thymus; Arch of aorta; Eight and left innominate veins; Left carotid artery; Deep cardiac plexus. Posteriorly. (Esophagus. The arteries are derived from the inferior thyroid and bron- chial. The veins empty into the thyroid and bronchial plexuses. The nerves Are from the pneumogastrie and sympathetic. The lymphatics empty into the mediastinal glands. The bronchi are the continuation of the trachea from its bifurcation to the hilus of the lung. Bight hronchus is shorter, about one inch in length, placed more horizontally, and wider. It divides at the hilus into three short branches, corresponding to the divisions of the right lung. Left hronchus is longer, nearly two inches in length, placed more obliquely, and narrower than the right. It divides iato two long branches corresponding to the number of lobes : — Relations. — ^The right bronchus has the right auricle and the supe- rior vena cava in front of it, the right pulmonary artery at first below, then passing also in front, and the vena azygos arching over it from behind ; The left bronchus has the arch of aorta above, the left pulmonary artery at first above, then in front, and the oesophagus, thoracic duct and descending aorta behind. The arteries, veins, nerves, and lymphatics are the same as for the trachea. The two bronchi subdivide into the bronchial tubes, or bron- chioles, which ramify throughout the lungs, dividing and sub- dividing, to end finally in the primary lobules, where they communicate with the intercellular air-passages. Their carti- lages consist of thin plates, distributed irregularly along the tubes, and in the finer tubes disappearing entirely. The mucous membrane of the bronchi and bronchial tubes is lined throughout with columnar ciliated epithelium. THE LUNGS. The lungs, the organs of respiration, are two in number, occupying the lateral cavities of the chest, separated from each other by the heart and structures within the mediastinum. They VOCAL AND RESPIRATORY APPARATUS. 253 accurately fill the cavity of the chest at all times, and are covered by the pleura. They are conical in shape, presenting each a base, apex, two borders, and two surfaces. The apex extends upward above the level of the first rib; the base occupies the convex surface of the diaphragm; the external^ or thoracic surface is accurately applied to the wall of the thorax ; the inner surface is in contact with the pericardium, and is marked by a depression, the hilum pulmonis, at the root of the lungs. The posterior bor- der rests on either side of the spiaal column, and the anterior border is thin and overlaps the pericardium. The root of each lung, situated near its middle, is composed of the following structures, surrounded by a reflection of pleura : — Bronchus ; Pulmonary artery; Pulmonary veins; Bronchial glands; Bronchial vessels; Posterior and anterior pulmonary plexuses of nerves; Connective tissue. The root of the right lung is behiud the right superior cava and the vena azygos arches over it. The root of the left lung lies below and in front of the arch and descending aorta. The relative positions of the pulmonary veins, pulmonary artery, and bronchus on either side are: — Both sides, from before backward — V. Pulmonary veins, A. Pulmonary artery, B. Bronchos. Eight side, from above downward — B. Bronchus, A. Pulmonary artery, V. Pulmonary veins. Left side, from above downward — A. Pulmonary artery, B. Bronchus, V. Pulmonary veins. The weight of the lungs varies according to many conditions. In the adult the approximate weight is about forty-two ounces, the right being two ounces heavier than the left, and their total capacity about three hundred cubic inches. The right lung has three lobes, the left but two, of which the lower is the larger. 254 HUMAN ANATOMY. The lungs in infancy are of a pale-rose color, but later become of a leaden hue, variegated with dark slate and bluish-black. They are highly elastic, crackle on pressure, and float in water, having a specific gravity of 0.345 to 0.746. In the foetus before birth, and also in certain diseases, they sink in water. The structure of the lung consists of an external serous coat, the visceral layer of the pleura, a subserous areolar tissue, highly elastic, and the parenchyma, or proper substance of the lungs. The latter is made up of small polyhedral primary lobules, which imite to FlO. 122. Heart and lUBgs: 1, right Tentricle; 3, right auricle; 5, pulmonary artery; 9, aorta; 10, superior cava; 20, root of lung; 21, 22, 23, upper, middle, and lower lobes ot right lung; 24, 25, upper and lower lobes ot left lung. form secondary lobules, the latter giving rise to the markings on the surface. Both the primary and secondary lobules are held together by connective tissue. A primary lobule represents the structure of the entire organ, consisting of a hronchiole and in- fundibula, or air-passage, communicating with numerous air- cells. The air-cells are minute polyhedral cavities, from one two- hundredths to one-seventieth of an inch in diameter, separated from each other by a thin lamina, and communicating freely with the infundibula. They vary much in size, being largest on the surface, the thin borders, and the apices. Each air-cell is com- posed of the basement membrane, lined with squamous epi- VOCAL AND RESPIRATORY APPARATUS. 255 thelium, and surrounded by a minute capillary net-work of blood-vessels, which intervene between the terminal branches of the pulmonary artery and the commencement of the pulmonary veins. Between the cells the capillary net-work forms a single layer. The pulmonary artery conveys the venous blood to the lungs and terminates in the capillary net-work about the air-cells, from which proceeds the pulmonary vein, which proceeds along the bronchial tubes to the left auricle of the heart. The bronchial arteries supply the structure of the lungs. They are derived from the aorta, and follow the course of the bronchial tubes, the ironchial veins returning the blood to terminate in the hemi- azygos or superior intercostal vein on the left. The lymphatics are numerous and consist of a superficial set converging to the root and a deep set along the course of the tubes, and both enter the bronchial glands. The nerves are derived from the anterior and posterior pul- monary plexuses of the pneumogastric and sympathetic, the lat- ter the larger. Ganglia are found upon these nerves. THE PLEUE^. Bach lung is invested by a delicate serous membrane, the pleura, which lines the internal wall of the thorax and is reflected at the root of the lung over that organ. It consists essentially of two layers, a parietal, or pleura costalis, and visceral, or pleura pulmonalis. It adheres accurately to the subjacent structures, and is called, from its position, costal, diaphramatie, mediastinal, and pulmonary. The space between the two layers of each pleura, known as the cavity of the pleura, contains a thin, serous secretion. A fold extending downward from the root of the lung to the diaphragm forms the so-called pulmonary ligament, or liga- mentum latum pulmonis. Each pleura is a closed sac ; the right is wider, shorter, and extends higher in the neck than the left. They do not meet in the median line, except opposite the upper part of the gladiolus, but have a space between them known as the mediastinum. The arteries are from the intercostal, bronchial, pericardiac, internal mammary, musculo-phrenie, and thymic. The veins accompany the arteries. The lymphatics are numerous, and empty into the intercostal and posterior mediastinal glands. The nerves are from the phrenic and sympathetic. 256 HUMAN ANATOMY. MEDIASTINUM. The mediastimim is the space left between the two pleurse in the median line between the stenmm and vertebral column, and contains the heart within its pericardium and all the thoracic viscera except the limgs. The intervals in the mediastinum from their position are named anterior, middle, posterior, and superior mediastinal cavities. The boundaries and contents of the four mediastinal cavities are as follow: — Anterior mediastinum, bounded in front by the sternum; laterally, pleura; behind, the pericardium. It contains: — Triangularis sterni muscle; Remains of thymus gland ; Left internal mammary artery and vense comites ; Lymphatic vessels from convex surface of the liver; Areolar connective tissue. Middle mediastinum, bounded in front by the anterior mediastinum; laterally, by the pleura; behind, posterior media- stinum. It contains : — Ascending portion of aorta; Superior vena cava; Heart, inclosed in the pericardium; Bifurcation of trachea; Pulmonary artery and veins; Phrenic nerves; Arteriae comites nervl phrenici, from the internal mammary. Posterior mediastinum, bounded in front by the pericardium and root of the lungs; behind, vertebral column; laterally pleura. It contains: — (Esophagus; Vena azygos major; Thoracic duct; Vena azygos minor; Descending aorta; Superior intercostal veins; Lymphatic glands and vessels; Pneumogastrio nerves; Great splanchnic nerves. The superior mediastinum is that portion of the mediastinal space above the upper border of the fifth thoracic vertebra. It is bounded in front by the manubrium sterni ; behind, by the upper dorsal vertebra ; laterally, by the pleura. It contains : — Origin of stemo-thyroid muscle; Left superior intercostal vein; Origin of stemo-hyoid muscle; Left recurrent laryngeal nerves; Lower end of the longus colli muscle; Cardiac nerves; Innominate artery; Pneumogastrio nerves; Left carotid artery; Phrenic nerves; Subclavian artery; CEsophagus; Transverse portion of the aorta; Trachea; Innominate veins; Thoracic duct; Superior vena cava; Remains of the thymus gland; Lymphatics. THE GENITO-URINARY APPARATUS. The genito -urinary apparatus consists of the urinary organs and the male and female generative organs. THE DEINAET 0K6ANS. The urinary organs consist of the kidneys, ureters, bladder, and urethra. The kidneys are two glandular organs, situated deeply in the lumbar region, opposite the second or third lumbar verte- bra and last dorsal, the right a little lower than the left. They are surrounded by fat and held in position by the blood-vessels and loose connective tissue. The right kidney is ia relation in. front with the liver, descending portion of the duodenum and ascending colon. The left is in relation with cardiac end of stomach, lower end of spleen, tail of pancreas, and descending colon. Bach kidney measures about four inches in length, two in width, and one in thickness, and weighs from four and one-half to six ounces in male, four to five and one-half in female. Their shape is characteristic. The notch at the inner side is called the hilus, and communicates with the interior cavity, the sinus, at which the blood-vessels, nerves, and ureter have their passage. The kidneys have a special fibrous coat, slightly adherent, which extends into the hilus and becomes continuous with the fibrous coat of the blood-vessels and ureters. The relative position of the arteries, veins, and ureter at the hilus are, from above down- ward : — A. Artery, V. Vein, U. Ureter; from before backward : — V. Vein, A. Artery, U. Ureter. The general structure of the kidney consists of a cortical substance and a medullary substance, the latter arranged into conical masses called renal pyramids, with their bases in the cortical substance, and their apices, the renal papillse, projecting (257) 258 HUMAN ANATOMY. into the calices of the pelvis within the sinus of the kidney. The renal pyramids of Malpighi, about ten to fifteen in number, corresponding to the foetal lobules, are arranged into three irregu- lar rows. The substance, or parenchyma of the kidney, is com- posed of urinif erous tubules, held together by comparatively little —A. Pia. 123. Longitudinal section o£ kidney: 1, cortex; 2, medulla; S, section of tubules In boundary layer; 4, fat of renal sinus; B, renal ar- teriole; A, branch of renal artery; C, renal calyx; U, ureter. connective tissue. The terminal orifices of these, several hundred to each papilla, open on its summit. Course of the Tuhuli Urinif eri. — ^Beginning at the Mal- pighian capsule within the cortex, each tubule pursues a very tortuous and complicated course before it terminates in the renal papillaa, the outline of which is as follows (after Gray) : — THE GENITO-URINAEY APPARATUS. 259 fa) Neck — the constricted portion below the capsule, before it ter- minates in t}ie renal papillae. (b) Proximal convoluted tube, i Within the cortical struc- (cj Spiral tubule of Schachowa, f ture. (d) Descending limb of Henle's loop, -i Within the medullary (ej Henle's loop, | structure. (f) Ascending limb of Henle's loop, i Partly in medullary and partly in cortical. (g) Irregular tubule, \ (h) Distal convoluted tubule, I Cortical structure. { i) Curved tubule, J (}) Straight collecting tubule — descends through the medullary structure to open by an orifice in a renal papilla. The straight, collecting or receiving tubes converge as they descend, and unite and reunite until there are about one dozen, but they vary from eight to twenty, one opening on the summit of each papilla. In the cortical portion they form groups, or medul- lary rays, giving the appearance of conical masses in the cortical substance with their apices toward the periphery — the so-called pyramids of Ferrein. The tubules are composed of a basement membrane, lined with pavement epithelium. In the neck the epithelium becomes continuous with that of the Malpighian capsule, and at the glomerule the wall is reflected upon the in- closed tuft, or Malpighian corpuscle. In other situations the epithelium varies much in shape and size. The renal arteries subdivide at the hilum, cross the sinus, and enter the renal sub- stance between the renal papillfe. They terminate in the renal glomerules, or Malpighian corpuscles. Each one of these re- markable convolutions of capillaries is about one one-hundredth of an inch in diameter and inclosed in a pouch-like dilatation of the uriniferous tubules or Malpighian capsule, or capsule of Bowman. The efferent vessels form, together with others, a capillary net-work between and around the uriniferous tubules. They are made up of three sets: (a) the plexuses around the tubuli contorti ; (b) the veins beneath the capsule ; and (c) the plexuses about the apices of the pyramids of Malpighi. These form the venw interlobulares, ivhich join the venw rectce, and together form the renal veins, vencB propria renales. Those in the sinus unite together to form the renal vein, and open iato the inferior vena cava, the left crossing in front of the abdominal aorta. The ureter commences as a compressed pouch, the pelvis. Within the sinus the pelvis divides, and subdivides into several small funnels, or calices, each one of which receives one or two projecting renal papillse. The calices, pelvis, and ureter are all of the same structure, consisting of a fibrous and unstriated mus- cular and a mucous coat. The fibrous coat becomes continuous Avitli the capsule above and below in the fibrous structure of the 260 HUMAN ANATOMY. bladder. The muscular coat ceases at the base of the renal papillae, but the mucous membrane becomes continuous with that of the uriniferous tubules. Its epithelium is of the "transi- T and 8. Ascending limb of Henle'B loop tube. 15. Tabnle of Bellini. |b Sulieapsiilnr layer with- out Malpighian cor- puscles. 12. First part of col- lecting tube. 11. Distal convoluted tubule. A. Cortex. 10. Irregular tubule. 3. Proximal convo- luted tubule. 9. Wavy part of as- cending limb. 2. CoDstnction or neck. 4. Spiral tnbnia. 1. Malpighian surrounded tuft Bovrmana capsule. 8. Spiral part of as- cending limb of Henle's loop. B. BouiroABT Zone. 6. Pescending limb of Henle's loop tube. 0. Benis B loop. C Fafillart Zonb. Diagram of uriniferous tubules: 1, Malpighian capsule; 2, proxi- mal convoluted tubule; 2 b, distal convoluted tubule; 3, descending limb of Henle's loop; 4, ascending limb of Henle's loop; B, irregular tubule; 6, collecting tube; a, apex of pyramid; b, base of pyramid; o. cortical portion. THE GENITO-URINARY APPARATUS. 261 tional" variety, consisting of several layers of cells of varying shapes, resting upon a basement membrane. The nerves are derived from the renal plexus of the sympa- thetic, formed by branches in the semilunar ganglion, the solar plexus, and the lesser and smallest splanchnic nerve. The lymphatics communicate with the lumbar glands. The ureter proper is a musculo-membranous tube, sixteen to eighteen inches in length, about the diameter of a goose-quill, extending from its origin in the pelvis of the kidney to the basal or posterior angle of the vesical trigone, which it enters by passiag obliquely through the muscular and mucous walls of the bladder. Course. — It descends upon the psoas magnus muscle, be- neath the peritoneum, being crossed by the spermatic vessels. About the first division of the sacrum it crosses the external or common iliac artery, passing behind the sigmoid flexure on the left and the ileum on the right side, to enter the pelvis, where it reaches the bladder withia its posterior false ligament. In the male it passes behind the vas deferens. In the female it crosses the uterine artery ^/g to V^ i^^h from the cervix uteri and passes along the side and upper part of the vagiaa to the bladder {vide Vagina). Muscles of the Ureters. — Two oblique muscles. Origin, be- hind the orifices of the ureters ; insertion, into the middle lobe of the prostate gland ; action, they guard the orifices of the ureters, and prevent the reflux of uriae. Suprarenal capsules {vide Ductless Glands). THE BLADDEH, The bladder is a musculo-membranous sac, the reservoir for the urine, situated in the anterior part of the pelvis, behind the pubes, in front of the uterus and vagina ia the female, and the rectum in the male. It measures, moderately distended, five inches in length, three in width, and holds about one pint. It consists of a body, summit, base, and neck. The body is partially invested with peritoneum behind, but in front it is wanting, the body being in relation with the sym- physis pubis, triangular ligament, and internal obturator muscles. The summit, or apex, is rounded, and directed upward and forward, being connected to the umbilicus by the urachus, the remains of the foetal allantois, and also by the obliterated hypo- gastric arteries, one on either side. 262 HUMAN ANATOMY. The space bounded by the pubic surface and the sides of the bladder uncovered by peritoneum, filled by the recto-vesical fascia, is known as the space of Eetzius. The base, or fundus, in the male is situated upon a triangular space upon the second portion of the rectum, its base formed behind by the recto-vesical fold, its apex by the prostate gland, and its sides by the vas deferens and vesiculee seminales. In the female it is situated in contact with the cervix uteri and anterior wall of the vagina, adhering closely to the latter. Its upper portion has a peritoneal covering. The nech, or cervix, is the contracted portion, continuous with the urethra. It is encircled in the male by the prostate gland, and is directed in both obliquely forward and downward. The ligaments of the bladder consist of five true ligaments derived from the pelvic fascia and the urachus; and five false ligaments derived from the peritoneum. The true ligaments are: — Two anterior (pubo-prostatic), two folds of recto-vesical fascia passing from the pubic symphysis to the cervix and upper surface of prostate gland; Two lateral folds of recto-vesical fascia connecting the lat- eral surfaces of prostate gland with sides of base of bladder; The urachus, an obliterated fcetal structure, passing as a fibro-muscular cord from the apex of the bladder to the um- bilicus. The false ligaments are : — Two posterior ligaments are peritoneal folds passing be- tween sides of rectum in male, sides of uterus iu female, to postero-lateral surface of bladder, inclosing obliterated hypogas- tric arteries, ureters, vessels, and nerves ; Two laterals are folds of peritoneum passing from iliac fossae to sides of bladder ; The superior is a single peritoneal fold inclosing the oblit- erated hypogastric arteries from apex of bladder to umbilicus. The structure consists of four coats, a serous, muscular, sub- mucous, and mucous. The serous coat is derived from the peritoneum. It covers the posterior surface from the ureters to the summit, and is re- flected from the sides to the walls of the pelvis and abdomen. The muscular coat is made up of unstriated fibres arranged in three layers: — ■ (a) Longitudinal external layer, named the detrusor urinm muscle; (b) Circular middle layer, forming at the neck the sphincter vesica; THE GENITO-TJRINAEY APPARATUS. 263 (c) Longitudinal internal layer. The submucous or cellular coat, made up of fibrous and elastic tissue, supports the mucous coat and unites it with the muscular. The mucous membrane is smooth and of a pale-rose color. Its epithelium is continuous with that of the ureters and pelvis of kidneys; its superficial layer, large, tessellated, polyhedral cells, its deep layer of club-shaped and spindle-shaped cells. The inner surface of the base presents a triangular space, the vesical trigone, or trigonum vesicce, apex in front, formed by the orifice of the urethra, its basal angles behind about two inches apart, and each about one and one-half inches behind the urethral orifice, formed by the orifices of the ureters : — U I'A IV," U U 2" The uvula vesicce is an elevation of mucous membrane pro- jecting from the floor near the apex of the trigone into the orifice of the urethra. The arteries are the superior, middle, and inferior vesical, with branches from the obturator and sciatic in the male, and branches from the vaginal and uterine in the female. The veins from the vesico-prostatic empty into the internal iliac vein. Lymphatics follow the course of the vessels and enter the lumbar glands. The nerves to the base and neck are from the third and fourth sacral, and to the summit from the hypogastric plexus of the sympathetic. GENERATIVE APPAEATU8. Male Organs. The male organs of generation consist of the testes, vasa deferentia, vesiculse seminales, and penis. The testicles (testes) are two glandular bodies which secrete the spermatic fluid, and are suspended by the spermatic cords within the scrotum, the left a little larger and lower than the right. They are oval, compressed laterally, measure one and a half to two inches in length, one inch in breadth, one and one- fourth inches in their antero-posterior diameter, and weigh from six to eight drachms. 264 HUMAN ANATOMY. They each consist of a body, or testicle proper, and an epididymis. The latter consists of a head, body, and tail. The scrotum consists of the skin marked in the median line by the raphe, formed by the union of the genital folds, and the dartos, a reddish, contractile tissue which surrounds the tester and extends from the raphe to the under surface of the penis, forming the septum scroU, which divides it into two cavities for the testes. The spermatic cord is made up of the following struc- tures : — External spermatic, or intercolumnar fascia; Cremaster muscle; Internal spermatic fascia, or fascia propria; Vaa deferens; Spermatic artery; Deferential artery from superior vesicle; Cremasteric artery, from epigastric; Vasa spermatica; Lymphatic vessels; Spermatic plexus of nerves [from renal and aortic plexuses] ; Genital branch of genito-crural ; Scrotal branch of the ilio-inguinal. It commences at the internal abdominal ring, passes ob- liquely through the inguinal canal, emerges at the external ab- dominal ring, descends into the scrotum, and terminates at the posterior border of the testes. The external spermatic or intercolumnar fascia is derived from the aponeurosis of the external oblique muscle. This mus- cle arises within the inguinal canal from Poupart's ligament and pubic spine, and descends along the cord in loops. Its fibres are striated. The cremaster muscle, or cremasteric fascia or middle sper- matic fascia, is derived from the lower border of the internal oblique and transversalis muscle. The internal spermatic fascia, or fascia propria, is the in- fundibuliform process of the transversalis fascia. The proper coverings or tunics of the testicle are three — tunica vaginalis, tunica albuginea, and tunica vasculosa : — The tunica vaginalis is a serous membrane surrounding the testes, and at the posterior portion is reflected on itself to form a sac. The outer portion of the latter is loosely attached, except at the lower part of the testicle, where the gubernaculum testis binds it down. The tunica albuginea is a dense, white, fibrous structure, continuous at the upper part with a similar structure on the epididymis. At the back part it projects into the glandular sub- THE GENITO-UKINARY APPARATUS. 265 stance of the testicle to form the mediastinum testis (or corpus Highmorianum), from which numerous septula (trabeculae) Fig. 125. Male organs of generation: A, interior of blaflder; B, sympliysia pubis; C, integument: D, vas deferens; E, vesiculse seminales; F, prostate gland; G, corpus cavernosum; H, scrotum; I, verumontanum; J, Cowper's gland; K, orifice of ejaoulatory duct; L, bulb of corpus spongiosum; M, corpus spongiosum; N, urethra; O, prepuce; P, glans penis; Q, suspensory ligament; K, space of Retzlus; S, peritoneal told. 266 HUMAN ANATOMY. diverge, and with similar cords from the tunica albnginea serve to maintain the shape of the gland and divide it into lobules. The tunica vasculosa (pia mater testis) lies within the tunica albuginea, and is the source and termination of the blood- vessels to the secretory substance of the gland, resembling the pia mater of the brain. The glandular structure of the testis is divided into several hundred (two hundred and fifty to four hundred) pyramidal lolules (lobuli testis), each one of which consists of from one to three tubuli seminiferi. At the conical extremity of the lobules the tubules end in straight tubes, vasa recta, which enter the mediastinum to form the plexus retiformis. Prom the upper part of this the vessels unite into from twelve to twenty vasa efferentia, which pass out of the testicle to the epididymis. Within the epididymis these efferent canals form a series of spermatic cones, the coni vasculosi. These end in a coarse, convoluted tube, about twenty feet in length, forming the body and tail of the epididymis, and ending in the spermatic duct, or vas deferens. These tubes are lined with columnar ciliated epithelium. The vas aberrans of Haller is a spermatic cone joining the epididymis, but unconnected with the testicle. The hydatids of Morgagni are one or two pediculated bodies attached to upper part of testis or head of epididymis, and sup- posed to be the remains of Miiller's duet. Structure of the Tubules. — The tubuli seminiferi are minute convoluted tubules, closed at one extremity, consisting of a hya- line membrana propria lined with several layers of epithelial polyhedral cells, the seminal cells, from which the seminal or spermatic fluid is directly elaborated. These cells undergo a process of indirect division to form the spermatoblasts, which are arranged into bundles, and are con- verted into spermatozoids, the nuclei becoming the head, and the tail, or ciliary appendages, being afterward developed. These cells are constantly cast off and replaced. Descent of the Testes. — In early foetal life the testes are placed at the back part of the abdomen, below and in front of the kidneys, and behind the peritoneum. About the third month a peculiar structure, the gubcmacu- lum testis, appears, attached to the lower end of the epididymis, and extending as a cord to the bottom of the scrotum. It is supposed to cause the descent of the testicle. It reaches its full development between the fifth and sixth month, at which time the testicle reaches the iliac fossa. It enters the internal abdom- inal ring by seventh month, and the scrotum by the eighth THE GENITO-UEINAEY APPARATUS. 267 month, carrying before it a fold of peritoneum, which is after- ward shut, off, forming the tunica vaginalis testis. Other cover- ings of the testicles are also derived in this manner. In the female a structure similar to the gubernaculum forms the round ligament. The vas deferens has three coats — (1) an external fibrous coat, (2) an unstriated muscular coat, and (3) a lining mem- mu scle: FlO. ]2«. The testicle and eplflldymis: A, testis; B, epididymis; C, tunica vaginalis: D, vas deferens; E, spermatic artery and veins; P, artery of cord, brane of columnar epithelium. It is about one and one-half feet long and one line in diameter. From the tail of the epididymis it ascends in the cord, forming an important part, and lying behind the blood-vessels. It ascends through the inguinal canal, and at the internal ring descends on the bladder, crosses the ureter and obliterated hypogastric artery, and runs forward to form with the duct of the seminal vesicle the ejaculatory duct. 268 HUMAN ANATOMY. The walls of the vas are very thick and the canal very small. Its course for the most part is straight, but beneath the bladder it becomes enlarged, tortuous, and more capacious, but again contracts near its termination. The arteries of the cord are: — The spermatic, to the testicle from the aorta; The vas deferens, or deferent artery, from the superior vesical ; The cremasteric, from the deep epigastric. The veins from the testis — spermatic veins — ^unite with branches from the epididymis to form the pampiniform plexus, which terminates on the left side, passing beneath the sigmoid flexure in the left renal vein, on the right side in the inferior vena cava. The lymphatics are numerous, and terminate in the lumbar glands. The nerves are from the sympathetic system, and form the spermatic plexus. The arteries of the scrotum are derived from the Cremasteric branch of epigastric; Superficial external pudic, from the femoral; Deep external pudic, from the femoral ; Superficial perineal, branch of internal pudic, from internal iliac. The seminal vesicles are two membranous pouches, closely adhering to the under surface of the bladder. Their size varies, but they usually measure two and one-half inches in length, one-half inch in breadth, and two to three lines in thickness. Each consists of a tube closed at one end, about four to sis inches long, the diameter of a quill, convoluted into a mass. Its structure is analogous to that of the ducts, but thinner. The ejaculatory ducts, two in number, are formed by the union of the vasa deferentia with the duct of the vesiculse semi- nales. Each duct is three-fourths to one inch in length, and passes through the prostate gland to terminate at the margin of the sinus pocularis by a slit-like orifice. The arteries are derived from the inferior vesical and middle hsemorrhoidal. The veins and lymphatics correspond, and the nerves are from the hypogastric plexus of the sympathetic. The sepiinal vesicles are not only reservoirs, but secrete a fluid to dilute the spermatic liquid. The semen is a viscid, whitish liquid, composed of a color- less liquid, the liquor seminis, and the spermatozoa and seminal granules. THE GENITO-URINAKY APPARATUS. 269 The seminal granules are about one four-thousandth of an inch in diameter. The spermatozoa are the essential elements, and consist of an oval head, a body or middle piece, and a tail. The penis is composed of three columnar bodies, two cor- pora cavernosa and one corpus spongiosum, invested with skin, and filled with a peculiar, vascular, sponge-like structure. It is attached to the pubic arch and symphysis by its root, has a free expanded extremity, the glans penis, or head, and an intermediary portion, the body. Fig. 127. Vertical section ol testicle. (After Ora]/.) The upper surface is the dorsum, or back. The glans is a olunt, cone-like body, expanded at its base, the corona, and attached by its cervix, or neck. The summit has a slit-like aperture, the meatus^ or orifice of the urethra. The ?kin adheres loosely to the organ, and has a loose, cup- like fold over the glans, called the prepuce, after which it is reflected into the cervix and glans, becoming continuous with 270 HUMAN ANATOMY. the mucous membrane of the urethra at the meatus, to which it is attached below by a band or bridle — frenum or frenulum The skin covering the glans resembles mucous membrane, is very vascular and sensitive, but about the cervix and corona has numerous preputial glands — glandulw Tysoni odoriferi, which secrete the smegma. The superficial fascia is thin, free from adipose tissue, and continuous with that of the abdomen and scrotum. From the front of the pubis a fibro-elastic fasciculus de- scends to the root of the penis, called the suspensory ligament. The corpora cavernosa, or cavernous bodies, constitute more than two-thirds of the bulk of the organ. They spring as crura from the rami of the ischium and pubis, swell out into the hulb of the cavernous hody, or corpus cavernosum, join each other in the median line, to terminate in a conical extremity, which re- ceives the glans. They are grooved above for the dorsal vein, and below for the corpus spongiosum. They each have a strong outer fibrous membrane, which, on uniting, forms a thick partition, complete behind, but incom- plete and comb-like in front — septum pectiniforme. From this septum fibro-elastic bands diverge in all directions and form trdbeculce, which with the blood-vessels occupying their intervals, form the erectile tissue of the corpora cavernosa. The corpus spongiosum commences as a bulb below the crura and in front of the triangular ligament. It expands anteriorly into the glans penis, which fits upon the conical termination of the corpora cavernosa. The urethra enters above and anteriorly to the bulb, and traverses its entire length to the summit of the glans, opening at the meatus. The corpus spongiosum consists of an external fibrous coat, thinner and more elastic than that of the corpora cavernosa, and an interior erectile tissue. A thin, muscular layer lines the ex- ternal fibrous coat, and another is found beneath the mucous membrane of the urethra. The corpora cavernosa get their blood from the arteries of the corpora cavernosa, and branches from the dorsal artery of the penis, from the internal pudic. The corpus spongiosum is supplied by the artery of the bulb. The arteries of all three bodies terminate finally in the erectile tissue. Many arterial branches, especially at the root of the penis, form short convolutions — the helecine arteries, which terminate in finer vessels, and subsequently open into the spaces of the erectile tissue. Others open directly. THE GENITO-URINARY APPARATUS. 271 From these spaces the veins begin; some wind around the side of the organ to the dorsal vein, while others pass under the pubis to join the prostatic plexus. The lymphatics are numerous; the superficial join the inguinal glands, the deep Join the lymphatic plexus about the prostate and other deep lymphatics of the pelvis. The male ueethea is the common canal for the emission of semen and urine, extending from the neck of the bladder to the meatus urinarius. It is from eight to nine inches in length, and consists of three portions — the prostatic, membranous, and spongy. The prostatic portion extends from the neck of the bladder to the anterior border (apex) of the prostate gland. It is one and one-quarter inches in length, and is the widest and most dilatable part. Its floor is raised, and presents : — Veru montanum, or caput gallinaginis, an elevated ridge; Prostatic sinus, on each side of the veru montanum ; Orifices of the prostatic ducts, in the floor of the prostatic sinuses ; Sinus pocularis, a depression in the median line in front of the veru montanum, presenting the slit-like openings of the ejaculatory ducts. This cul-de-sac is one-quarter of an inch in length. It is homologous with the uterus, and has received the name of utricle, or uterus masculinus. The membranous portion is about three-quarters of an inch in length, extending from the apex of the prostate gland to the corpus spongiosum above and in advance of the bulb. It passes out of the pelvis beneath the symphysis pubis, traversing the triangular ligament. It is the least dilatable portion and has four .n C.fnam / . y^m^tu .. Dp tr IS- -t W ^h-s.u V.m.fffr Pig. 132. Surface anatomy of the myelencephalon. (Roman numerals refer to cranial nerves.) C. 1, first cervical spinal nerve; C. mam., corpus naammlllare; Op. n., optic nerve; Inf., infundibulum; Op. ch., optic chiasm; Subs, int., substantia interpeduncu-geniculate body s Sub. b., basilar sulcus; M. p. c, middle peduncle of cerebellum; P., flocculus; P. h., horizontal fissure; C, olive; V. 1. s., ventrolateral sulcus; Pyr., pyramid; V. m. t., ventral median fissure. (Aflcr Tan Gehuchteti.) (Whitclieacl.) fourth ventricle, and on either side, under the name of the crus cerebelli, it passes to the cerebelhim, forming its middle peduncle. OEREBELLFir. The cerchelJiim. or little brain, occupies the inferior occipital fossse beneath the great cerebral lobes, from which it is separated by the tentorium. Its average weight is a little over five ounces in the male, and is proportioned to the greater brain about one to twenty. It is oblong, flattened from abova downward, and THE NERVOUS SYSTEM. 289 divided into two lateral hemispheres connected by a central por- tion, the vermiform process. It is composed of gray and white matter — the former upon the surface. It is not convoluted, like the cerebrum, but con- sists of a number of thin plates, folia, arranged in a series of crescentric curves, with the concavity forward. The cerebellum consists of a central lobe, or vermis, and two lateral hemispheres. The latter are separated on the inferior surface of the cerebellum by a deep hollow, the vallecula, which lodges the medulla oblongata. The floor of the vallecula is formed by the inferior portion of the vermis. The incisura cerebelli anterior, anterior cerebellar notch, separates the hemispheres in front, and rests against the corpora quadrigemina ; The incisura cerebelli posterior, posterior cerebellar notch, or incisura marsupialis, separates the hemispheres behind, and receives the upper portion of the f alx cerebelli ; The great horizontal fissure separates the cerebellum into a superior and an inferior surface. The upper surface of the cerebullum shows the superior vermis, with its corresponding lobes in the hemispheres. These latter are separated from each other by fissures. The following table gives the arrangement of the lobules : — Great horizontal fissure. Pre- Post- Pre- Post- Frenui.um. Ala. Great Anterior crescentic lobule. Posterior crescentic lobule. LlJfGUI.A. Lobus centralis. Lobus eulminis. Lobus clivi. Folium cacuminis. FnEJSfULUM. Ala. Anterior crescentic lobule. Posterior crescentic lobule. Posterior superior lobule. -central fissure, -central fissure. -clival fissure. -clival fissure. horizontal ~ fissure. The inferior surface of the cerebellum presents the deep depression, the vallecula, which renders the connection between 290 HUMAN ANATOMY. the vermis and the hemisiDheres less intimate. The divisions of the vermis and their corresponding lobules in the* hemispheres are as follows : — Great Post- Pre- Post- Great Slender lobe, post gracile pos- terior inferior lobule. Lobulus biventer. Amygdala. Flocculus. Tuber valvulse. Pyramis. Uvula. Nodule. Posterior Inferior lobule. Lobulus biventer. Amygdala. Flocculus. horizontal fissure. pyramidal fissure. pyramidal fissure. nodular fissure. horizontal fissure. The cerebellum is connected with the encephalon by the peduncles of the cerebellum^ three in number, from above down- ward, as follows : — Crura ad cerebrum, superior, or processus e cerebello ad testes, pass to the cerebrum; Crura ad posterior, middle, or processus ad pontem, the transverse fibres of the pons Varolii, connect the hemispheres ; Crura ad medullam, inferior, or processus a cerebello ad medullam, the restiform bodies of the medulla oblongata, connect with medulla oblongata. The internal structure of the cerebellum consists of an arbor vitse arrangement of gray matter, inclosing a white mass. In centre of the latter is found a grayish, dentated mass, the corpus dentatum, an irregular capsule of gray matter opening anteriorly. THE CEREBRUM consists of a large, ovoidal mass, divided into two lateral halves, or hemispheres, by the great longitudinal fissure, connected by a white, transverse coinniissure — the corpus callosum. The sur- face is irregularly marked by convolutions, or gyri, separated from each other by irregiilar depressions, fissures, or sulci. The outer surface is composed of gray matter, which, from its loca- tion, is called the cortical substance. The interior surface, for the most part, is white. THE NERVOUS SYSTEM. 291 The principal fissures of the brain are five, as follows : — 1. The great longitudinal fissure, separating the two hemispheres from one another. 2. The great transverse fissure of Bichat, between the cerebellum and the cerebrum, admitting the pia mater to form the velum inter- positum. 3. The fissure of Bylvius, beginning at the anterior perforated space and ascending obliquely. 4. The fissure of Rolando, descends from near the middle of the great longitudinal fissure to join the fissure of Sylvius. ^.f.ns. -^^^p. S..postei Ji.ant:cisc.5. S.temp.. ■(S.'fefnp.Tn^ Pig. 133. Left Bide of human cerebrum. S. fr. B., superior frontal sulcus; S. praec, precentral sulcus; S. R., central sulcus of Rolando; S. poste., Post-central sulcus; S. interp., Interparietal sulcus; S. p. o., parieto- occipital fissure; Ram. post. S., posterior ramus of fissure of Sylvius; S. temp, m., middle temporal sulcus; S. temp, s., superior temporal sulcus: F. S., fissure of Sylvius; R. ant. asc. S., ascending anterior ramus of fissure of Sylvius; R. a. h. S., horizontal anterior ramus of fissure of Sylvius; S. fr. inf., inferior frontal sulcus. (After Tan Oehuchten.) (Whitehead.) 5. The parieto-occipital fissure, on the postero-lateral aspect of the cerebrum. 6. The calloso-marginal fissure. 7. The collateral fissure. 8. The limiting sulcus of ReiL 292 HUMAN ANATOMY. The principal lobes of the brain are six, as follows : — 1. Frontal lobe, on the outer surface of the brain; it is bounded below by the fissure of Sylvius, and behind by the fissure of Rolando. On the mesial surface it is bounded by the callaso-marginal fissure, and on the inferior surface it is bounded behind by the stem of the Sylvian fissure. On the outer surface it is divided into the (a) Gyrus frontalis ascendens; Cb) Gyrus frontalis superior; (c) Gyrus frontalis medius; (d) Gyrus frontalis inferior. suice. Sulsiibf' .,,Sulcalmgiy. Fi!s.csk. Convolutions and fissures of the median and tentorial surfaces of the right cerebral hemisphere. Fis. S., fissure of Sylvius; Sul. cal. marg., calloso-marginal sulcus; Sul. subt., subf rental sulcus; Sul. C. C, sulcus of corpus callosum; Sul. marg., marginal sulcus; Sul. Bubp., subparietal sulcus; Fis. p. o., parieto-occipital fissure; Fis. calc, calcarine fissure; Fis. den., dentate fissure; Fis. col., collateral Assure. (A^er Tan Oehuchten.) iTVMtehead.) On the mesial surface it is divided into (a) Gyrus marginalis; (b) Paracentral lobule. On the orbital surface it is divided into (aj Gyrus orbitalis internus; (b) Gyrus orbitalis anterior; (c) Gyrus orbitalis posterior. 2. Parietal lobe, lies between the fissure of Rolando, the parieto- occipital, and the fissure of Sylvius, and consists of five gyri: — (a) Ascending parietal; "j Supramarginal, (b) Superior parietal; V Angular, (cj Inferior parietal, J Post parietal; THE NERVOUS SYSTEM. 293 3. Occipital lobe, lies at the posterior aspect of the cerebrum and is divided into first, second, and third occipital convolutions. 4. Temporo-sphenoidal lobe, occupies the middle fossa of the skull. Sul.pl/. -'^^ul.temp.inj' '- -Fis.co2. FlO. 135. Inferior aspect of cerebral hemisphere. Sul. olf . Sul. temp, inf., inferior temporal sulcus; Pis. col., (Aiter Fon Oehuchten.) (WMteliead.) olfactory sulcus; Collateral fissure. 5. Island of Reil, or central lobe, lies within the fissure of Sylvius. It consists of six convolutions — the gyri operti. 6. The limbic lobe surrounds the corpus callosum. Its extremities are united hj the roots of the olfactory tract. 294 HUMAN ANATOMY. The inner of median surface of the hemispheres presents five fissures, as follows : — 1. Calloso-marginal. 2. Parieto-occipital. 3. Calcarine. 4. Oocipito-temporal or collateral. 5. Dentate fissure, or sulcus hippocampi. The lobes on the iaternal surface are six in number, as follows : — 1. Gyrus fornicatus, or convolution of the corpus callosura, de- scends as the gyrus hippocampi and terminates as the uncinate gyrus. 2. Marginal, or first frontal convolutions. 3. Quadrate, or prseeuneus. 4. Cuneus, or occipital lobule. 5. Uncinate gyrus. 6. Temporo-sphenoidal lobe. The inferior surface of each hemisphere is divided into three lobes — the anterior, middle, and posterior. The two former occupy the anterior and middle fossa of the sloill and the poste- rior rests upon the cerebellum, separated from it by the tento- rium. This surface presents for study from before backward the following points: — The longitudinal fissure, separates the two hemispheres ; Corpus callosum, the great transverse commissure of the cerebrum, extending by means of its peduncles to near the Sylvian fissure ; Lamina cinerea, is a thin, gray layer, forming the anterior part of the inferior boundary of the third ventricle; Olfactory nerve, with its bulb; Fissure of Sylvius, between the anterior and middle lobes of the cerebrum, and lodges the middle cerebral artery; Anterior perforated space, transmits vessels to the corpus striatum; Optic commissure, is formed by the junction of the optic tracts; Tuber cinereum, is a gray eminence between the corpora albicantia and optic tracts, and forms part of the floor of the third ventricle; Infundibulum, is a tube of gray matter connecting the pituitary body with the third ventricle; Pituitary body, is a small, vascular, bilobed body, connected by the infundibulum and occupying the sella Turcica (for histologv vide "Ductless Glands"); °^ Corpora albicantia, or mammillaria, are two white, rounded masses, formed by the folding of the anterior crura of the fornix, and are sometimes called the bulbs of the fornix; Posterior perforated space, allows the passage of blood-vessels to the optic thalami; Crura cerebri, or cerebral peduncles, connect the cerebrum with the medulla, cerebellum, and spinal cord; it consists of the anterior portion, THE NERVOUS SYSTEM. 295 or crusta, and the posterior portion, or tegmentum, between which is a mass of gray matter — the locus niger; Pons Tarolii, covers up the posterior portion of the cerebral lobes. Interior of the Cekebkum. — The interior of the cerebrum, viewed above the level of the corpus callosum, presents a white surface — the centrum ovale mintis, the margins of which are con- voluted gray matter, and are called lahia cerebri. It is studded throughout with minute blood-vessels — puncta vasculosa. The hemispheres, viewed on a level with the corpus callosum, present a large white mass — the centrum ovale majus — in the centre of which is the connecting band, the corpus callosum. Corpus Callosum. — This connecting band forms the roof of the lateral ventricles. It is about four inches in length and varies from an inch and a half to two inches in width, presenting in front a bend, or genu, below which it terminates in the tuber cinereum through the lamina cinerea. Posteriorly it forms a thick, rounded fold — ^the splenium, or pad — which is continuous with the fornix. The peduncles of the corpus callosum are two reflected bun- dles of white matter given off near the anterior termination of the corpus, and each passing backward across the anterior per- forated space of its own side to the fissure of Sylvius. The superior surface of the corpus callosum shows a depres- sion — the raphe — ^bounded on each side by elevated bands, the strice longitudinales, or nerves of Lancisi. External to these are the strice longitudinales laterales. On either side of the raphe are many transverse lines — ^the lineje transversse, which indicate the direction of the fibres of the corpus. The cerebral commissures are connecting bands of gray and white matter, pursuing either a transverse or antero-posterior course. Transverse: — Anterior, middle, and posterior commissures; Corpus callosum; Optic chiasm; Fornix; Pons Varolii; Posterior medullary velum. Antero-posterior: — Corpus callosum (nerves of Lancisi) ; Fornix ; Fasciculus uncinatus; Tsenia semicircularis; Gyrus fornicatus; 296 HUMAN ANATOMY. Fasciculus longitudinales inferior; Olfactory tracts; Crura cerebri; Peduncles of pineal gland; Processus e cerebello ad testes. VENTRICLES OF THE BRAIN. The interior of the brain contains five distinct cavities, named the ventricles of the brain, situated as follows : Two lat- eral ventricles in the upper part, within the substance of the hemispheres, the third ventricle between the optic thalami at the base of the brain, the fourth ventricle between the medulla oblongata and the cerebellum, and the fifth ventricle within the septum lucidum between the two lateral ventricles. The ventricles intercommunicate- — the two lateral ventricles with the third by means of the foramen of Monro, the third with the fourth ventricle by means of the iter a tertio ad quartum ventriculum, and, in the foetus, with the fifth, and through the infundibulum with the cavity of the pituitary body. The lateral ventricles are bounded as follows: — The roof, the corpus callosum; the floor is formed by the follow- ing parts from before backward: corpus striatum, taenia semicircu- laris, optic thalamus, choroid plexus, corpus fimbrlatum and fornix; internally, by the septum lucidum; externally, in front and behind by the brain-substance. Each lateral ventriole presents three cornua — the anterior cornu, posterior cornu, or digital cavity, and the middle cornu. The anterior cornu curves outward and forward over the corpus striatum and into the anterior lobe. The middle cornu passes into the middle lobe, ascending to the transverse fissure at the base of the brain. Its course is back- ward, outward, downward, forward, and inward (B., 0., D., F.,L). The posterior cornu runs backward into the posterior lobe, its course being backward, outward, and inward (B., 0., I.). Parts of Lateral Ventricle — Corpus Callosum. — Described above. Septum Lucidum. — Forms the internal boundary of the lateral ventricle. It consists of two layers of white and gray matter, and is attached above to the under surface of the corpus callosum, below to the fornix, and anteriorly to the prolongation of the corpus callosum. B.etween the laminse forming the septum is a narrow interval — the fifth ventricle. The Corpus Striatum. — Situated in the lateral ventricle, its broad end directed forward into the fore part of the body and THE NERVOUS SYSTEM. 297 anterior cornu of the ventricle, its narrow end directed outward and backward, and separated from its fellow by the thalami optici. The intraventricular portion is called the caudate nucleus, the extraventricular the lenticular nucleus, the two separated by the internal capsule. —Antcaif ^JfuclcauJ JVelint Chp} ' Post.cor. Fio. 136. The lateral ventricles and choroid plexus. St. term., stria termln- alis; Thai., thalamus; Hipp., hippo-campus; Fimb., fimbria; Ant. cor., anterior cornu Of lateral ventricle; Nucl. caud., nucleus caudatus; Col. for., columns of- thfe fornix; Vel. int.. Velum interpositum; Ch. pi., choroid plexus; Cal. av., calcar avis; Post, cor., posterior cornu of lateral ventricle. (After Gray.) (.Whitehead.) The internal capsule is a large layer of white fibres, derived from the medulla and crura cerebri, separating the lenticular nucleus from the caudate nucleus anteriorly, and the lenticular nucleus from the optic thalamus posteriorly. 298 HUMAN ANATOMY. The external capsule is a small layer of white fibres on the outer surface of the corpus striatum, between the lenticular nucleus and the elaustrum. The elaustrum is a convoluted layer of gray fibres between the external capsule and the island of Eeil. The Tcenia Semicircularis, or Horny Band of Tarinus. — A band of medullary substance in the furrow between the corpus striatum and the optic thalamus. Its anterior portion descends with the anterior pillar of the fornix, its posterior portion passes into the descending horn. Beneath it is the vena corporis striati. The choroid plexus, a vascular membrane, occupying the margin of a fold of pia mater, known as the velam interpositum. It runs across the floor of the lateral ventricle, and communicates with its fellow of the opposite side through the foramen of Monro. Posteriorly it descends into the middle horn of the lateral ventricle. The corpus fimbriatum (tfenia hippocampi), a narrow white band behind the choroid plexus. It is the lateral edge of the posterior pillar of the fornix. The fornix, a lamella of white fibrous matter, beneath the corpus callosum, continuous with it posteriorly, but separated from it anteriorly by the septum lucidum. It consists of two symmetrical halves which join to form the iody, each half having an anterior and posterior crus where they do not join. The anterior crura curve down to the base of the brain, where each crus spreads out and curves upon itself to form the corpus albicans of that side. Prom this point it passes to the corresponding optic thalamus. The posterior crura, at their commencement, are joined to the under surface of the corpus callosum. They pass downward into the descending horns of the lateral ventricles, being con- tinuous with the concave borders of the hippocampi majores. The lateral edge of the posterior crus is called the corpus fimbriatum. The lyra is a series of lines, some transverse, others longi- tudinal and oblique, on the under surface of the fornix, between the diverging posterior crura. Optic Thalamus. — The thalami optici are two large gan- glionic masses, situated between the diverging portions of the corpora striata. Each thalamus rests upon the corresponding crus cerebri. The thalamus is bounded externally by the corpus striatum and taenia semicircularis, and internally forms the lateral bound- ary of the third ventricle. THE NERVOUS SYSTEM. 299 Its upper surface is partly covered by the fornix. Its under surface forms the roof of the descending horn of the lateral ventricle. Its posterior and inferior part exhibits two rounded emi- nences, the external and internal geniculate bodies. Its anterior extremity forms the posterior boundary of the foramen of Monro, which foramen connects the two lateral ventricles with the third. Velum interpositum, a vascular membrane, reflected from the pia mater into the interior of the brain through the trans- verse fissure. It passes beneath the posterior border of the corpus callosum and fornix, and above the corpora quadrigemina, the pineal gland, and the optic thalami. It forms the roof of the third ventricle. Its anterior extremity passes on each side into the corresponding lateral ventricle, forming the anterior extrem- ity of the choroid plexus. The vascular fringes of the velum interpositum projecting into the third ventricle are called the choroid plexuses of the third ventricle. It has two veins, the venee Galeni, which run along its under surface and are formed by the veins of the choroid plexuses and the venae corporis striata. The vense Galeni unite to form a single trunk and empty into the straight sinus. The posterior cornu of the lateral ventricle runs into the substance of the posterior lobe. On the floor of this horn is an eminence corresponding to a sulcus between two convolutions, and called the hippocampus minor. Between the posterior and middle horns is another emi- nence — ^the eminentia collateralis, or pes accessorius. The hippocampus major, or cornu ammonis, a white emi- nence running the entire length of the floor of the middle horn. This eminence is the doubled-in surface of the gyrus fornicatus. The lower extremity of the hippocampus major is called the pes hippocampus. The fascia dentata, the gray and serrated edge of the middle lobe. It is really external to the cavity of the middle cornu. The third ventricle is a mere flssure in the median line of the cerebrum, situated between the optic thalami. It com- municates with the lateral ventricle by the foramen of Monro and with the fourth ventricle by the iter a tertio ad quartum ventriculum. The cavity is crossed by three commissures — the anterior commissure, a white, rounded cord; the middle or soft commissure, composed of gray matter; and the posterior com- missure, a white band connecting the two optic thalami. It is bounded by the following structures: — 300 HUMAN ANATOMY. The roof, by the vlum interpositum, suspending tTiQ choroid plex- uses of the third ventricle, and laterally the peduncles of the pineal gland; floor, by the parts inclosing the interpeduncular spax:e at the base of the brain, viz. : the lamina, cinerea, tuber cinereum and infundibu- lum, corpora albicantia and the posterior perforated space ; laterally, by the optic thalami; in front, by the anterior commissure and anterior crura of the fornix; behind, the posterior commissure and the iter a tertlo ad quartam ventriculum. The fourth ventricle is a diamond-shaped cavity between the cerebellnm behind and the posterior surface of the medulla oblongata and pons in front. It is inclosed behind by the pia mater, which contains an opening for the exit and entrance of ■C.hp|l»oi,d > /P.'^eal Stria ^^-^.^^^^ /P'lea) Kece*& of fir ^nt. .^/^^^ Pineal Body ^mf^^ jL^S'H.p. Corp. O.M.aJ, W^^>^::^^r^^-^r^;^'^^^i ^f Sylvia ^ 1 )KSii^""^' \ ■fe FlO. 137. Mesial section of brain and brain stem. 1, anterior commissure; 2, middle commissure; 3, posterior commissure: 4, pituitary body; IV, fourth ventricle. the subarachnoid fluid from the subarachnoidean space of the brain and spinal cord, and a vascular fold of pia matfer — the choroid plexus. It communicates in front with the third ven- tricle by the iter a tertio ad quartum ventriculum. It is bounded as follows : — The roof, valve of Vieussens and the cerebellum, containing in front the foramen of Magendie, by which it communicates with 'the subarach- noidean space; the floor, the posterior median fissure of the medulla and pons, the fovea posterior, the orifice [ventricle of the central canal of the cord] of Aurantius, the locus caeruleus, and the taenia violacea. THE NERVOUS SYSTEM 801 the calamus scriptorius and the fasciculi teretes; anteriorly, the pons Varolii and medulla oblongata; posteriorli') the cerebellum j laterally, the processus a cerebello ad testes, the restiform bodies, and posterior pyramids of the medulla. Its lining membrane is continuous with that of the third ventricle. The gray matter of the floor of the fourth Yentricle presents masses for the origin of certain cranial nerves. The fifth ventricle is a narrow fissure, formed within the two lamina of the septum lucidum, being origiaally a part of the great longitudinal fissure. It is bounded : — Above, by the under surface of the corpus callosum; Below, by the anterior part of the fornix; Laterally, by the lateral ventricles, from which it is separated by the septum lucidum. It is not lined with epithelium. The mesencephalon includes those portions of the brain SAibstance which connect the cerebrum, cerebellum, and medulla oblongata together, and comprises the following structures : — Crura cerebri, before described {ante, p. 294). The valve of Vieussens, or anterior medullary velum, is a thin layer of white matter stretched between the processes e cerebello ad testes, and forming the roof of the iter a i-ertio ad quartum ventriculum. It presents an elevated ridge descending on its upper part from the corpora quadrigemina — the frcenulum. Corpora or tubercula quadrigemina, or optic lobes, are four spherical eminences, placed in pairs above the valve of Vieussens, ind behind the third ventricle, beneath the posterior border of the corpus callosum. The two anterior are called the nates, the two posterior the testes. The brachia (anterior and posterior) are two white cords connecting them with the optic thalamus and beginning of the optic tracts. The processus a cerebello ad testes connect them with the cerebellum. To the outer side of the optic lobes are two small masses called the corpus geniculatum externum and internum. The pineal gland, or epiphysis cerebri, is a small, reddish, conical body resting upon and between the nates. It represents the rudiment of a median eye of certain extinct lizards and am- phibia (Spencer), well marked in Hatteria. It is attached to the cerebrum ]jy its two peduncles, and is held in position by a fold of the pia mater. It contains a cavity, filled with a viscid fluid and secretory matter composed of phosphate and carbonate of lime, phosphate of magnesia, ammonia, and a little animal matter — the acervuliis cerebri. 302 HUMAN ANATOMY. THE INTERNAL STRUCTURE OP THE MEDULLA OBLONGATA. The anterior and lateral pyramidal tracts of the cord are continued into the medulla oblongata as the pyramids. The lateral pyramidal tracts decussate in the lower part of the me- dulla. The anterior ground bundle is continued upward • into the medulla as the posterior longitudinal bundle. It lies behind the pyramids in the medulla. The sensory columns of the cord (Goll and Burdach) are continued into the medulla oblongata and terminate in the funiculus gracilis and cuneatis. They increase in size and each develops a uncleus : the clava and cuneate nucleus, respectively. These fibres partly form the restiform bodies. The decussation of the sensory fibres takes place at a higher plane than that of the motor fibres. It is also called the decus- sation of the fillet and it consists of the decussating fibres derived from the clava and the cuneate nucleus. These fibres are called deep arcuate fibres. After decussating the fibres are continued upward behind the pyramids, displacing backwards the posterior longitudinal bundle. In the medulla on cross-section are seen the olivary bodies, which contain the dentate nuclei. The restiform bodies, or inferior cerebellar peduncles are formed by the direct cerebellar tract, the posterior superficial arcuate fibres, the anterior superficial arcuate fibres, and the cerebello-olivary fibres. The formatio reticularis is seen behind the pyramids and the olivary bodies in the medulla oblongata. It is composed of the deep arcuate fibres, fibres of Gower's tract, and the antero lateral ground bundle. The gray matter of the cord is continued into the medulla. The anterior horns are cut off and displaced by the decussation of the lateral pyramidal tract and the posterior horns are displaced outward by the increase in size of the posterior sensory tracts. The latter are known as the nucleus of Eolando and are capped by the substantia gelatinosa Eolandi. The central canal is expanded into the fourth ventricle. The gray matter forms nuclei for the cranial nerves in the floor of the fourth ventricle. THE NERVOUS SYSTEM. 303 „ THE IXTEKXAL STRUCTURE OF THE POXS YAEOLII. On cross-section the pons is seen to consist of a dorsal tegmental part, and a ventral part, or crusta. The crusta is composed of transverse fibres (fibres from the cerebellum to the pons and from the nucleus pontis to the cere- bellum), which go to form the middle cerebellar peduncles; lon- gitudinal fibres, which belong to the pyramidal tracts, much scat- tered; and gray matter, which here forms a nucleus called the nucleus pontis. The tegmentum of the pons contains a thick layer of gray matter, which forms the floor of the fourth ventricle, and from which cranial nerves take their origin; formatio reticularis, which is the continuation upward of the same from the medulla ; the superior olivary nucleus; the posterior longitudinal bundle; the fillet; the superior cerebellar peduncle; and the corpus trapezoides. The fillet occupies a position between the crusta and tegmen- tum, and to differentiate it from a tract that makes its appear- ance above the nucleus of the third nerve is called the me- sial fillet. The other tract is named the lateral fillet. The lateral fillet is composed of longitudinal fibres which take origin from the nucleus of the eighth cranial nerve (coch- learis) of the same side, from that of the opposite side, and from the superior olive. They end in the inferior quadri- geminal body, the internal geniculate body, and a few in the superior quadrigeminal body. The mesial fillet has been described. It takes its origin in the medulla from the cuneate and gracile nuclei of the opposite side. Some of its fibres end in the superior quadrigeminal body, but the remainder pass through the subthalamic tegmental region into the posterior part of the lateral nucleus of the optic thalamus. Some end here, while some are continued through the thalamus, enter the corona radiata, and pass to the posterior central gyrus of the Eolandic region. THE STRUCTURE OF THE CEREBELLUM. On section the gray matter of the cerebellum is found to occupy the cortex; to its arborescent appearance the term arbor ritce has been applied. The white matter, medullary body, of each hemisphere con- tains a nucleus of gray matter : the corpus dentatum. The middle peduncles connect the cerebellum with the pons. 304 HUMAN ANATOMY. The inferior peduncles, or restiform bodies, connect the medulla and cord with the cerebellum. The superior peduncles connect the cerebellum with the cerebral cortex. After decussating below the corpora quadri- gemina some of the fibres pass to the opposite red nucleus in the tegmentum of the crura cerebri passing through the optic thalamus to end in the Eolandic region, and others end in the optic thalamus. Each superior peduncle contains both aflEerent and efferent fibres. The cerebellar cortex consists of two layers : an outer, molecular layer, and an inner, granular layer. Between these two layers is a single layer of large cells, the cells of Purkinje. THE STEUCTUEE OF THE illDBEAIN. The midbrain consists of a dorsal part, the corpora quad- rigemina, and a ventral part, the crura cerebri. It is tunneled by the aqueduct of Sylvius, which connects the fourth ventricle with the third ventricle. The upper end of the crura cerebri is encircled by the optic tract. On section the crura cerebri show a ventral and lateral portion, — the tegmentum, — and a dorsal portion, the lamina quadrigemina. The ventral portion contains the substantia nigra. The fissure of Sylvius is surrounded by gray matter, from which the third and fourth cranial nerves take origin. The inferior or posterior quadrigeminal bodies contain the fibres of the lateral fillet posteriorly and the brachium laterally. The superior or anterior quadrigeminal bodies consist of four strata : stratum zonale, stratum cinereum, stratum opticum, and stratum lemnisci. The mesial and upper fillet in part ends in the latter, which also contains large nerve-cells. Some fibres from the occipital lobe and fibres from the retina, conveyed by the superior brachium, end in the superior quadrigeminal bodies. The superior cerebellar peduncles have been described ; they connect the cerebellum with the cerebral cortex. The red nucleus is found in the tegmentum. Some of the fibres of the superior cerebellar peduncles end there. The posterior longitudinal bundle lies in tlie tegmentum and is the continuation upward of the tract of the same name in the medulla and pons. It is connected with the nuclei of the motor nerves of the muscles of the eyeball. The lateral fillet is continued upward from the pons in the tegmentum. Its connections with the superior olivary nuclei and the nuclei of the eighth nerve have been described. THE NERVOUS SYSTEM. 305 The mesial fillet likewise is eontinued upward from the pons in the tegmentum of the crura. The course of the fibres has been described. The crusta of the crura contains the pyramidal fibres and the cortico-pontine fibres. H.nuci.caiid-. Ant con. Tnuc/. caud: . ■St term Paly. : /^ntcol .. Post Col ^n}:cdH ..-Hnuclcaui/. -J-ilnt.cap. £xt.cap. Is/. MecL ?ent ■ Claus. -■-fi Lint cap. Thai. ■ C.jf.m. -• T.nud.caudl Tinih -Hipp. Post, con Fig. 138. Interior of mesencephalon. For., -fornix; Sep. pel., septum pellu- cidum; Col. for., column of fornix; H. nucl. caud., head of nucleus caudatus; Ant. com., anterior commissure: T. nucl. caud., tail of nucleus caudatus; Mas. int. massa intermedia; Thai., thalamus; St. term., stria terminalis; Pulv., pulvinar; Ant. col., anterior colliculus; Post, col., posterior colliculus; IV, Trochlear nerve; Ant. cor., anterior, cornu of lateral ventricle; A. 1. int. cap., anterior limb of internal capsule; Ext. cap., external capsule; Isl., island of Reil; Nucl. lent., nucleus lentiformis; Claus., claustrum; P. 1. int. cap., posterior limb of internal capsule; Hipp., hippocampus; Post, cor., posterior cornu of lateral ventric' 'After Landois.) Whitehead.) 306 HUMAN ANATOMY. SENSORV MOTOR AREA > INTERNAL CATSVtB FIRST TEMF ORAl. OYBUS ' SENSORY DECUSSATION !?■ OF CEREBELLUS , •'•'^'■^ J\ CERE BELLAR :- ^ CORTEX ''•'... y DENTATE NUCLEUS FIFTH NERVE EIGUTH NERVE NINTH NERVE TENTH NERVE NUCLEUS GRACILIS , POSTEROMEDIAN COLUMN POSTEROLATERAL COLUMN -DIRECT CEREBEUJ\R TRACT POSTERIOR NERVE ROOT FlQ. 139. Diagram of sensory tracts from spinal cord to brain (original). THE NERVOUS SYSTEM. 307 CORTSX OF THE BBNSORY MOTOR AREA CORONA BADIATA INTERNAL CAPSULE >7V.'v''/'' ■-,■■■.■. t- '^•*'™-*TE NUCLEUS ^^imi I \''\ ' ' •] °™C THALAMUS LENTICULAR NUCLEUS roURTH NERVE •PINAL COED DECUSSATION OF PYRAMIDS ANTERIOR PYRAMIDAL TRACT ANTERIOR COMMISSURE THIRD NERVK FIFTH NERVE SIXTH NERVE SEVENTH NERVE NINTd NERVE TENTH NERVE ELEVENTH NERVE TWELFTH NERVB UTERAL PYRAMIDAL TRACT ANTERIOR-NERVE ROOT Fig. 140. Diagram of motor tracts from brain to spinal cord (original). 308 HUMAN ANATOMY. THE STRUCTURE OF THE CEREBRUM. On cross-section the cut brain surface shows an outer gray cortex and the inner white matter and cavities, one to each hemisphere: the lateral ventricles. The white matter contains "iF CZ-— • cj[—Wi CM— Fia. 141. Scheme of the nuclei and root-fibres of the cranial nerves. {After Edinger.) {Whitehead.) certain masses of gray matter : the optic thalami and the corpora striata, which latter are on each side of the brain divided into the caudate and lenticular nuclei by the internal capsule. Be- tween the optic thalamus and caudate nucleus on the inside and the lenticular nucleus on the outside passes the broad band of THE NERVOUS SYSTEJI. 309 white fibres known as the internal capsule. As it nears the cortex the fibres spread out. To this is given the name corona radiata. Between the external capsule and the cortex of the island of Eeil is seen a thin sheet of gray matter : the claustrum. The white matter between the claustrum and the lenticular nucleus is known as the external capsule. On horizontal section of the brain the internal capsule is seen to be bent upon itself, the interval between the optic thalamus and the caudate nucleus. This bend is called the genu. One-third of the capsule lies in front of this bend, and is called the anterior limb; the portion behind the genu is called the posterior limb. The anterior limb, contains (1) fibres that pass from the optic thalamus to the lenticular and caudate nucleus, (2) fibres that pass from the optic thalamus to the cortex of the frontal lobe, and (3) fibres that pass from the cortex of the frontal lobe to pass to the nucleus pontis. The posterior limb contains (1) a continuation upward of a portion of the mesial fillet and the superior cerebellar peduncles, (2) the pyramidal tracts or motor fibres from the Eolandic area, (3) the fibres of the optic radiation, (4) the fibres of the auditory radiation, and (5) the temppro-pontine tract. The opposite sides of the brain are connected by the com- missural fibres. Convolutions on the same stde are connected by association fibres. Projection fibres are those which unite the cerebral cortex with nuclei in lower levels. They pass princi- pally through the corona radiata. The cerebral cortex consists of four layers: the stratum zonale, the layer of small pyramidal cells, the layer of large pyramidal cells, and the layer of polymorphic cells. CRANIAL NERVES. The cranial nerves consist of twelve pairs, as follows: — 1. Olfactory, 8. Auditory (portio mollis), 2. Optic, 9. Glosso-pharyngeal, 3. Motores oculorum, 10. Pneumogaatric (vagus, or 4. Pathetici, par vagum), 5. Trifacial, 11. Spinal accessory, 6. Abducentes, 12. Hypoglossal. 7. Facial (portio dura), 1. Olfactory nerve, special nerve of smell. A number of nerves (20) arise from the olfactory bulb or lobe of the brain. Superficial origin of the tract by roots: internal from frontal lobe, middle or gray root from the olfactory tubercle between 310 HUMAN ANATOMY. the other roots of the tract, and external from the middle lobe; deep origin, from gyrus fomicatus, uncinate gyrus of limbic lobe; course, roots unite, pass forward, and form bulbus olfac- torius (from this the olfactory nerves are given off) ; exit, fora- mina of cribriform plate of ethmoid; distriiution, by three Fig. 142. Base of brain and cranial nerves: 1, olfactory bulb; J, opOc nerves; 4, tractus opticus; 5, crus cerebri; 6, third pair ot nerves; 7, fourth pair of nerves; 8, fifth pair of nerves; 9, sixth pair of nerves; XO, pyramid; 11, olivary body; 22, pons Varolii; 24, seventh and eighth pairs ot nerves; 25, ninth, tenth, and eleventh pairs of nerves; 26, twelfth pair of nerves; 27, cerebellum. groups to mucous (Schneiderian) membrane of nares, inner to septum nasi, middle to roof of nasal fossa, outer to superior turbinated bone. H,l\„-h,uJ ,",^ S,^/p Fiii. u:i. Intii; Ofiiloinot.ir, tr.)clilc;iv, unci liiliuial iirivcs. THE NERVOUS SYSTEM. 311 2. Optic nerve, special nerve of sight; superficial origin, optic chiasma or commissure formed by union of the optic tracts. The commissure and tracts contain intereerebral fibres, the cere- bro-retinal fibres of the same side, and the cerebro-retinal fibres of opposite sides. Deep origin, the optic tracts arise from optic thalamus, the upper corpora quadrigemina, and corpora genicu- lati; course, riiverge and pass forward; exit, optic foramen; distribution, to ganglion cells in the retina. 3. Motor oculi, motor nerve; superficial origin, inner sur- face of crus cerebri ; deep origin, from the oculo-motor nucleus in floor of aqueduct of Sylvius; course, from inner side of crus cerebri passes forward, descends along external wall of cavernous sinus ; exit, sphenoidal fissure between two heads of external rec- tus muscle; distribution, by superior and inferior divisions, to all the ocular muscles (including the iris) except the external rectus and superior oblique. 4. Teochleae (pathetic), motor nerve; superficial origin, from valve of Vieussens, on outer side of crus cerebri; deep origin, from floor of aqueduct of Sylvius; course, outer side of crus cerebri, through outer wall of cavernous sinus; exit, sphe- noidal fissure ; distribution, to superior oblique muscle. 5. Trifacial, or Trigeminus^ common sensation, taste and motion ; superficial origin, by two roots, like a spinal nerve, from the side of the pons Varolii; deep origin, the motor root, from (1) a nucleus in the fioor of the aqueduct of Sylvius (descending motor r&ot), and (3) from a nucleus in the pons (these join to form the motor root) ; the sensory root ends in two terminal nuclei — (1) the sensory nucleus of the fifth nerve in the pons and (3) the substantia gelatinosa Eolandi in the pons, medulla, and the spinal cord as far down as the second cervical nerve; course, passes forward to apex of petrous portion of temporal bone, where the sensory root enters Gasserian ganglion, the motor passing beneath, and later joins a branch of the ganglion. It divides into three branches — ophthalmic, superior maxillary, and inferior maxillary; exit, ophthalmic by sphenoidal fissure, superior maxillary by foramen rotundum, inferior maxillary by foramen ovale. Ophthalmic nerve, entirely sensory, supplies lachrymal gland, upper eyelid, skin and muscles of forehead, eyebrow, nose, eyeball (ciliary muscle, iris, etc.), mucous membrane of eyelids and nose, and the ciliary ganglion. Its branches are : — Lachrymal, Frontal, Kasal. 312 HUMAN ANATOMY. Superior maxillary nerve, entirely sensory also; supplies sensation to upper jaw, teeth, hard and soft palates, tonsils, gums, antrum of Highmore, muscles, skin and mucous membrane of lower eyelid, muscles and skin of cheeks and upper lip, and mucous membrane of floor of nares. Its branches are : — Orbital, Middle dental. Palpebral, Spheno-palatine, Anterior dental. Nasal, Posterior dental, Labial. Inferior maxillary nerve consists of two portions : anterior, or motor, and posterior, or sensory, having a threefold function, because one of its branches, the lingual, is joined by the chorda tympani nerve (given off from the facial nerve), which conveys gustatory fibres to the anterior two-thirds of the tongue. It supplies motion to all the muscles of mastication (except to buccinator) , anterior belly of digastric, and mylo-hyoideus ; sen- sation to skin of ear, lower part of face, lower lip, and tongue. Its branches are: — Anterior Portion. Posterior Portion. Masseteric, Auricujo-temporal, Deep temporal (2), Inferior dental. Buccal, Lingual. Pterygoid (2). The fifth nerve has four ganglia connected with it : — (a) Ophthalmic, or lenticular; (bj Spheno-palatine, or Meckel's; (c) Otic, or Arnold's; (d) Submaxillary {vide Sympathetic System). 6. Abddcens, motor ; superficial origin, pyramidal body and pons Varolii; deep origin, floor of fourth ventricle; course, passes forward in cavernous sinus; exit, sphenoidal flssure; dis- tribution, to external rectus muscle. 7. Facial, motor nerve; superficial origin, lateral tract of medulla and pons Varolii; deep origin, from a nucleus in the pons, deeply placed, from which the fibres ascend close to the floor of the fourth ventricle, where they form the eminentia teres arch over the nucleus of the sixth nerve, and then emerge (this nerve carries some sensory fibres, probably gustatory fibres from the tongue, through the chorda tympani nerve to the pass inter- media, near the nucleus of the ninth nerve) ; course, forward and outward, through internal auditory meatus, aqusductus Fallopii, THE NERVOUS SYSTEM. 313 and inner wall of tympanum; exit, stylo-mastoid foramen; dis- tribution, to muscles of expression, buccinator, and platysma, posterior belly of digastric, stylo-hyoid, attrahens and retrahens aurem muscles, to laxator tympani and stapedius through tym- panic branch, dorsal surface of tongue through chorda tympani, and levator palati, and azygos uvulae muscles through Vidian. In its course through the temporal bone it communicates with many important nerves; in the internal auditory meatus, with audi- tory; in aquseductus Fallopii, with otic ganglion, by the small petrosal; Meckel's ganglion, by the large petrosal nerve; the sympathetic of the great meningeal, by the external petrosal nerve; and at its exit from the stylo-mastoid foramen, with the glosso-pharyngeal, pneumogastric, auriculo-temporal, auricularis magnus, and the carotid plexus. Its branches are : — On the Face. In Aquwductus FallopU. "Cb^ Tympanic, ^ ^ Chorda tympani. Kl « Temporal, Temporo-facial, -j Molar, { InfraorbitaL Buccal, Cervico-facial, -j Supramaxillary, Inframaxillary. Posterior auricular, Stylo-hyoid, Digastric. 8. Auditory, special nerve of hearing, consists of two por- tions — (1) cochlear, or auditory portion, and (2) vestibular, or fibres whose function is to localize position. The former pass from the cochlea to the (1) accessory auditory nucleus in the medulla, and (2) to the lateral acoustic tubercle in the medulla. These fibres, by means of the lateral fillet, communicate with the inferior corpora quadrigemina. The vestibular fibres pass from the vestibule to the external and internal dorsal nuclei in the floor of the fourth ventricle, and by the sensory decussation to the nucleus euneatus. Superficial origin, from groove between olivary and restif orm bodies ; course, winds around restiform body, and passes forward to internal auditory meatus, with the facial; distribution, to in- ternal ear by two branches : — Vestibular, and Cochlear {vide Ear, p. 370). 9. Glosso-pharyngeal, or ninth, nerve of motion, common sensation, and taste; superficial origin, from medulla oblongata, between olivary and restiform bodies; deep origin, from motor 314 HUJIAN ANATOMY. FlO, U4. THE NERVOUS SYSTEM. 315 Fig. 145. 316 HUMAN ANATOMY. and sensory gray nuclei in floor of fourth ventricle; course, passes outward across flocculus; exit, jugular foramen, in sepa- rate sheath, in front of pneumogastric and spinal accessory, hav- ing two gangliform enlargements upon it — Jugular and petrosal, petrous, or ganglion of Andersch ; distribution, to mucous mem- brane of fauces, tonsil, pharynx, and middle ear, muscles of the pharynx, and special sense of taste to the base and sides of the tongue. Its branches are: — Tympanic (Jacobson), Pharyngeal branches. Carotid branches. Tonsillar branches. Muscular branches. Lingual branches. 10. Pneomogastkio (vagus, or par vagum), tenth nerve, motor and sensory; superficial origin, from groove between olivary and restiform bodies by a dozen filaments; deep origin, from nuclei in floor of fourth ventricle; course, passes outward across the flocculus; exit, jugular foramen in a common sheath with the spinal accessory nerves; distribution (not definitely known), motor nerve to the organs of respiration and voice, and a motor and sensory nerve to heart, oesophagus, pharynx, and stomach. Its branches are : — Auricular (Arnold's), Thoracic cardiac. Pharyngeal, Anterior pulmonary, Superior laryngeal, Posterior pulmonary. Recurrent laryngeal, (Esophageal, Cervical cardiac, Gastric. It also communicates with the left hepatic sympathetic plexus. 11. Spinal accessory, eleventh, motor nerve; superficial origin, from lateral tract of medulla and spinal cord as low as sixth cervical nerve; deep origin, spinal portion from anterior horn of gray matter, accessory portion from gray nucleus in floor of fourth ventricle; course, the spinal portion enters the skull through the foramen magnum, and joins the accessory por- tion in the jugular foramen; exit, jugular foramen in sheath with the pneumogastric; distiibution, to sterno-cleido-mastoid and trapezius muscles, communicating with the cervical plexus and pharyngeal and laryngeal branches of pneumogastric. 13. Hypoglossal, or twelfth nerve, motor nerve of tongue; superficial origin, from groove between olivary and pyramidal bodies by about a dozen filaments ; deep origin, from gray nucleus at lowest part of floor of fourth ventricle; exit, anterior condy- loid foramen; distribrition, to the omo-hyoid (both bellies), Bterno-hyoid, sterno-thyroid, thyro-hyoid, and muscles of the THE NERVOUS SYSTEM. 317 tongue — styloglossus, hyoglossus, genio-hyoid, genio-hyoglossus, communicating with the pneumogastric, sympathetic, first and second cervical, and gustatory nerves. Its branches are : — bescendens hypoglossi. Muscular, Thyro-hyoid, Meningeaj. Membranes of the Coed. — The membranes of the spinal cord are three — the dura mater, arachnoid, and pia mater. Dura mater is a loose fibrous sheath, continuous with the dura of the brain, and from which it differs in not inclosing the venous sinuses, not dipping into the fissures of the cord, and not being adherent to the bony canal. From the latter it is separated by the venous plexuses and some connective tissue. It extends the whole length of the canal, from the foramen magnum (to which it is attached) to the top of the sacrum. Arachnoid is a delicate serous sac, continuous above with the cerebral arachnoid, inclosing the pia mater, from which it is separated by an interval — the suharachnoidean space. It is filled with the cerebrospinal fluid. The outer surface of the arachnoid is in contact with the dura, the space between them being called the subdural space. Pia mater is a fibrous membrane, closely adhering to the cord and forming its neurilemma. Over the anterior median fissure it is strengthened by a fibrous band — the linea splendens — • and laterally has the ligamentum denticulatum. It terminates below the cord as the.filum terminale — a slender filament. THE SPINAL COED. The spinal cord (medulla spinalis) is the elongated portion of the cerebro-spinal axis contained in the spinal canal. Its length is about sixteen to eighteen inches, extending from the medulla above to the lower border of the first lumbar vertebra below, where it terminates in the cauda equina by a slender pro- longation of gray substance, called the conus medullaris. It presents two enlargements, the upper or cervical, extend- ing from the third cervical to the second dorsal vertebra, and the lower or lumbar, about the position of the second or third dorsal vertebra. It is divided into two lateral halves by the anterior and posterior median fissures, united in centre Ijy the commissure. The lateral portions are subdivided by antero-Iateral and postero-lateral fissures into anterior lateral and posterior lateral columns, and posteriorly a narrow fissure separates the 318 HUMAN ANATOMY. posterior median column from the posterior median fissure. The gray substance occupies the centre of the cord, and is arranged into two crescentie masses connected together by the gray com- missure. The posterior horn forms the apex cornu, from which arises the posterior root of the spinal nerves. The anterior horn is thick and short, and affords origin to the anterior root of the nerve. The gray commissure contains throughout its whole length a minute canal — the central canal, or ventricle of the cord, continuous above with the fourth ventricle. SPINAL NERVES. The spinal nerves consist of thirty-one pairs, arranged in the following order : cervical, eight pairs ; dorsal, twelve pairs ; lumbar, five pairs ; sacral, five pairs ; coccygeal, one pair. Pig. 146. Section of spinal cord and membranes; 1, dura mater; 2, aracb- noid membrane; 3, ganglion on posterior root; 4, anterior root of spinal nerve; 6, 5, subarachnoid space; 6, posterior branch of spinal nerve; 7, anterior branch of spinal nerve. Each of the spinal nerves arises by two roots, an anterior or motor and a posterior or sensory. The fibres of the anterior root arise from the antero-lateral columns, originating deeply in the gray matter of the cord. The posterior roots arise at the postero-lateral fissure, also originating deeply in the gray matter of the cord. The posterior roots have each a ganglion developed upon it — except sometimes the first cervical. These roots unite and the nerve then subdivides into two branches, both having motor and sensory fibres. The posterior branches are smaller and unimportant ; they supply the skin and muscles of the back. The anterior branches supply the neck, front and sides of the trunk, and the extremities. Cervical Plexus. — The cervical plexus is formed by the anterior divisions of the first to the fourth cervical nerves. It is covered by the sterno-mastoid muscle, and rests upon the THE NEPvVOUS SYSTEM 319 ^11 II 'si II 1 Ji:i III il;^ V iH iliS :§\\.A ^ .1 «5..-" Fio. 147. 320 HUMAN ANATOMY. scalenus medius and levator anguli scapulae muscle. It gives ofi:— Superficial. Superficialis colli. Occipitalis minor, Auricularis magnus. Supraclavicular. Deep. Communicating, Communicantes hypoglossi. Muscular, Communicating, Phrenic, Muscular. The phrenic nerve, or internal respiratory of Bell, is derived from the third and fourth cervical nerves, with a branch from the fifth. It descends into the chest between the subclavian artery and vein, between the pericardium and the pleura, to the diaphragm, to which it is distributed. The course of the two nerves differs in the thorax {vide Mediastinum). The bkachial plexus is formed by the anterior branches of the four lower cervical and the first upper dorsal nerves — the fifth, sixth, and seventh forming one cord, and the eighth cervical and first dorsal another cord. Below the line of the clavicle, both these trunks divide, the adjacent cords of the two upper uniting to form the posterior, and the remaining cords forming the outer and inner cords respectively, receiving their names from their relative position to the subclavian artery. Each of these cords again bifurcates, the adjacent divisions at the outer ending of the cords uniting over the artery to form the median nerve, the other divisions forming the musculo-cutaneous, ulnar, circumflex, and musculo-spiral, the two latter being the divisions of the posterior cord. Its branches are: — Above the clavicle : — Comrriunicating, arises from the fifth cervical, and passes to the phrenic ; Muscular, supply the scaleni, rhomboidii, longus colli, and Bubclavius ; Posterior thoracic, long thoracic, or external respiratory of Bell, arises by five roots from the fifth to seventh cervical nerves, which unite within the scalenus medius muscle and descend to supply the serratus magnus ; Suprascapular, passes beneath the trapezius, through the suprascapular notch, to supply the shoulder-joint and supra- spinatus muscle. Below the clavicle : — The anterior thoracic nerves are two in number : an external branch from the outer cord and an internal from the inner cord. THE NERVOUS SYSTEM. 821 322 HUMAN ANATOMY. The former supplies the pectoralis major and the latter the pec- toralis minor, and sending branches to the pectoralis major. The subscapular nerves are three in number. The upper subscapular supplies the subscapular muscle, the lower sub- scapular nerve supplies the teres major and subscapularis, the middle or long subscapular supplies the latissimus dorsi. The circumflex nerve accompanies the posterior circumilex artery, passing through the space formed between the teres major, teres minor, and long head of the triceps to supply the shoulder- joint and skin over the shoulder and the neighboring muscles. It gives off two branches, an upper and a lower branch. The musculo-cutaneous nerve pierces the coraco-brachialis muscle and descends the arm to supply the brachialis anticus, biceps, coraco-brachialis, and the skin of the forearm. At the outer border of the tendon of the biceps, above the elbow, it be- comes superficial and divides into two branches — ^the anterior descending the radial side of the forearm to the wrist and supply- ing the skin of that part, the posterior branch descending the back part of the radial side of the forearm to the wrist to supply that part. The internal cutaneous nerve descends the inner side of the arm together with the basilic vein to about its middle, where it becomes cutaneous, supplying the skin of this region. It has an anterior and a posterior branch. The lesser internal cutaneous nerve of Wrisberg is derived from the inner cord, and receives filaments from the eighth cer- vical, first dorsal, and the intercosto-humeral nerve, and is dis- tributed to the skin on the inner side of the arm. The median nerve, formed by a root from the outer and inner cord of the brachial plexus, descends the arm, crossing the brachial artery from its outer to its inner side at the bend of the elbow. Its branches are: — Muscular branches, supply all the superficial anterior muscles ex- cept the flexor carpi ulnaris ; Anterior interosseous, supplies all the deep anterior muscles except the inner half of the flexor profundis digitorum; Palmar cutaneous, crosses above the annular ligament, divides into two branches to supply the ball of the thumb and the palmar surface of the hand; Branches to the muscles of the thumh, supplies the opponens, outer head of the small flexor, and the abductor; Digital hi-ancJies, supply both sides of the thumb, index and middle and the radial side of the ring finger. The ulnar nerve descends the inner side of the axillary artery to the middle of the arm, where it crosses the internal head of the triceps, and accompanies the inferior profunda artery to THE NERVOUS SYSTEM. 323 the elbow, where it passes between the heads of the flexor -carpi ulnaris. Its branches are, in the forearm: — Articular (elbow), supplies the elbow-joint; Muscular, to inner half of the deep flexors and flexor carpi ulnaris; Cutaneous, supplies the skin of the palm; Dorsal cutaneous, supplies both sides of the little finger and the ulnar side of the ring flnger; Articular (wrist), supply the wrist-joint. In the hand : — Superficial palmar, supplies the skin of both sides of the little and the ulnar side of the ring finger and palmaris brevis muscle ; Deep palmar, supplies the interossei, lumbrieales, adductor poUicis, and inner head of flexor brevis poUicis. The musculo-spiral nerve, derived from the posterior cord of the brachial plexus, descends the arm in front of the teres major and latissimus dorsi muscle, accompanies the superior profunda artery through the musculo-spiral groove to the outer side of the elbow, between the supinator longus and the brachialis anticus, where it divides into the radial and posterior interosseous nerves. Its branches are : — Muscular, supply the supinator longus, external carpi radialis longior, triceps, anconeus, and brachialis anticus; Radial, descends in the course of the radial artery to three inches above the wrist, where it becomes superficial, and supplies the adjoining sides of the thumb and index, index and middle, middle and ring fingers. Cutaneous, supply the outer side of the arm, elbow, and radial side of forearm and wrist ; Posterior interosseous, passes through the supinator brevis muscle, and supplies, as it descends, all the posterior brachial and radial muscles, except those supplied by the muscular branches of the ulnar. It has a ganglion upon it. Dorsal Nerves. — The posterior divisions of the dorsal nerves subdivide into the external and internal branches to supply the muscles of the skin of the back. The anterior divisions of the dorsal nerves (intercostal nerves), twelve in number, supply the walls of the abdomen and thorax. They are divided into two sets — ^the six upper, distributed for the most part to the walls of the chest, and the six lower, to the walls of the chest and abdomen. The upper six dorsal nerves run between the two sets of the intercostal muscles, accompanied by the intercostal vessels, to supply the skin over the front of the chest and the mammae. Their branches are : — Lateral cutaneous, divide into two branches, anterior and posterior; Anterior branches, to the skin of the chest and mammse principally; Posterior branches, to the skin over the scapular and lower dorsal regions. 324 HUMAN ANATOMY. The jirst intercostal nerve is not distributed to the skin, hut crosses the axilla under the name of the intercosto-humeral nerve; it joins the brachial plexus. The six lower dorsal nerves have received the name of lower, or abdominal intercostal nerves, from their distribution. The last dorsal one is of large size, and gives off a branch, the dorsi lumbar nerve, to join the lumbar plexus. Lumbar Nerves. — The posterior divisions of the lumbar nerves have the same distribution as the other spinal nerves. The anterior divisions of the upper four lumbar nerves unite to form the lumbar plexus. The fifth, with a branch from the fourth, joins the sacral nerves to form the lumbo-sacral cord. The lumbar plexus is formed by the communicating loops from the anterior branches of the first four liunbar nerves and a branch from the last dorsal. Its branches are: — 1. Ilio-hypogastric, divides into two branches : — (a) Iliac branch, supplies the skin of the gluteal region and the oblique muscles of the abdomen ; (h) Hypogastric branch, supplies the skin of the hypogastric region and the oblique muscles. 2. Ilio-inguinal, supplies the skin of the inner and upper part of the thigh, the scrotum, and labium (in female). 3. Genito-crural, passes through the psoas muscle and di- vides into two branches: — (a) Genital branch, follows the spermatic cord to supply the cre- master muscle ; in female, supplies round ligament ; (h) Crural branch, descends in the sheath of the femoral vessels to supply the skin in front of the thigh. 4. External cutaneous emerges below the anterior superior spine of ilium and divides into : — (a) Anterior branch, to skin of outer and front aspect of thigh; Cb) Posterior branch, to skin of outer and back aspect of thigh. 5. Obturator, follows the brim and outer wall of pelvis to foramen (obturator), which it pierces to enter thigh, (a) Anterior branch, supplies the femoral artery; (i) Posterior branch, supplies the adductor muscles ; (c) Articular branch, supplies the synovial membrane of knee-joint. 6. Accessory obturator, sends a branch to the hip-joint and one to join the anterior branch of the obturator nerve. 7. Anterior crural, the largest branch of the lumbar plexus, descends through the psoas muscle, beneath Pouparfs ligament. Trom Las/ D.irjf.i/ ^-_Toffidvpa7i^offpojitorT/uij7i 7'o sk I iL oTvr uumrAnhU • Fig. Utf. Lunihiir plexus ii'id liranches. THE NERVOUS SYSTEM. 325 into the thigh, where it divides into an anterior or cutaneous branch, and a posterior or muscular branch. It supplies all the muscles and the front of the thigh, excepting the tensor vaginae femoris, gives branches to the knee, and supplies the skin of the inner side in front of the thigh, and to the leg and foot. Its branches are, from the anterior division : — (a) Middle cutaneous, to sartorius muscle and skin of fiont of thigh; Cb) Internal cutaneous, supplies the skin in the inner aspect of the leg; (c) Long saphenous, or internal saphenous, passes through Hunter's canal, accompanies the internal saphenous vein, to supply the skin of the inner side of the foot. From the posterior division: — (a) Muscular, to the muscles of the anterior and lateral aspects of the thigh; Cb) Articular, to the knee-joint. The saceal nerves, five in number, divide into anterior and posterior nerves. The four upper sacral nerves, with the fifth lumbar, and a filament from the fourth, the two latter forming the lumbo-sacral cord, together form the sacral plexus. It lies upon the anterior surface of the pyriformis muscle, and is separated from the viscera, the sciatic and pudic branches of the internal iliac artery, by the pelvic fascia. Its branches are : — 1. Muscular, supply the obturators, gemelli, quadratus femoris, and pyriformis. 3. Superior gluteal, supplies the glutei muscles and tensor vagina femoris. 3. Pudic, accompanies the internal pudic artery, passing out of the great saero-sciatic foramen, around the spine of the ischium to re-enter the lesser sacro-sciatic foramen. It gives off:— (a) Inferior hsemorrhoidal, supplies the skin of the anus; (b) Perineal, follows the course of the superficial perineal artery to supply the perineal structures generally; (o) Dorsal nerve of the penis, follows the course of the correspond- ing artery to supply the skin of the glans, prepuce, and penis generally. 4. Small sciatic, supplies the skin of the thigh, leg, perin- eum, and the gluteus maximus muscle. Its branches are : — (a) Inferior gluteal, to the gluteus maximus; (i) Internal cutaneous branches, to the skin of the inner and upper part of the thigh, and one branch, the inferior pudendal, supplies tha skin of the scrotum in the male and labium in the female; 326 HtJMAN ANATOMY. (e) Ascending, cutaneous branches supply the skin over the gluteus maximus. 5. The great sciatic nerve, the largest in the body, passes out of the great saero-seiatic foramen, and descends between the tuberosity of the ischium and the great trochanter to the lower third of the thigh, where it divides into the internal and external popliteal. It gives off: — (a) Muscular branches, to the muscles on the inner and posterior aspect of the thigh; (i) Articular branches, to the hip-joint. The internal popliteal, descends through the popliteal space to the arch of the soleus muscle, where it becomes the posterior tibial. It gives off : — (a) Articular branches, to knee-joint; (h) Muscular branches, to the muscles on the posterior aspect of the leg; (c) External, or short saphenous nerve, passes between the two heads of the gastrocnemius muscle, and descends the leg, receiving the eommunicans peronei branch from the external popliteal, around the outer malleolus to supply the skin of the outer side of the foot. The posterior tibial nerve descends the leg in company with the posterior tibial vessels to below the inner ankle, where it divides into the external and internal plantar nerves. It gives off:— 1. Muscular branches, to the deep muscles of the calf ; 2. Plantar, cutaneous branch, supplies the skin on the 'inner side of the sole and heel; 3. Articular brauch, to ankle-joint. The internal plantar accompanies the corresponding artery to the inner side of the foot and gives off: — 1. Cutaneous branches; 2. Muscular branches; 3. Articular branches ; 4. Four digital branches, supplying both sides of the first three toes and the inner side of the fourth. The external plantar supplies by a superficial branch the outer side of the fourth and both sides of the fifth toes, and gives off a deep, or muscular branch. The external popliteal or peroneal nerve descends to the inner side of the biceps tendon, pierces the peroneus longus about offYt'atattxLa4canAToe, FJi.i. 150. Harral plexus ; ijcl hirariches THE NERVOUS SYSTEM. 327 one inch below the head, and divides into the anterior tibial and musculo-cutaneons. It gives off : — 1. Articular branches to knee-joint; 2. Cutaneous branches, one of which is the eommunieans peronei, which joins the short saphenous. The anterior tibial accompanies the corresponding artery on its outer side to the ankle, where, after giving off an articular branch, it divides into : — 1. External or tarsal branch, to the tarsal and metatarsal joints, and 2. Internal, to the dorsal adjoining sides of the great and second toes. The musculo-cutaneous gives off : — Internal branches, supply skin of inner side of foot and ankle, and inner side of great toe; 2. External branch, to the dorsum of the adjacent sides of the second, third, fourth, and fifth toes. 328 HUMAN ANATOMY. Table of the Spinal Nerves. First cervical nerve, Second cervical nerve, Posterior division (suboccipital). Anterior division (occipital). Posterior division. Anterior division. h Complexns, recti, and obliqui muscles. } Unites with the second cervical nerve, and supplies the anterior recti and rectus lateralis muscles. } External brajich. Spleniua, complexus, trachelo-mastoid muscles. Internal branch (great . oEcipital), With branch from third cervical, supplies the skin of the scalp. Small occipital, Branch to great auricular. Branch to superficial cervical. Branch to communicans nom. Third cervical nerve. Fourth to eighth cervical ' nerves. Posterior division. Anterior division. Posterior divisions, Anterior division of fourth. External and internal branches. Splenius, semispinalis, complexus, trachelo-mas- toid muscles. Branches to form the great auricular, superficial cervical, and communicantes minor. External branches ', } Internal \ branches, J Muscles of the side of the neck. Skin and larger muscles of the neck. Anterior 1 iivisiona, [■ i to eighth, J Branch to phrenic. Branches to the trapezius, scalenus medius, levator anguli scapulae mus- cles. Unite with the first dorsal nerve to form the brachial plexus. Cervical plexu8.«{^,,'JP"gh"g^ < {Superficialis colli, Auriculasis magnus, Occipitalis minor. Descending, ■< Supraclavicular. ' Formed by the anterior divisinna of the first to the fourth cervical nerves. THE NERVOUS SYSTEM. 329 Table of the Spinal Neeves. — (Continued.) Internal set. Cervical plexus f Deep [continued), \ branches. " Above the clavicle. Brachial plexus,i< Below the clavicle. Communicating, Muscular, { ^u^^jes. Phrenic, Communicans nonL „_ , T i f Muscular, External set. | Communicating. \ Tovphrenio. (• Communi- cating, Muscular, Posterior thoracic. Supra- scapular, "1 Scaleni., rhomboidei. Ion- J gus colli, subclavius. > Serratus magnus. ' (External respiratory nerve of Bell.) 1 Shoulder-joint and supra- / spinatus muscle. External f Pectoralis branch, \ major. "lor cic. Internal branch. Pectoralis minor, Pectoralis major. Three sub- scapular. Circum- flex, Musculo- cutaneous, Subscapular muscle. Upper, I Lower, < Teres major. L Middle, Upper, Lower, Anterior branch. Posterior . branch. { Latissimus dorsi. Shoulder- joint, and muscles and skin about it. Integument of palmar sur- face of the wrist. Integument of dorsal sur- face of the wrist. > Formed by the anterior branches of four lower cervical and first dorsal nerves. 330 HUMAN ANATOMY. Table of the Spinal Nerves. — (Continued.) Brachial plexus, below the clavicle, contin'd, Internal cutaneous. Anterior and posterior branches, Integument of inner side of the arm. Lesser internal cutaneous (Wrisberg's) :i Integument of the inner side of the arm. Median, Ulnar, in forearm. {Superficial anterior mus- cles, except flexor carpi ulnaris. I Deep anterior muscles, ex- cept the inner half of flexor profundus digi- torum. Palmar f Ball of thumb and palmar cutaneous, \ surface of the hand. Branch to f Opponens, outer head of the muscles of -j small flexor and ab- the thumb, I ductor. r Both sides of thumb, in- Digital dex and middle fingers, branches, and radial side of ring finger. Articular, i (Elbow). TiT,.„„„i„- / Inner half of deep flexors. Muscular. | ^^^ ^^^^^ ^^^ ^j^^^j^; Ulnar, in hand. Cutaneous, < Skin of the palm { ■[ (Wrist) Dorsal cutaneous. Both sides of little finger, and ulnar side of ring finger. Articular, Superficial palmar. Deep palmar. Skin of both sides of little finger, and ulnar side of ring finger, and palmaris brevis muscle. Interossei lumbricales, ad- ductor pollicis, inner head of flexor brevis pollicis. THE NERVOUS SYSTEM. 331 Table of the Spinal Nerves. — (Continued.) Brachial plexus, below the clavicle, cont'd. Miisculo- spiral. Dorsal nerves. Posterior divisions. Anterior divisions' Muscular branches. Radial, Lumbar Posterior divisions. Anterior divisions. '■{ Cutaneous, Posterior interosseous. External branches. Internal branches. Upper six (thoracic intercostals). Supinator longus, extensor carpi radialis longior, tri- ceps, anconeus, brachialis anticus. Supplies adjoining sides of thumb and index finger, index and middle, middle and ring fingers. Inner side of arm, elbow, radial side of forearm, and wrist. Posterior brachial and radial muscles, except those sup- plied by ulnar (muscular branches). Muscles and skin of the back. First intercostal crosses the axilla to join the lesser in- ternal cutaneous (Wris- berg's). Lower six (thoraeico- abdominal intercostals). External branches. Internal branches. Skin of chest and mammae. Skin over scapula and lower dorsal region. The last dorsal gives off the dorsi lumbar to the quad- ratus lumborum muscle. Ante- rior, Lateral >- cutane- - f ous. Poste- L rior. . Muscles of back. the skin of the The four upper unite to form the liunbar plexus. The fifth, with a branch from the fourth, joins the sacral nerves to form the lumbo-sacral cord. ' To the walls of chest and abdomen. 332 Luinbar plexus. HUMAN ANATOMY. Table of the Spinal Nerves. — (Continued.) Iliac, nio- hypogastrie, ' Anterior crural, f Skin of gluteal region, ob- \ lique muscles of abdomen. Hypo- / Skin of hypogastric region rie, I gastric and oblique muscles. Ilio- inguinal, Genito- crural. External cutaneous. Obturator, Accessory obturator, f Skin of inner and upper part of thigh, scro- \ turn, labium (in female). Genital i Cremaster muscle, round liga- ' \ ment (in female). Crural, i Skin of front of thigh. Anterior, ( S"^™. °^ ""^^"^ ^'■°"* ^^P^''* °* ' \ thigh. Posterior, ( ^^J". °t ""^^'^ ^^"^ ^^P^<=* °* ' I, thigh. Anterior, < Femoral artery. Posterior, < Adductor muscles. { Articular, { ^^^°^X^ membrane of knee- Branch to hip-joint, branch to anterior branch of obturator nerve. Anterior division. Posterior division, Middle cutaneous. Internal cutaneous, Long saphenous. Muscular, Sartorius muscle, skin of thigh (ant.). Skin of inner aspect of leg. Skin of inner side of foot. Muscles of the an- terior and lateral aspect of thigh. Articular, < Knee-joint. Sacral nerves. " Posterior divisions, Muscles and skin of the back. Sacral plexus, THE NERVOUS SYSTEM. 333 Table of the Spinal Nerves. — (Continued.) f External ■» -j and internal > '' (. branches, J r / The four upper j< divisions, \ cord form the sacral plexus. Muscular / Obturators, gemelli, quadratus femoris, and muscular, ^ pyriformis. f^ufpa?' f Crlutei muscles, and tensor vaginse femoris. hemorrhoidal, } ^'^^^ °^ *'^« *""«• Perineal, \ Perineal structures. Dorsal nerve of 1 oi • r ii. > Skin of the penis. Pudic, the penis. Articular, | To hip- joint. Small sciatic. Inferior gluteal. Internal cutaneous, > Gluteus maximus. Branch to skin of inner and upper part of thigh. Inferior pudendal, skin of scrotum, labium in fe- male. Ascending cutaneous, Great sciatic. > Skin over gluteus maximus. Articular, < Hip-joint. Muscular, -f Muscles of inner and pos ' \ tenor part of thigh. Bifurcation, { ^^^^^^^ } PopliteaL Internal popliteal. brSet } TO knee-joint. Muscular \ To muscles of the posterior aspect of branches, / the leg. 334 HUMAN ANATOMY. Internal popliteal {contintied), Table of the Spinal Nerves. — (Continued.) '^"^litlt''* } S'^'" °^ ""t^' ^^^^ °f *^« ^°°*- Muscular. { De^P^ muscles of the Plantar f Skin of inner side of cutaneous, \ sole and heel. Articular, < To ankle-joint. Posterior tibial (continuation) , Bifurca- f Internal plantar, tion, \ ~ Internal plantar. External plantar. External plantar. Cutaneous, Muscular, Articular, . Four digitaL Supplies the outer side of the fourth and both sides of the fifth toes. Mus- cular branch. External popliteal (peroneal), Articular, < To knee-joint. < Communicans peroneL ] Two cutaneous. Anterior tibial. Musculo- cutaneous, External or tarsal branch, Tarsal and metatar- sal joints. (Inner and dorsal, ad- joining sides of great and second toes. Internal f To skin of inner side branches, \ of foot and ankle. C To dorsum of adja- External cent sides of third, branch, fourth, and fifth toes. Ir,/>T,iiT! Ctjralid^irtery y,, .'£«>-„' "■— ..^-^^ ridiatt^tiTe(u SfihfniJ Pai^rtinj 6analiOit _ _ , . . . ., ^ \ ^ ~ XarvfSuB/^A.^i.aLPwtfBsai^m^iaJXjm' To iiaiff^i'i/i it/"^./! a/'J'tit h mafOAtr aniiitt bra./rchfs J'iV.-.'i.f'f' S,tr/u/.V, , I , /hnn (V..V,.,,v./.r. THE NERVOUS SYSTEM. 335 THE SYMPATHETIC NERVOUS SYSTEM. The sympathetic system, like the cerebro-spinal axis, is double, consisting of a gangliated cord on each side, extending the entire length of the vertebral column, and numerous nerve- fibres, both communicating, by which it anastomoses with the cerebro-spinal axis, and distributory, by which the blood-vessels and viscera are supplied. Branches of the ganglionated cords ascend through the carotid canal to communicate with the ganglia of the fifth cranial nerve, and also with each other through the ganglion of Eihes, situated upon the anterior communicating artery. They also communicate below in the ganglion impar, located in front of the coccyx. The ganglia of each cord correspond very nearly in position and number to the vertebrse, except in the cervical region, where there are but three: cervical three, dorsal twelve, lumbar four, sacral five. From the ganglionated cords three important plexuses are given off — the cardiac, solar, and hypogastric — situated in the thoracic, abdominal, and pelvic cavities respectively. Cranial Ganglia — Ganglia Connected with the Fifth Cranial Nerve. — Besides the Gasserian ganglion upon the root there are four — ophthalmic, spheno-palatine, otic, and submaxillary — each of which have three branches of communication or roots — ^motor, sensory, and sympathetic — ^besides several branches of distribu- tion. The ophthalmic, or ciliary, ganglion is situated in the orbit between the external rectus muscle and optic nerve upon the first division of the fifth cranial nerve. Its sensory root is from the nasal branch of ophthalmic, motor root from the motor oculi or third and sympathetic from the cavernous plexus. Its branches are the short ciliary nerves (^vide Bye). Spheno-palatine, or Meclcel's, the largest, is situated in the spheno-maxillary fossa upon the superior maxillary, or second division of the fifth. Its sensory root is derived from the superior maxillary, its motor root, from the facial, through means of the Vidian, and its sympathetic from the carotid plexus. Its branches are : — 1. Ascending; „ T-, ,. (■ Anterior, or large palatine, 2. Descending, or I j^j^jj^^ ^^ external palatine, paianne, ^ Posterior, or small palatine; 336 HUMAN ANATOMY, 3. Internal, 5 Superior nasal, ' I Naso-palatine; 4. Posterior, Vidian, Large superficial petrosal. Deep petrosal, Pharyngeal branch. The Vidian nerve passes from the back part of Meckel's ganglion through the foramen lacerum medium, where it divides into the large superficial and deep petrosal nerves. It gives off the upper posterior nasal branches to the mucous membrane of septum, orifice of Eustachian tube, and roof of the nose. Or, more correctly, the Vidian is formed by the union of the great petrosal (large superficial petrosal) from the facial and the large deep petrosal from the carotid sympathetic, runs forward through the Vidian canal, and joins the spheno-palatine ganglion (Meck- el's). In this description, the nerves given off to the nasal mucous membrane must be considered branches from the gan- glion inclosed in the same sheath. The large superficial petrosal branch enters the cranium through the foramen lacerum medium, passes beneath the dura mater and Gasserian ganglion, enters the hiatus Fallopii, re- ceives a branch from the tympanic branch of the glosso-pharyn- geal (Jacobson's), and through the aquseductus Fallopii to termi- nate in the geniculate ganglion of the facial nerve. The large deep petrosal branch crosses the foramen lacerum medium to the carotid canal, where it joins the carotid plexus of the sympathetic. The pharyngeal or ptery go-palatine nerve desends from the back part of the ganglion through the pterygo-palatine canal, to supply the upper part of the pharynx. Besides the Vidian and its branches there are two other petrosal nerves {vide Facial Nerve), the small and external petrosal. The small petrosal connects the geniculate ganglion of the facial, within the aquaeductus Fallopii, with the otic ganglion. The external petrosal connects the geniculate ganglion of the facial, within the same canal, with the sympathetic plexus of the middle meningeal plexus. The otic ganglion (Arnold's) is placed below the foramen ovale upon the inferior maxillary nerve, or third division of the fifth. Its sensory root is derived from the auriculo-temporal branch of the inferior maxillary; the motor root, from the internal pterygoid branch of the same, the sympathetic root, from the plexus on the middle meningeal artery. Branches are distributed to the tensor palati and tensor tympani muscles. THE NERVOUS SYSTEM. 337 The submaxillary ganglion is placed above the deep portion of the submaxillary gland. Its sensory root is derived from lingual branch of inferior maxillary ; its motor root, from the facial nerve through a branch of the chorda tympani; and its sympathetic, from the plexus of the facial artery. Beside these ganglia, situated upon branches of the trifacial nerve, the following are found within the cavity of the cra- nium : — Ganglion of Ribes, on the anterior communicating artery; Ganglion of Bidder, on the middle meningeal artery ; Ganglion of Laumonier, on the internal carotid artery; Ganglion of Cloquet, on the naso-palatine nerve in the incisor fossa; Ganglion of Bochdaleck, near the spheno-palatine ganglion. Cervical Ganglia. — The cervical ganglia consist of three, — superior, middle, inferior, — of which the superior is the largest and the middle the smallest. These are connected above with the cranial ganglia, below with the thoracic ganglia, and com- municate with each other. The superior cervical ganglion lies opposite the second or third cervical vertebra, behind the internal carotid artery. It is connected with the middle ganglion, the upper four cervical, and tenth and twelfth cranial nerves, distributes branches to the carotid (internal), cavernous, and pharyngeal plexuses, and gives off the superior cardiac nerve to the cardiac plexus. The middle cervical ganglion lies upon inferior thyroid ar- tery, opposite fifth cervical vertebra. It is connected with the upper and lower ganglia and spinal nerves, and gives off the middle cardiac nerve to cardiac plexus. The inferior cervical ganglion lies internal to the superior intercostal artery, below the last cervical vertebra. It is con- nected to the middle ganglion, first thoracic, lower cervical nerves, forms the vertebral plexus, and gives off the inferior car- diac nerve to cardiac plexus. Thoracic ganglia lie upon the heads of the ribs on each side of the vertebral column. They are connected with the inferior cervical ganglion above, the lumbar ganglion below, the dorsal spinal nerves behind, and give off internal branches divided into two sets of six each — upper and lower. Internal branches from upper set are distributed to the pulmonary and aortic plexuses, and internal branches from lower set unite to form the three splanchnic nerves, — great splanchnic, lesser splanchnic, renal splanchnic, — distributed respectively to the semilunar ganglion, renal and suprarenal plexiises, and to renal and cceliac plexuses. 338 HUMAN ANATOMY. The lunibar ganglia lie along inner margin of the psoas muscle. They communicate above and below with the other ganglia and with the lumbar spinal nerves, and give ofE internal branches which form the hypogastric plexus. The sacral ganglia are situated internal to the anterior sacral foramina. They unite below in front of the coccyx in the coccygeal ganglion, or ganglion inipar. They communicate with the sacral nerves, join the pelvic plexus, and send branches on the middle sacral artery. Cakdiac Plexus. — The cardiac plexus, formed from the superior, middle, and inferior cardiac nerves from the cervical ganglia, and the cardiac branches from the pneumogastric and recurrent laryngeal, consists of two portions — ^the superficial and deep. The superficial cardiac plexus, situated beneath the arch of the aorta, is formed by the left superior cardiac nerve and the left inferior cervical cardiac nerve, and contains the cardiac ganglion of Wrisberg. It forms part of the anterior coronary plexus, and sends branches to the left anterior pulmonary plexus. The deep cardiac plexus lies between the arch of the aorta and trachea, and receives all the cardiac nerves except the two mentioned above. It forms part of the anterior coronary and posterior coronary plexuses. The SOLAR PLEXUS^ or "abdominal brain," consists of ganglia and a net-work of nerve branches, formed chiefly from the branches of the two great splanchnic nerves and branches from the right pneumogastric. It is situated between the aorta and the crura of the diaphragm and the stomach, and surrounds the superior mesenteric artery and cceliac axis. Its ganglia are two crescentic ganglionic masses — ^the semi- lunar ganglia (the largest in the body) — situated in front of the crura of the diaphragm. They are composed of smaller ganglia aggregated together. From the solar plexus and semilunar gan- glion are derived numerous branches which are distributed as plexuses over all the abdominal arteries, as follows : — Coeliao, Phrenic, Aortic, Gastric, Suprarenal, Superior mesenteric. Hepatic, Renal, Inferior mesenteric. Splenic, Spermatic, Ovarian. The HYPOGASTRIC PLEXUS is situated below the bifurcation of the aorta, in front of the sacrum, and is formed by branches from the lumbar ganglia and aortic plexus. It divides into two parts which, with branches from the sacral ganglia and nerves, become the pelvic plexuses. THE NERVOUS SYSTEM. 339 The inferior htpogastkic or pelvic plexus on each side gives ofE the following branches : — Vesical plexus, Inferior hsemorrhoidal, T)_„„i„4.;^ i„„ ( Small cavernous nerves, "I 4.„ ^„- Prostatic plexus, ■( t ' > to penis, ^ ' I. Large cavernous nerves, I *^ ' Vaginal plexus, Uterine plexus. ORGANS OF SPECIAL SENSE. THE NOSE. The nose, the special organ of smell, consists of two parts, — the external prominence, or nose proper, and the internal cavities, or nasal fossse. BEPTA OF S AL branch) SUPERIOR r NASAL i Fig. 152. The olfactory nerves and nerves of common sensation to the nose, (Eclcleu.) The nose proper is made up of a cartilago-osseous frame work covered with muscles {vide Muscles) and skin, and lined with mucous membrane. The osseous portion is formed by the mar- gins of the anterior meatus {vide Osteology). The cartilaginous portion consists of an upper and lower lateral cartilage on either side, and the nasal septum dividing the nasal cavity into two nasal fossfe. Arteries are from nasal branch of ophthalmic and infra- orbital, nasal artery from superior coronary, and branches of the lateralis nasi. Veins empty into the ophthalmic and facial veins. Nerves from iufratrochlear, infraorbital, and facial. (340) ORGANS OF SPECIAL SENSE. 841 Fig. 153. Nasal meatuses and accessory cavities. 1, right termination of the left frontal sinus; 2, right frontal sinus; 3, probe extending from the right frontal sinus through the infundibulum into the right nasal fossa; 4, ethmoid cells; 5, large opening into the maxillary sinus; 6, anterior antrum of the sphenoid bone; 7, posterior sphenoid antrum; 8, middle nasal meatus ; 9, inferior meatus ; 10, inferior turbinated bone; 11. probe extending through the nasal duct. {Bishop.) 342 HUMAN ANATOMY. The nasal fossce are the two bony cavities already described (vide page 3C). They are lined throughout with mucous mem- brane called Schneiderian membrane, which is continuous with the antrum of Highmore, frontal, ethmoidal, and sphenoidal sinuses, with the conjunctiva through the lachrymo-nasal duet, with the pharynx through the posterior nares, and with cavity of the tympanum through the Eustachian tube. Its epithelium is columnar, ciliated in the lower part and the sinuses, and is Sis as front. Ximmlna pspyr. OBa. ethmoid. Foramfna ethToofcI. Froo. ortit. oaa. palat. Corpus osa. sphenoid. For. optic, Flamra orbit. Bnp. Sella turcica V Spina trooWear. PAra orbit, oss. front. Fosea boccI lacrito. Oa nosald 'acies orbit. xOl. 00. nncia. ose. ethmoid. Froo. ethmoid, conohao naeal. inf. Froo. maxill. conchaa naeal. inf. For epbenopalat. Fossa pterygopalat. lamina lat. proo. pteryg. Froo. pyramid, osb. palat. maxilla rls Farfl perpendio. oas. palat. Sonde in canal, palat. Fig. 1B4. Inner wall of orbit and adjacent parts. (His.) (Ball.) columnar but not ciliated in the membrane to which the olfactory nerve is distributed. Arteries are anterior and posterior ethmoidal from the ophthalmic to the roof, frontal and ethmoidal sinuses, spheno- palatine from the internal maxillary to the mucous membrane of the spongy bones, septum, and meatuses, and alveolar from the internal maxillary to the antrum. Veins empty into facial, ophthalmic, and beginning of great longitudinal sinus. ORGANS OF SPECIAL SENSE. 343 Nerves are : — Olfactory, or first cranial, the special nerve of smell, is dis- tributed over the upper third of the septum, and superior turbin- ated bones. Anterior dental branch of superior maxillary to the inferior turbinated bone and inferior meatus. Nasal branch of ophthalmic, to the outer walls and septum. Besides these, the spheno-palatine ganglion, the vidian, the supe- rior nasal branch, naso-palatine, and anterior palatine also send branches to the parts. Horizontal section of right orbit. 1, Horner's muscle; 2, septum or- bitale; 3, fibres of the orbicularis palpebrarum muscle; 4, tarsal plate; 6, conjunctival sac; 6, outer palpebral ligament; 7, temporal muscle; 8, wall of the orbit; 9, external rectus muscle; 10, orbital fat; 11. optic nerve; 12, internal rectus muscle; 13, Inner check ligament; 14, inner wall of the orbit; 16, attachment of the capsule of Tenon (16) to the conjunctiva; 17, Tenon's space; 18, outer check ligament; 19, lachrymal gland; 20, ciliary process; 21, iris. (After Gerlach.) {Ball.) THE EYE. The ORGAN'S of sight comprise the eyeballs and their ap- pendages and the optic nerves. The eye is a spherical organ, situated in the anterior part of the skull, protected in front by several appendages, acted upon by muscles, and supplied by blood-vessels and nerves. Within the orbit it rests upon a bed of fat, from which it is separated by a membranous sac — ^the capsule of Tenon. 344 HUMAN ANATOMY. It presents the following: — Diameters. — Antero-posterior, in the adult, .950 inch; transverse, .925; vertical, .915; oblique, .943. Anterior and posterior poles are the geometric centres of the cornea and fundus respectively. Optic axis is a straight line passing through the cornea and posterior pole of the eye. Line of vision, or visual axis, is an imaginary line which connects the point of fixation with the fovea centralis, through the nodal point, usually to the outer side of the centre of the pupil. It forms with the optic axis as it cuts the cornea, the visual angle — an angle of from 3° to 7°. Antero-posterior section of EyebaU, 1, optic nerve; 2, sclera; 3, cornea; 4, spaces of Fontana; B, choroid; 6, ciliary muscle; 7, ciliary processes; 8, iris; 9, retina; 10, Jacob's membrane; 11, anterior cham- ber; 12, posterior chamber; 13, pupillary area; 14, aqueous humor; IB, hyaloid membrane; 16, canal of Stilling; 17, canal of Petit; 18, vitreous humor; 19, capsule of the lens; 20, fluid of Morgagni; 21, lens. (Liveilli.) {.Ball.) Nodal point is an imaginary point — the centre of curvature of the refracting media — where all the luminous rays pass with- out deviation. Equatorial plane, an imaginary plane passing through the centre of the eyeball at right angles to the optic axis, dividing the globe into two hemispheres — -the anterior and posterior. Equator is the line upon the surface of the globe where the equatorial plane cuts it. Meridional planes are imaginary antero-posterior planes co- inciding with the axis. ORGANS OF SPECIAL SENSE. 345 Meridians are the lines upon the surface where these merid- ional planes cut it. The capsule of tenon, tunica vaginalis oculi, is a fascia between the eyeball and the walls of the orbit, isolating the eye- ball and allowing free movement. It is continuous in front with the subconjunctival connective tissue, and behind with the dura mater, through the sphenoidal fissure and optic foramen, and consists of two layers — a visceral layer investing the posterior portion of the eyeball, and a parietal layer lining the cushion of fat on which the eye rests. The inner aspect is lined with flattened endothelial cells, and incloses a lymph space communicating with subdural and sub- arachnoidean lymph spaces of the optic nerve-sheath. It sup- ports the lachrymal gland, is strengthened by numerous fibrous bands, and is pierced by the ocular muscles, inclosing them in imperfect sheaths. The capsule of Bonnet is the name given to the portion posterior to the passage of the tendons, and The capsule of Tenon is then applied to the anterior socket- like half. The eyeball is composed of a large, opaque segment of a sphere, forming about five-sixths of the globe, for the protection of its contents, and a smaller transparent segment of a sphere, implanted upon and continuous with it in front. The optic nerves enter the eyeballs to their nasal side, in the direction of the axes of the orbit. It is composed of three tunics or coats: — 1. Sclerotic and cornea, 2. Choroid, iris, and ciliary processes, 3. Retina; And three humors, or refracting media :— 1. Aqueous humor, 2. Crystalline lens (and capsule), 3. Vitreous humor, or body. The sclerotic coat, so called from its extreme hardness, is thicker behind (one twenty-fifth of an inch) than in front (one-sixtieth of an inch), and presents two surfaces for study — the external and internal. The external surface is smooth and white, and has attached to it the various muscles of the eye. The internal surface is grooved for the passage of the ciliary nerves, and connected to the choroid by a fine areolar layer — the lamina fusca. 346 HUMAN ANATOMY. Behind, and a little to the nasal side, where the optic nerve enters, it presents a perforated appearance — ^the lamina cribrosa — the larger opening of which — ^the porus opticus— transmits the arteria centralis retinae; the others transmit the ciliary vessels and nerves. It is continuous in front with the cornea, over- lapping it a little on its outer margin. In structure it is made up of white, iibrous tissue, with a small quantity of elastic fibres, and connective-tissue corpuscles. It probably contains no nerves. The cornea is the convex, transparent, nearly circular tissue forming the 8,nterior one-sixth of the globe. It is from one twenty-second to one thirty-second of an inch in thickness. Its thickness at the periphery is 1.13 millimetres, hence its pos- terior surface is more curved than the anterior. Its transverse diameter is a little greater than the vertical, owing to the over- lapping of the sclerotic above and below. It is composed of four layers : — 1. Conjupctival epithelium; 2. Cornea proper, { ^^^^^^ '^^^^' I: ^^ t^, '^"^"^ } ^-^-- °^ ^— *• The conjunctival epithelium consists of several layers of cells (columnar, polyhedral, and squamous) covering the ante- rior surface of the cornea, continuous with the conjunctiva. The anterior elastic lamina is the name given to the outer epithelial layer of the cornea proper. The cornea proper is made up of a transparent fibrous struc- ture, identical with the sclerotic, and consisting of about sixty layers or lamina, connected by a cement substance inclosing cor- neal spaces, each of which contains a corneal corpuscle. The posterior elastic lamina consists of an elastic homo- geneous membrane, internal to the proper structure of the cornea, and constituting, with the epithelial lining, the membrane of Des- cemet, or Demours. The structure of the cornea is non-vascular, being nourished by channels representing lymphatic vessels, and continuous with the corneal spaces. The nerves derived from the ciliary nerves are numerous, and form between the outer surface of the cornea proper and the epithelial covering the subepithelial plexus, from which is given off the intraepithelial plexus. The second tunic consists of the choroid, lining the sclerotic coat throughout; the iris, the circular curtain suspended in the ORGANS OF SPECIAL SENSE. 347 aqueous humor and perforated by the pupil ; and the ciliary liga- ment and muscle at the junction of the two. The choroid is a dark, reddish-brown, highly vascular membrane, lining the sclerotic coat from the optic nerve to the ciliary ligament, where it projects backward in a number of processes, the ciliary processes. It is in contact externally with the sclerotic, to which it is attached by the membrana fusca, and internally with the retina. Pig. 157. Vessels of the eye. A, conjunctival vessels; B, anterior ciliary vessels; C, vena vorticosa; D, posterior ciliary arteries; E, central retinal vessels. iBall.) The structure consists chiefly of a dense capillary net-work. The choroid proper consists of an external and an internal layer, and is separated from the sclerotic by the lamina supra choroidea, continuous with the lamina fusca of the sclerotic, and from the pigmentary layer of the retina by the lamina vitrea. The external layer consists of a capillary net-work, derived from the larger branches of the short ciliary arteries, inclosing between the meshes large, star-shaped pigment cells, which are connected together by a delicate stroma. They terminate in the 348 HUMAN ANATOMY. venm verUcosw, which emerge through the sclerotic midway be- tween the optic nerve and the margin of the cornea. The internal layer is also composed of a iine capillary net- work, derived from the short ciliary arteries, and continuous in front with the vessels of the ciliary processes. It is called the tunica Ruyschiana. The vitreous membrane lamina vitrea is a thin, structure- less layer, separating the pigmentary layer of the retina from the membrana Euysehiana. The arteries are from posterior ciliary, short ciliary (prin- cipally in the external layer), and recurrent branches of long and anterior ciliary. The veins converge from the venfe verticosse to form four or five trunks, which pierce the sclerotic midway between the optic nerve and corneal margin to join the cavernous sinus. The nerves are the long and short ciliary. The ciliary processes are a series of sixty to eighty pig- mented vascular processes arranged circularly around the lens behind the iris, and composed of the two internal layers of the choroid folded inward. They are continuous in front with the iris, and are connected behind with the suspensory ligament of the lens. The structure is similar to that of the choroid. The iris is a thin, circular curtain, suspended in the aqueous humor, between the lens and cornea, and perforated by a circular aperture — the pupil. Its circumference is connected with the choroid, and also by means of the ciliary ligament with the sclerotic and cornea. The sclerotic contains in this position, near its Junction with the cornea, a circular canal (lymph canal, or venous sinus), the sinus circularis iridis, or canal of Schlemm. The circumference of the iris is also connected in front with the cornea by ligamentuni pectinatum iridis, derived from the membrane of Descemet. The structure is composed of four elements : — 1. A basement membrane and polyhedral cells, continuous with the membrane of Descemet ; 2. Stroma, consisting of bundles of fibrous tissue and cells; Circular fibres, forming the sphincter of the pupil; Radiating fibres, forming the dilator of the pupil; 4. Pigment, consisting of polyhedral or round pigment cells, dis- tributed for the most part to the posterior surface, which, from its deep-purple tint, has been called the uvea. 3. Muscular fibres, ORGANS OF SPECIAL SENSE. 349 The arteries are from the long and anterior ciliary and from the ciliary processes (from the short ciliary). The veins join those of the ciliary processes and anterior ciliary veins. The nerves are the long ciliary from the nasal branch of the ophthalmic (first division of fifth), and ciliary branches from the lenticular or ophthalmic ganglion. The circular fibres are supplied by the fibres from the third or motor oculi, ihe radiating fibres are supplied by the sympa- thetic. The menibrana pupillaris is a delicate, vascular membrane, closing the pupil in the foetus, continuous at its margin with the iris. It usually disappears about the eighth month, but occa- sionally, persists. 'Hhe. ciliary muscle is a grayish, circular band of unstriated fibres, attached to the fore part of the choroid. It consists of two sets of fibres — ^the external radiating fibres, dilator pupillce — arising from the junction of the sclerotica and cornea, and is inserted into the choroid opposite the ciliary processes; the in- ternal circular fibres, sphincter pupillcB, pursue a circular course around the insertion of the iris. It is the muscle of accommodation, its contraction drawing on the ciliary processes, relaxing the circular fibres described as the ligament of the lens, and allowing the anterior surface of the lens to become more convex by its inherent elasticity. The retina is a delicate, white, nervous membrane, the ex- pansion of the optic nerve. It lines the eyeball, being in con- tact externally with the choroid and internally with the vitreous body. It terminates in front near the ciliary ligament in an irregular margin, the ora serrata, but is prolonged forward as the pars ciliaris to the iris. Its internal posterior surface presents at a point corresponding to the axis of the eyeball a small, round, elevated spot of yellowish color, the macula lutea, or yellow spot of Sommering, in the centre of which is the fovea centralis, a central depression, the region of most acute vision, and about one- tenth of an inch to its nasal side, at a point corresponding to the axis of the orbit, the entrance of the optic nerve, an oval, bluish- white depression, with distinct, often pigmented margins, a re- gion destitute of vision and called the optic dish. Its centre presents the arteria centralis retinae, giving branches to the nasal upper and lower side of the macular region. The structure of the retina is exceedingly complex, con- sisting microscopically of ten distinct layers from within out- ward, as follows: — 350 HUMAN ANATOMY. 1. Membrana limitans interna, the most internal layer of the retina, forms the base of the connective-tissue frame-work of the retina, from which it is derived. 2. The fibrous layer consists of nerve-fibres, the termination of the optic nerve-fibres. FiQ. 158. The structure of the human retina. I, pigment epithelium layer; II, rods and cones; III, granules ot the visual cells; IV, outer plexl- torm layer; V, layer of horizontal cells; VI, layer of bipolar cells; VII, layer of amaUrine cells; VIII, Inner plexiform layer; IX, gan- glion-cell layer; X, layer of nerve-flbres; 1, diffuse amakrine cell; 2, diffuse ganglion cell; 3, centrifugal nerve-fibre; 4, amakrine asso- ciation fibres; 5, neuroglia-cells; 6, Miiller'a radial fibres. (A.tter ereeff.) IBall.) ORGANS OP SPECIAL SENSE. 351 3. The vesicular layer, composed of a layer of large, flask-shaped ganglionic cells. It is wanting in the muscular region. 4. The inner molecular layer is composed of a layer of glandular- like structure, forming a reticulum inclosing minute granules. 5. The inner nuclear layer is composed of three sets of nuclear bodies, the first resembling bipolar nerve-cells, the second without branches, and the third continuous with the radiating fibres, or fibres of Miiller. 6. The outer molecular layer resembles closely the inner molecular layer, from which it differs only by containing branched stellate cells. FIG. 169. Diagram of the retinal vessels. 1, superior temporal artery: 2, superior temporal vein; 3, superior nasal vein; 4, superior nasal ar- tery; 5, inferior nasal vein; 6, inferior nasal artery; 7, inferior tem- poral vein; 8, interior temporal artery; 9, macula lutea; 10, macular veins. (Ball.) 7. The outer nuclear layer is composed of several layers of nuclear cells, separable into two kinds, the rod granules and cone granules, both continuous with the rods and cones of Jacob's membrane. 8. The memhrana Umitans externa is, like the internal limiting membrajie, derived from the radiating fibres, or fibres of MuUer. 9. Jacob's membrane, or the layer of rods and cones, consists of two distinct kinds of cells — the rods and cones, distributed alternately throughout this layer, the rods being much more numerous. The rods are solid, stand perpendicularly to the surface, and con- sist of two portions — an outer striated and an inner granular. 23 352 HUMAN ANATOMY. The cones are flfisk-shaped, with their pointed extremities toward the choroid. They also consist of two portions — an outer striated and an inner granular. 10. The pigmentary layer was formerly described as a layer of the choroid. It has received the name of tapetum nigrum, consisting of a layer of pigmented hexagonal epithelial cells. In many of the lower animals, this layer on the posterior surface is destitute of pigment, and is called from its iridescent lustre the tapetum lucidum. The radiating fibres, or fibres of Muller, consist of connective-tissue fibres, connecting all the layers together, and forming the membrana limitans interna and the membrana limitans externa. The structure of the retina at the macula lutea differs in the following manner: the cones only of Jacob's membrane are present ; the vesicular layer consists of several layers, and in the outer nuclear layer only the cone-fibres are present. At the fovea centralis only the cones of Jacob's membrane, the outer nuclear layer, and the internal granular layer are present. The arteria centralis retina supplies only as far as the inner nuclear layer. THE HUMOKS OF THE EYE. The aqueous humok is a transparent, alkaline, serous fluid, small in quantity, weighing four to five grains, and filling the anterior and posterior chambers of the eye. It is composed of water, 96.7; extractive matters, principally chloride of sodium and albumen, 0.1. The anterior chamber is the space (about 2.7 millimetres in depth) between the iris and the cornea. The posterior chamber is the space between the anterior sur- face of the lens and the iris. It amounts to a space only at the circumference, the two being in contact at the posterior surface. The crystalline lens is a biconvex, transparent, elastic body, suspended within its capsule, surrounded by the ciliary processes. It is more convex on its posterior surface (anterior surface has an average radius of ten millimetres, the posterior six millimetres), and is received into a hollow depression in the hyaloid membrane. It measures about one-third inch in diameter, one-fourth inch in axis, and weighs from four to four and one-half grains. Its index of refraction increases from the periphery to the centre, with an average of 1.4371. Its composition consists of about 60 per cent, water ; soluble albuminous matter, 35 per cent. ; insoluble albuminous matter, S.5 per cent.; cholesterine and fat, 2 per cent ORGANS OF SPECIAL SENSE. 353 The structure of the lens consists of concentric lariiinse, made up of hexagonal prisms, about one five-thousandth inch in breadth, united laterally by dentated margins, and curving round the borders of the lens. The laminae are arranged into three triangular segments. The central portion, or nucleus. Is unstratified, while the outer layers, or cortex, each contain a nucleus, forming a nuclear layer. The capsule of the lens is a clear, elastic, brittle membrane, inclosing the lens, and held in position by the suspensory liga- ment. Its anterior layer is thicker, and is attached to the lens by a layer of polygonal nucleated cells, which break down post- mortem to form the liquor MorgAgni. The canal of Petit is about one-tenth of an inch in diam- eter, encircliiig the capsule of lens. Its base is formed by the capsule, and it is bounded in front by suspensory ligament, be- hind by the hyaloid membrane of the vitreous humor. The suspensory ligament, or zonula of Zinn, is a thin, trans- parent structure, extending from the margins of the hyaloid fossa, where it is continuous with hyaloid membrane, to the an- terior margin of the lens. The vitebous humor, or body, is a clear, albuminous fluid, filling the cavity of the retina, hollowed out in front — hyaloid fossa — for the lens, and inclosed in the hyaloid membrane. It is composed of water, with a few salts, and a little albumen. Its structure is finely reticular, particularly in the fcetus. The canal of Stilling, canal of Cloquet, or hyaloid canal, is a canal extending in the fcetus from the entrance of the optic nerve to the lens, containing fluid, and lined by a fold of hyaloid membrane. In the fcetus a canal extends parallel but independent of this, and transmits a minute artery to the capsule of the lens. The hyaloid mejribrane is a delicate capsular membrane, investing the vitreous body, excepting its anterior surface, sends fibrous septa into the structure of the vitreous, and is continuous in front with the posterior capsule, and with the anterior capsule through the suspensory ligament. MUSCLES OF THE EYEBALL. Rectus Superior. — Origin, sheath of the optic nerve and upper margin of optic foramen; insertion, into upper surface of sclerotic coat, three or four lines from corneal margin; action, rotates the eyeball upward; nerve, third cranial. 354 HUMAN ANATOMY. Eeotus Infeeioe. — Origin, from lower and inferior part of optic foramen (ligament of Zinn) ; insertion, into lower sur- face of sclerotic; action, rotates the eyeball downward; nerve, third cranial. Eeotus Inteenus. — Origin, same as inferior rectus; inser- tion, into inner surface of sclerotic; action, rotates the eyeball inward ; nerve, third cranial. fP Q,. ■! ' i?S. m — ''Rm. JXi ^•^^^ Fig. 160. The orbital muscles. Lp., levator palpebrse superlorls; Os, supe- rior oblique; Rs, superior rectus; Rl, external rectus; Oi, inferior oblique; Ri, interior rectus; Rm, internal rectus; Tr, trochlea. (Ball.) Eeotus Exteenus. — Origin, by two heads — lower, from ligament of Zinn and lower margin of sphenoidal fissure ; upper, from outer margin of optic foramen; insertion, into outer sur- face of sclerotic; nerve, abducens, or sixth cranial. Passing between the two heads are the ophthalmic vein, the tliird, nasal branch of fifth, and sixth nerves. ORGANS OF SPECIAL SENSE. 855 Sdpeeioe Oblique. — Origin, from inner margin of optic foramen. Its tendon passes through a pulley near the internal angular process of the frontal bone; insertion, into sclerotic between external and superior recti, midway between entrance of optic nerve and the cornea; action, rotates the eyeball on its axis; nerve, fourth, or patheticus. Infeeiob Oblique. — Origin, orbital plate of superior maxilla; insertion, near that of superior oblique, between ex- ternal and superior recti ; action, rotates the eyeball on its axis ; nerve, third cranial. Bupra-orbital tAorimal MuBcuInr and oiliary Anterior meningeal Ophthalmic ^ " Nasal SnpeTlor reotas Iievator palpebrie Bupeiieria Anterior ethmoidal Mnecular trunk Ciliary Arteria centralis Foatericr ethmoidal Optic nerve Internal carotid Fig. 161. Opthalmlc artery and branches. (After MerTcel and KalUus.) (Ball.) The AKTERiES of the globe of the eye are : — 1. The short ciliary, enter through the sclerotic around the optic nerve, to supply the choroid and ciliary processes. 2. The long ciliary arteries, two in number, pierce the sclerotic, run forward between the choroid and sclerotic to the ciliary muscle, which they supply, and where they form an anas- tomotic circle about the iris. 3. The anterior ciliary arieries, five or six in number, enter the sclerotic in front, supply the ciliary processes, and anasto- mose about the iris. 4. The arteria centralis retina, supplies the retina, dividing into four or five branches, which enter as deeply as the inner nuclear layer. 356 HUMAN ANATOMY. The other arteries supplying the eye and its appendages are derived from the ophthalmic branch of the internal carotid (the same source as the ciliary arteries), the anterior cerehral branch of the internal carotid, and the infraorbital branch of the in- ternal maxillary. The branches of the ophthalmic are the Lachrymal ; Supraorbital ; Anterior and posterior ethmoidal; Palpebral; Frontal; Nasal. (For description see Arterial System.) The veins of the eyeball emerge as the vence vorticosce, unite with the other veins to form two main trunks — ^the ophthalmic and inferior ophthalmic veins — ^to terminate in the cavernous sinus. The ophthalmic vein anastomoses freely with the angular vein, the commencement of the facial at the inner angle of the orbit. The nerves of the eye and its appendages are nerve of special sense, the optic or second cranial; motor nerves, the third and fourth; branches of the fifth and sixth, and filaments from the sympathetic. Sensory nerve, ophthalmic division of the trifacial, or fifth cranial {vide Cranial Nerves). Sympathetic branches, derived chiefly from the ciliary gan- glion, Meckel's ganglion, and the cavernous and carotid plexuses {vide Cranial Sympathetic Ganglion). APPENDAGES OF THE EYE. The appendages of the ej'e, or the tutamina oculi, include : — The eyebrows (supercilia), Lachrymal gland. The eyelids (palpebrw). Lachrymal sac, CJonjunetiva, Nasal duct. The eyebrows are the elevated arches of skin surmounting the upper margins of the orbits, covered with short, thick hairs. The eyelids are two movable fokls, covering and protecting the front of the eye. The upper lid is larger, longer, and most movable, being supplied with a special muscle — the levator pal- pebrae superioris. The lids are separated by an elliptical lissure — the fissuia palpehiwum — and connected at their angles of junction bv the Older and inner cantlii. ORGANS OF SPECIAL SENSE. 357 ' On the nasal side the lids are separated by a triangular space — the lacus lachrymalis — at the outer angles of which are the lachrymal papillcB, surmounted by the puncta lachrymalm, the commencements of the lachrymal canal. The lacus lachry- malis is filled by the coruncula lachrymalis, a mass of follicles resembling the Meibomian glands, and covered by the con- junctiva. The structure of the eyelids, from without inward, is as follows : — Skin, subcutaneous areolar tissue, fibres of the orbicularis muscle, tarsal cartilage, fibrous membrane, Meibomean glands, vessels, and nerves; The sMn is thin, and attached by a very loose areolar tissue to the muscle beneath; The fibres of the orbicularis are pale, thin, and act involuntarily; The tarsal cartilages are two plates of firm connective tissue; the superior, the larger, is semilunar in shape, the inferior is elliptical. The inner margins are fixed to the orbit by tendo oculi; The fibrous merribrane of the lids, or tarsal ligament, passes over the anterior surface of the tarsal cartilage, being attached to its free margin below and to the margin of the orbit externally; The Meibomian glands, about thirty in number in the upper, a few less in the lower lid, are arranged vertically on the inner surface of the cartilages; they axe straight, sebaceous follicles, into which open a number of secondary follicles, terminating above in a blunt extremity, and opening below on the free margin of the lids by small foramina, cor- responding to the number of tubules; The eyelashes, or cilia, are arranged on the free border of the lids in two or three rows. The conjunctiva is the mucous lining membrane of the front of the eye. It consists of two portions — the ocular, re- flected over the sclerotic and cornea, and the palpebral portion, lining the internal surface of the lids. The ocular portion is loosely connected with the sclerotic, but over the cornea it becomes very thin, consisting only of the epithelial layer. The palpebral portion is thick, highly vascular, and contains many papillae. At the inner angle of the eye it forms a semi- lunar fold, the plica semilunaris — the rudiment of the nictitating membrane of birds, the membrana nictitans. The point of reflection is •called the fornix conjunctivae, and the reflected portions the superior and inferior palpebral folds. The lachrymal apparatus consists of the lachrymal gland, canals, sac, and nasal duct. The lachrymal gland is an oval, glandular body, about the shape and size of an almond, situated in a depression in the upper surface of the orbit near the external angular process. 358 HUMAN ANATOMY. The anterior portion of the gland is sometimes described as a separate lobe — the palpebral portion of the gland, or ac- cessory gland of Bosenmuller. The gland is attached to the bony roof of the orbit by the tarso-orhital fascia. The ducts, seven to ten in number, open upon the con- junctiva near its point of reflection by minute orifices arranged in a row. The lachrymal canals commence at the puncta lachrymalia, at the summits of the papilla3 lachrymales, and descend by two canaliculi to empty into the lachrymal sac. DUCTS OF LACHRYMAL GLAND VIMAL s»e -NASAL DUCT Fig. 162. The lachrymal apparatus. (.Oerrish, after Testut.) {Eckley.) The superior canal descends obliquely inward and down- ward, while the inferior descends at first and then passes nearly horizontally inward. The lachrymal sac is the oval, dilated^ upper portion of the nasal duct, lodged in a deep groove formed by the nasal process of the superior maxilla, and the lachrymal bone. It is crossed by the tensor tarsi muscle, which acts as a compressor, and receives a fibrous expansion from the tendo oculi. Its structure is made up of a fibrous elastic coat, lined by mucous membrane continuous with the nose and conjunctiva. The nasal duct is a membranous tube about three-quarters of an inch in length, extending from the lachrymal sac to the inferior meatus of the nose, and lining the bony lachrymo-nasal ORGANS OF SPECIAL SENSE. 359 canal. It passes backward, downward, and outward, and is pro- tected at its inferior extremity by a valve of mucoTis membrane — the valve of Hasner. The mucous lining is continuous with that of the sac, but instead of having squamous epithelium it is ciliated in the duct. THE EAE. The eab — the organ of hearing — is contained in the petrous and mastoid portions of the temporal bone, and consists of three divisions, — external ear, middle ear or tympanum, and internal ear or labyrinth: — ■ , _, . , f Auricle, or pinna, 1. J!.-xternal ear, ^ External auditory canal or meatus; 2. Middle ear or tympanum. Membrana tympani. Tympanic cavity. Mastoid sinuses, , Eustachian tube; 3. Internal ear or labyrinth. {Vestibule, Semicircular canals, Cochlea; (Utricle, Semkircular canals. Cochlea. The external ear includes two parts, the auricle and the external auditory canal. AUEIOLB, OR PINNA. The external prominent portion is composed of cartilaginous segments, comieeted together by ligaments and muscles richly supplied with blood-vessels, nerves, and lymphatics, and covered with skin. It presents the following elevations and depres- sions : — Fossa of the helix, a depression between the helix and anti- helix ; Fossa of the antihelix, a depression above the bifurcation of the antihelix; Tragus, the prominence in front of the concha, usually cov- ered with hair; Antitragus, a small, conical eminence behind the tragus, from which it is separated by the incisura intertragica ; Lobule, the soft, rounded portion depending below the anti- tragus ; 360 HUMAN ANATOMY. Concha, a deep cavity, surrounded by the helix, and leading into the exteraal meatus. Muscles. — The muscles of the auricle include two sets, the extrinsic muscles, already described {vide Muscles), of which there are three — the attolens aurem, attrahens aurem, and re- trahens aurem — and the intrinsic muscles, but slightly devel- oped, of which there are two sets, four on the anterior surface of the auricle — the tragicus, antitragicus, helicis major, and Fig. 163. External, middle, and internal ear. 1, external ear; 2, middle earj S, internal ear; 4, pinna; 6, helix; 6, antihelix; 7, fossa navicularis; 8, fossa innominata; 9, tragus; 10, antitragus; 11, concha; 12, lobe; 13, meatus auditorius externus; 14, tympanic membrane; 16, promon- tory; 16, foramen rotundum; 17, posterior wall of the tympanum; 18, ossicula auditus; 19, Eustachian tube; 20, narrow canal; 21, vestibule; 22, semicircular canals, the superior, posterior, and horizontal; 23, ampullae; 24, cochlea; 25, prominence caused by the scala vestibuU, 26, scala tympani. (Boennmg.) helicis minor — and two on the posterior surface, transversus auricula and obliquus auriculae. Arteries, auricular branch, from the occipital; posterior auricular, from the external carotid; and anterior auricular, from the temporal. The veins correspond to the arteries. Nerves, auricular branch, from the pneumogastric ; auriculo- temporal branch, from the inferior maxillary; occipitalis major ORGANS OF SPECIAL SENSE. 361 and occipitalie minor; auricularis magnus, from the cervical plexus ; and posterior auricular, from the facial. Auditory canal, or meatus auditorius externus, is an osseo- cartilaginous canal, about one and one-fourth inches in length, extending from the concha to the tympanic membrane. The osseous portion forms about two-thirds of the passage, and consists in greater part (anterior and lower) of a curved plate of bone — the annulus tympanicus. The anterior and upper part of the ring present two spines — spina tympanica, major and minor, for the ligaments of the malleus. The sulcus iympanicus is a furrow on the inner edge of the ring, for the attachment of the tympanic membrane. The cartilaginous portion forms about one-third of the pas- sage, and consists of the inverted cartilage of the tragus and concha, the upper and back part of which is deficient, the cleft being filled with fibrous tissue. The canal, including the tympanic membrane, is lined throughout with skin, containing short hairs, vascular papillae, sebaceous and ceruminoxis glands. Arteries — branches of the internal maxillary, posterior auricular, and temporal branches of external carotid. Nerves — from the auriculo-temporal branch, the inferior maxillary, and auricular branch of the pneumogastric. MIDDLE EAR, OR TYMPANUM, is an irregular cavity, situated between the auditory canal and the labyrinth, communicating with the pharynx through the Eustachian tube, and also with the mastoid cells. It contains a chain of movable bones, part of the chorda tympani nerve, and is filled with air. Its average diameters are about half an inch in height and width, and a line or two in depth from without inward. It is lined with mucous membrane, continuous with that of the Eustachian tube and mastoid cells, and which is reflected over all the tympanic contents. It is bounded by a roof, floor, and four walls, which present the following points : — Boof, a very thin plate of bone, separates the tympanum from the cranial cavity, and corresponds to a depression on the anterior wall of the petrous bone ; Floor, forms the bottom of the jugular fossa, and pre- sents : — The opening for Jaco'bson's nerve, the tympanic branch of the glosso-pharyngeaL 362 HUMAN ANATOMY. Outer wall, formed by the tympanic membrane, Shrapnell's membrane, and the annulus tympanicus surrounding it : — 'Notch of Bivinus, incomplete upper ring of bone ; Iter chordw posterius, gives entrance to chorda tympani nerve; Iter chorda) anterius, or canal of Hugier, givea exit to same ; Olaserian fissure, above and in front of annulus tympanicus, re- ceives the anterior ligament of the malleus, the long process of the malleus, and the tjTUpanic arterj' ; Pouches of the memtrana tympani, are two or three pockets formed by the doubling of the mucous membrane around the chorda tympani nerve. Inner wall, separates the tympanum from the labyrinth : — Fenestra ovalis, an oval window, leading to the vestibule, closed by the membrane to which the base of the stapes is attached; Fenestra rotunda, a round window, leading to the scala tympani, but closed also by a membrane — membrana tympani secundaria. Promontory, two grooves, marking the first turn of the codilea, and separating the fenestrse ovalis and rotunda; Prominentia canalis facialis, a small conical eminence above the fenestra ovalis, which contains within it the aquaeductus Fallopii, the long canal for the facial nerve. Posterior wall: — Openings of the mastoid cells, three or four in number, connect these sinuses with the tympanum. Anterior wall, is a thin plate of bone, which separates the tympanum from the carotid canal. Openings of the canal for tensor tympani, lies above and parallel with the canal for the Eustachian tube, and transmits the tensor tym- pani muscle; Opening of Eustachian tube, the larger of the two, is separated from the former by a thin plate of bone, the processus cochleariformis. The tympanum is divided into two parts — the attic of the tympanum and the atrium. The former is situated on a plane directly above the atrium, and is bounded above by the tegmen, externally by auditory plate, and internally by prominence of the tympanum. On the outer part of the atitic is a smooth surface called the sciite (Leidy). The ErsTACHiAN tube is an nssco-cartilaginous canal, one and one-half to two inches in length, extending from the tym- panum to the pharynx. It is about one-third osseous and two-thirds cartilaginous, and lined with ciliated epithelium continuous with the pharvnx ORGANS OF SPECIAL SENSE. 363 and tympanum. Its pharyngeal orifice is trumpet-shaped, and opens at the side of the pharynx, behind the inferior meatus. Muscles of the Eustachian tube: — Tensor palati (tensor veli), the most important has been described (vide Palatal Eegion). Its action is to pull the ante- rior wall of the tube outward and downward, and widen the canal. Internal Pterygoid {vide Muscles). — Some fibres are in- serted into the floor of the tube. It acts as tensor of the fascia. Ligamenta salpingo-pharyngeal, arises from the superior and middle constrictors of the pharynx, and is inserted by three to five tendinous cords into the pharyngeal orifice of the tube. Action, opens the tube. Arteries of Eustachian Tvhe. — Pharyngeal from the ex- ternal carotid, middle meningeal from the internal maxillary, and small branches from the internal carotid. Nerves, by branches from the glosso-pharyngeal, inferior maxillary, otic ganglion, and facial nerve. Membrana tympani is a thin, parchment-like membrane, stretched upon the annulus tympanicus, forming the bottom of the external auditory canal, and separating it from the tym- panum. It is placed obliquely downward and inward at an angle of 45°. Structure. — Its three layers are : — External or cuticular, derived from the skin of the meatus ; Middle or fibrous, consisting of two sets of fibres, radiating and circular, the latter forming a tendinous ring around the margin ; Internal or mucous, continuous with the mucous membrane of the tympanum. Its outer surface presents : — Manubrium of malleus, projecting downward and backward ; Processus gracilis of malleus, projecting outward above the manubrium ; Umbo, a depression of the membrane formed by the lower end of the manubrium ; Yellow spot, the cartilaginous end of the manubrium ; pyra- mid of light, a triangular cone of reflected light in the antero- inferior quadrant of the membrane. Inner surface (vide Outer Wall of Tympanum). Arteries, tympanic branch of iaferior, branch from the in- ternal carotid. Nerves, from the superficial temporal branch of the trifacial, and from the tympanic plexus. 364 HUMAN ANATOMY. OSSICLES OF THE TYMPANUM. The small bones of the tympamim are three in number, and connect the membrana tympani with the membrane closing the fenestra ovalis. They are the 1. Malleus, or hammer, consists of a head, neck, manubrium or handle, processus gracilis, and processus brevis. The manu- brium is attached to the membrana tympani, and has the tendon of the tensor tympani attached. Processus gracilis is lodged in the Glaserian fissure. A "^'^'"miim^^. Fig. 164. The auditory ossicles. A. G., external meatus; M, membrana tym- pani; n, handle of the malleus and its short process (p); h, head of the malleus; a, incus; K, its short process, with its ligament; 1, long process; s, os orbiculare; S, stapes; Ax, Ax, the axis of rotation of the ossicles, shown in perspective; t, line of traction of the tensor tympani. The other arrows show the movements of the ossicles when the tensor contracts. (Boenning.) 2. Incus, or anvil, consists of a body and long and short processes. The body articulates with the malleus and the long process by means of a rounded process. The os orbiculare ar- ticulates with the head of the stapes. 3. Stapes, or stirrup, has a head, neck, two branches (or crura) which unite into a cross-piece or base. The neck has in- serted into it the stapedius muscle, and the base is fitted to the fenestra ovalis. ORGANS OF SPECIAL SENSE. 365 Ligaments of the Ossicles: — Lig. mallei anterius, extends from the neck of the malleus to the outer wall of the tympanum; Lig. mallei suspensorium, suspends the head of the malleus to the roof; Lig. incudi posticum, attaches the short process to the posterior wall ; Lig. incudi suspensorium, suspends the incus to the roof; Lig. stapedi annularum, connects the base to the margin of the fenestra ovalis. Muscles. — The tensor tympani, stapedius. Tensor tympani (already described). Stapedius arises from the hollow of the pyramid on the inner wall, and its tendon is inserted into the neck of the stapes. Nerve from the tympanic branch of facial. Arteries : — Tympanic branch of internal maxillary; Stylo-mastoid branch of posterior auricular; Petrosal branch of middle meningeal; Tympanic branch from internal carotid; Branch from ascending pharjTigeal. Veins accompany the corresponding arteries and empty into middle meningeal and pharyngeal. Nerves: — Chorda tympani, from the facial, enters the iter chordae posterius, crosses the tympanum between the long process of the incus and the handle of malleus, and makes its exit at the iter chordfE anterius; Tympanic branch of glosso-pharyngeal (Jacobson's nerve) enters the floor, supplies the fenestrse and mucous membrane of tympanum and Eustachian tube; Tympanic branch from facial to the stapedius ; Branch from the otic ganglion to the tensor tympani. The tympanic plexus is formed upon the surface of the promontory, from the following nerves : — Jacobson's nerve, tympanic branch of the glosso-pharyngeal j Branch of the superficial petrosal, from the facial ; Branches from the carotid plexus of the sympathetic INTERNAL EAE, OK LABTEINTH. The internal ear, the essential part of the organ of hearing, consists of three complex cavities within the petrous portion of the temporal bone, filled with fluid, the perilymph, and contain- 366 HUMAN ANATOMY. ing a membranous sac filled also with fluid, the endolymph, which receives the distributions of the auditory nerve. Its divisions are : — Vestibule, Osseous portion, -i Semicircular canals. tion, -i Cochlea J I Utricle, Semicircular canals. Cochlea. Vestibule, is a common cavity of communication between the bony parts of the internal ear. It is situated betvreen the tympanum without, the cochlea behind, and the semicircular canals in front, and measures one-fifth of an inch in its antero- posterior and vertical diameters, and less from without inward. It presents : — Fenestra ovalis on its outer wall, closed by the stapes and its ligament; Fovea hemisph erica, a circular depression on its inner wall for the saccule, and perforated by the macula cribrosa, for the vestibular filaments of the auditory nerve; Orifice of the aquceductus vestibulce, on the inner wall for the transmission of a small vein; Eminentia pyramidalis, a vertical ridge on the inner wall separating the two foveas. Fovea semi-elliptica, in the roof lodges the utricle. Semioieoulae canals are three canals, forming each two- thirds of a circle one-twentieth of an inch in diameter, and named, from their position, the superior, posterior, and external. They are placed nearly at right angles to each other, and open into the vestibule by five apertures — two extremities uniting to form one. The superior and posterior are both vertical, the former being more anterior. The external is placed horizontally, its arch directed back- ward. AmpuUcB, are the dilated, flask-shaped extremities of the tubes, and are about one-tenth of an inch in diameter. The cochlea resembles closely a common snail's shell, placed with the base corresponding to the bottom of the meatus auditorius internus, and its apex directed outward and forward. It consists of two parallel tubes one and one-half inches in length, one-tenth of an inch in diameter, wound spirally for two and one- half turns around a central pillar — the modiolus. UKUANS OF SPECIAL SENSE. 367 The modiolus, or columella, is broad at the base, but tapers gradually toward the apex, where it terminates within the last turn or cupola in the infundibulum, a funnel-shaped lamina of bone. Its interior is traversed by numerous canals for vessels and nerves, one of which — the canalis centralis modioli — ^nins nearly the whole length, and transmits the arteria centralis modioli. Lamina spiralis, the partition between the two tubes, con- sists of two thin laminse of bone filled with cancellous tissue — Fig. 165. Organ of Cortl. N, cochlear nerve; K, inner and, P, outer halr- ceUs; n, nerve-flbrils terminating in F; a, a, supporting cells; d, cells In sulcus-spiralis; z, inner rod of Cortl; Mb. Corti, membrane of Corti. or the membrana tectoria; o, the membrana reticularis; H, Q, cells filling up the space near the outer wall. (Boenning.) lamina spiralis ossea — ^projecting from the modiolus half-way to the outer wall, the lamina being completed by the membrana basilaris. The osseous lamina terminates in the cupola in a hook-like process — ^the hamulus. The spiral canal, between the modiolus and the outer wall, is divided by the lamina spiralis ossea and membrana basilaris into two canals, or scales — ^the upper scala vestibuli, and the lower Bcala tympani. 368 HUMAN ANATOMY. The first turn of the spiral canal presents three openings : — 1. Fenestra rotunda, a circular aperture communicating with the tympanum, and closed by the membrana tympanum secundaria. 3. Fenestra ovalis, an oval opening, commimicating with the vestibule. 3. Aquceductus cochlearis, a small foramen openipg on the inferior surface of the petrous portion of the temporal bone, and transmitting a small vein from the cochlea to the jugular vein. Scala vestibuli, begins at the fenestra ovalis, and ascends in a spiral course above the lamina spiralis and membrana basilaris, to communicate vrith the scala tympani, through the helicotrema within the vestibule. It contains perilymph. Scala tympani begins at the fenestra rotunda, and ascends in a spiral course below the scala vestibuli, from which it is separated by the osseo-membranous lamina. It also contains perilymph. Helicotrema is an opening within the cupola, by which the two scalse communicate. It is formed by a deficiency of the osseous lamina. Canalis spiralis modioli, is a small canal which winds around the modiolus at the attachment of the osseous laminse. It con- tains a gangliform swelling — ganglion spirale — ^from which nerves pass to the organ of Corti. The mbmbkanous labyrinth corresponds closely to the osseous, the vestibule consisting of two membranous sacs — ^the utricle and saccule — communicating with each other, and the former receiving the five openings of the membranous semicir- cular canals, the latter communicating with the membranous cochlea through the canalis reuniens. The membranous labyrinth is surrounded everywhere with perilymph, filled with endolymph, and is composed of three coats — an outer fibrous layer, middle or tunica propria, and inner epithelial layer. Utricle, the larger of the two, is an elliptical sac lodged in the fovea hemi-elliptica. The semicircular canals communicate with it by five openings. Its wall is the thickest near the crista vestibuli, where the filaments of the nerve enter, the macula acoustica. Saccule, is a hemispherical sac lodged in the fovea hemi- spherica. It communicates with the cochlea through a small duct — the canalis reuniens. The otoliths, or otoconia, are small masses of crystals of lime carbonate, contained in the inner wall of the utricle and saccule opposite the distribution of the nerves. ORGANS OF SPECIAL SENSE. 369 Semicircular Canals. — These are three membranous tubes corresponding exactly in shape, number, and form to the bony canals in which they are contained. They are surrounded every- where with perilymph, except at the ampullae, where they are in contact with the bony canals. The inner, or epithelial "/ayer of- the membrane in the ampullae, is covered with columnar ciliated epithelium — auditory hairs. The membeanous cochlea begins at the base and ascends in a spiral course within the osseous cochlea to terminate in the cupola. It includes only the canalis cochlea — a part of the scala vestibulae. As before stated, the membrana basilaris extends from the margin of the lamina spiralis ossea to the outer wall of the cochlea, dividing the cavity into the two sealae — ^the scalae vestibulse above, the seals tympani below. The former is again subdivided by the membrane of Eiessner into two parts, the outer of which forms the canalis cocMece, or scala media — the mem- branous cochlea proper — on the floor of which is the organ of Corti covered by the membrana tectoria. The scalse are lined with periostium, and filled with perilymph. The scala tym- pani ends at the fenestra rotunda, but the scala vestibula com- municates freely with the vestibula. Limhus laminw spiralis is the periosteal margin of the lamina spiralis ossea, and consists of an upper lip — -the labium vestibulare — and a lower lip — ^the labium tympanicum — sepa- rated by a groove — the sulcus spiralis. Membrana basilaris extends from the labium tympanicum to the outer cochlear wall, to which it is attached by the liga- mentum spirale of Henle. Membrane of Reissner arises from the middle of the vestibu- lar lamina and passes obliquely at an angle of 40° to the outer cochlear wall, separating the canalis cochleae from the scala vestibula. Canalis cochlem extends as a spiral sac closed at both ends through the osseous cochlea, and contains the most important part of the labyrinth — the ultimate distribution of the auditory nerve in the organ of Corti. It is triangular on section, the membrana basilaris forming the base, the membrane of Eiessner the inner side, and the periosteum of the cochlea its outer side. Membrana tectoria, or membrane of Corti, commences as a delicate membrane, at a point between the origin of the mem- brane of Eiessner and the labium vestibule, and arches over to the -outer cochlear wall, inclosing the organ of Corti, but not in contact with it. Organ of Coeti consists of two sets of pillars — ^the inner and outer rods of Corti — extending upward upon the vestibular 370 HUMAN ANATOMY, surface of the membrana basilaris, and forming a series of arches. Between the bases of the arches is the zona arcuata. There are over three thousand arches, and within them is formed a tunnel extending the entire length of the lamina spiralis termi- nating at the hamular process. Inner rods of Gorti, rest upon the basilar membrane. Their upper extremities resemble the proximal extremity of the ulna, and each has on its inner side a row of epithelial cells — the inner hair-cells. Outer rods of Corti, are attached in a similar manner at the bases, and their extremities each resemble the head and bill of a swan, the former fitting into the cavity in the inner rods. On the outer side are four rows of ciliated cells — ^the outer hair- cells. Reticular memhrane, or lamina reticularis, is a delicate complex net-work of phalanges — fiddle-shaped structures — ex- tending from the inner rods to the external row of the outer hair-cells. The internal auditoet canal, is a short canal about one- third of an inch in length, extending outward from the meatus auditorius internus, on the posterior surface of the petrous por- tion of the temporal bone to terminate at a vertical, perforated plate — the lamina cribrosa. This plate is divided by a horizontal ridge into a lower and upper portion. The lower is perforated by numerous foramina for passage of the auditory artery and branches of the auditory nerve, some of which are arranged into a spiral-shaped depression, called the tractus spiralis foraminu- lentus; the upper has one large opening — ^the commencement of the aquaeductus Fallopii for portio dura, or seventh nerve. Auditory Nerve. — The auditory nerve enters the internal auditory meatus in company with the facial, and at the bottom divides into two branches — the vestibular and cochlear. Vestibular nerve subdivides into three branches, as fol- lows : — Superior, distributed to the utricle and ampullse of the superior and external semicirculax canals; Middle, enters the bottom of the fovea hemispherica, and is dis- tributed to the saccule; and the Inferior, is distributed to the ampulla of the posterior semicircular canal. Cochlear nerve, ascends in the canals in the modiolus, and divides into numerous branches which pass between the plates of the lamina spiralis ossea, forming a plexus which contains the ganglia spirale, from which filaments are distributed to the outer and inner hair-cells of the organ of CortL ORGANS OF SPECIAL SENSE. 371 Arteries of Labyrinth: — Internal auditory, from the basilar, divides into vestibular and cochlear branches, which accompany the corresponding nerves ; Stylo-mastoid, from the posterior auricular, enters through the stylo-mastoid foramen. THE TONGUE.* The tongue is the organ of the sense of taste, and lies in the floor of the mouth. It is composed of a mass of muscles, the free surface of which is entirely invested with mucous mem- brane. It consists of three parts : — Base, or root, attached to the hyoid bone, the epiglottis and the soft palate ; Body, the back or dorsum of which is marked by a median line or raphe, terminating behind in a cavity — ^the foramen cwcum; Apex, or tip, narrow and pointed, directed forward. The mucous membrane resembles skin, consisting of a corium or mucosa, containing numerous papilla, and covered with epithelium. The papillae are most numerous over the ante- rior two-thirds of the dorsum, and consist of three varieties : — Circumvallate, or largest, about ten in number, about one- twelfth inch wide, are arranged at the back part of the dorsum like the letter V, with the apex directed backward ; Fungiform, or medium, are scattered over the organ, espe- cially at the tips and sides ; Filiform, conical, or smallest, are distributed over the an- terior two-thirds of the dorsum. Mucous and serous glands and simple papillae, such as are found in the skin, are also present. Taste bulbs, or taste goblets, minute, flask-shaped bodies, about one three-hundredths of an inch in length, are situated in the circumvallate and fungiform papillae. Arteries, are branches of the lingual, facial, and ascending pharyngeal. 1. Nerves. — Hypoglossal, the principal motor nerve. 2. Lingual branch of glosso-pharyngeal, the special nerve of taste. 3. Lingual branch of the trifacial, to sides and anterior part; nerve of common sensation. 4. Chorda tympani, probably nerve of special sense. * Tide Lingual Muscles, page 119. 372 HUMAN ANATOMY. 5. Superior laryngeal, filaments from its internal branch to the base. THE SKIN AND APPENDAGES. The skin, besides being the special organ of the sense of touch, is an absorbing and excretory organ, and protects the underlying structures. It consists of two layers — ^the epidermis or cuticle, and the derma or true skin. The appendages are the nails, hair, sudoriferous and sebaceous glands, and their ducts. The epidermis, or cuticle, is an epithelial structure consist- ing of four layers : — 1. Rete Malpighii, the deepest layer, in contact with the corium, is composed of several layers of round or polyhedral epithelial cells. This layer is pigmented in the negro and other dark races. 2. Granular layer, is a single layer of compressed, spindle-shaped cells. 3. Stratum lucidum, are still more compressed into scales. 4. Stratum corneum, several layers of horny scales. The derma, or true skin, has two layers, the reticular or deeper, and the papillary or superficial. Eeticular layer is composed of interlacing bands of white fibrous tissue, mixed with yellow elastic fibres, and some un- striated muscular fibres where hairs are found, together with blood-vessels, lymphatics, and nerves. Papillary layer is covered with minute, vascular, highly sen- sitive elevations — ^the papillae — from one one-hundredth to one two-hundred-and-fiftieth of an inch in diameter. These papillae are in places arranged into parallel curved ridges, and have in the most sensitive situations some one of the "peripheral end organs" (already described) added to their structure. Nails are horny plaques of modified epidermis, firmly adapted to the derma on the dorsal surfaces of the terminal phalanges. Each nail is firmly implanted by its root into a fold of skin. The matrix is the derma beneath the body, which is vascular and highly sensitive. The lunula is the white, less vascular portion of the matrix near the root. Haies are a cylindrical modification of the epidermis, and consist of a root imbedded in the skin, a shaft, and a point. Boot is lodged in a pouch-like involution of epidermis — the hair-follicle. This is composed of an inner or cuticular and an ORGANS OF SPECIAL SENSE. 373 outer or dermic layer, and presents at the bottom a vascular papilla, on which the hair rests, and from which it grows. Shaft, consists of a pith or medulla in the centre, sur- rounded by a fibrous part containing pigment, and covered with a layer of scaly epithelium. Point, contains the two latter elements, but has no medulla. Sebaceous glands are minute, glandular bodies, situated in the substance of the corium, and each opening by a single duct into a hair-follicle, or upon the surface of the skin. They are most abundant where hairs are found. The Meibomian glands in the eyelids are the largest. SuDOEiFERODS or SWEAT GLANDS are minute, reddish, glandular bodies, situated usually in the subcutaneous connective tissue, each consisting of a single convoluted tube. Each gland opens on the surface of the skia. SURGICAL ANATOMY. THE TRIANGLES OF THE NECK. The general outline of the neck is somewhat quadrilateral. Bounded in front by the median line, behind by the trapezius muscle, above by the lower border of the body of the maxilla, and below by the clavicle. It is divided by the sterno-mastoid muscle running obliquely through it into two large triangles, an ante- rior and a posterior triangle. These are again subdivided by the crossing of the omo-hyoid muscle into two smaller triangles each. The anterior into the superior and inferior carotid tri- angles, and the posterior into the occipital and subclavian tri- angles. The digastric muscle, at the upper part of the anterior triangle, separates a triangular space called the submaxillary triangle. , The anterior triangle of the neck is hounded as follows: — Front, median line of the neck ; Behind, anterior border of the sterno-mastoid muscle; Above, lower border of the body of the lower jaw, and a line from its angle to the mastoid process, forming the base of the triangle ; Below, the top of the sternum, forming the apex. The floor is formed by the sterno-hyoideus, sterno-thy- roideus, thyro-hyoideus, inferior and middle constrictors of the pharynx, the anterior belly of the digastrieus, the mylo-hyoideus, stylo-hyoideus, and hyoglossus muscles. The roof is formed by the skin, superficial fascia, platysma myoides, and deep fascia. The inferior carotid triangle is the most inferior subdivision of the anterior triangle, and is hounded as follows : — Front, median line of the neck; Behind, anterior border of the sterno-mastoideus ; Above, anterior belly of the omo-hyoideus ; Below, the apex of the anterior triangle. It contains the following structures : — Thyroid gland and lower part of the trachea and larynx ; Inferior thyroid and common carotid arteries ; Inferior thyroid and internal jugular veins; Pneumogastric, recurrent laryngeal, descendens noni, com- municans noni, and sympathetic nerves. (374) SURGICAL ANATOMY. 375 The superior carotid triangle is the middle subdivision of the anterior triangle of the neck, and has the following bound- aries : — Above, posterior belly of the digastricus ; Below, anterior belly of the omo-hyoideus; Behind, anterior border of the stemo-mastoideus. It contains the following structures : — Termination of the common carotid; Pig. 168. Triangles of the neck. (Boenning.') The internal carotid; The external carotid, and the following branches : — Superior thyroid, lingual, facial, ascending pharyngeal, and occipital ; The internal jugular vein, and the following tributaries : — Lingual, facial, superior thyroid, pharyngeal, and occa- sionally the occipital ; 376 HUMAN ANATOMY. The pneumogastric, superior laryngeal, external laryngeal, descendens noni, spinal accessory, hypoglossal, and sympathetic nerves. The submaxillary triangle is the most superior of the sub- divisions of the anterior triangle, and has the following bound- aries : — Above, the lower border of the body of the jaw, and the imaginary line behind ; In front, the median line of the neck ; Behind, the posterior belly of the digastricus. It contains the following structures : — External carotid, internal carotid, facial, submental, mylo- hyoidean arteries; The internal jugular, the commencement of the external jugular, branches of the anterior jugular, and the facial vein and its branches ; . The inframaxillary branches of the facial nerve, the ascend- ing branches of the superficial cervical nerve, and the pneumo- gastric and glosso-pharyngeal and mylo-hyoid nerves; Portions of the parotid and submaxillary glands, and also submaxillary lymphatic glands and vessels. The posterior triangle of the neck occupies the space behind the posterior border of the sterno-mastoideus, and is hounded as follows: — Front, posterior border of the sterno-mastoideus; Behind, anterior border of the trapezius; Below, upper border of the middle third of the clavicle, forming its base; Above, the occiput, forming the apex. Its floor is formed from above downward, by the following muscles : — Splenius capitis, levator anguli scapuli, scalenius medius, scalenius posticus, and the upper portion of the serratus magnus. Its roof is formed by the superficial and deep fascia, and below by the platysma myoides. It contains the following structures : — The trans versalis colli artery and vein; Spinal accessory and superficial plexus of the cervical nerve; Lymphatic glands and vessels. The subclavian triangle, or the inferior division of the poste- rior triangle, is hounded — In front by the margin of the sterno-mastoideus, behind by the posterior belly of the omo-hyoideus, below by the clavicle. It contains the following structures : — The subclavian, transversalis colli, and transversalis humeri or suprascapular arteries; SURGICAL ANATOMY. 377 The transversalis colli, suprascapular, and external jugular veins ; The descending branches of the superficial cervical plexus, and the brachial plexus of nerves; Lymphatic neives and vessels. Triangle in Front of the Elbow-joint. Bounded — Externally, by supinator longus; Internally, by pronator radii teres; Base, above, by a line (imaginary) down through the con- dyles ; Apex, below, by crossing of supinator longus and pronator radii teres. It is covered in by skin, superficial fascia, and bicipital fascia; the floor is formed by oblique fibres of the supinator brevis and lower part of brachialis anticus muscles. It contains: — From within outward, median nerve, brachial artery, and venae comites, biceps tendon, and museulo-spiral nerve. Scarpa's Triangle is a large triangular space situated in the upper part of the anterior surface of the thigh, through which the femoral vessels descend. It is hounded — Externally by sartorius, internally by adductor longus, above by Poupart's ligament; below, apex is formed by crossing of boimding muscles. It is covered in by skin, superficial fascia lata, and cribri- form fascia, and its floor is formed by the following from with- out inward: iliacus, psoas, pectineus, and portion of adductor brevis muscles. It contains: — The femoral artery (with its profunda and cutaneous branches), inclosed in the femoral sheath, femoral vein (Joined by profunda and long saphenous veins), anterior crural nerve and its branches [from within outward being vein, artery, and nerve], deep lymphatic glands and vessels, and adipose tissue. Axilla. This is a pyramidal space between the upper and lateral part of the chest and the inner side of the arm. It is bounded — In front, by the pectoralis major and minor muscles; be- hind, by latissimus dorsi, teres major, and subscapularis; in- 378 HUMAN ANATOMY. temally, by the upper four ribs and intercostal and upper part of serratus magnus muscles, and its apex is directed upward, and is formed by the space between the first rib, clavicle, and upper border of the scapula, and its base by the skin and axillary fascia stretched across between the lower borders of the pectoralis major and latissimus dorsi muscles. no. 167. The axilla. 1, axillary artery; 2, brachial artery; S, acromlo- thoracic artery; 4, superior thoracic artery; 5, subscapular artery; 8, dorsalis scapulae artery; 7, posterior circumflex artery; 8, superior pro- funda artery; 9, posterior thoracic nerve; 10, long subscapular nerve; 11, median nerve; 12, cephalic vein; 13, musculo-cutaneous nerve; 14, teres major muscle. {Boenning.) It contains: — The axillary artery and vein, and their branches; the brachial plexus of nerves, and the branches given off below the SURGICAL ANATOMY. 379 clavicle; branches of the intercostal nerves; ten or twelve lym- phatic glands, and a quantity of loose adipose and areolar tissue. Popliteal Space. The popliteal space is a lozenge-shaped space situated at the back of the knee, and forms the ham. It is hounded — Externally, above the joint, by the biceps; below the joint, by outer head of gastrocnemius and plantaris; internally, above the joint, semitendinosis, semimembranosis, gracilis, and sar- torius; below the joint, by inner head of the gastrocnemius. It is covered in by the skin, superficial fascia, and fascia lata. Its floor is forined by the lower part of the posterior surface of the femur, the ligamehtum posticum Winslowi, and the pop- liteus muscle and its fascia. It contains: — The popliteal artery and branches ; popliteal vein receiving the external saphenous vein (vein to the outer side) ; internal and external popliteal nerves and branches ; articular branch of obturator nerve; branch of small sciatic nerve; four or six lymphatic glands, and a quantity of adipose and areolar tissue. ANATOMY OF HERNIA. Hernia is the protrusion of any' part of a viscus from its natural cavity through the enclosing walls of the cavity. As usually understood, it refers to the protrusion of the intestine or mesentery, or both, from the abdominal cavity. The most com- mon forms are: — External or oblique inguinal hernia; Internal or direct inguinal hernia; Femoral hernia; Umbilical hernia. 1. Oblique inguinal hernia escapes from abdominal cav- ity at the internal abdominal ring, carrying before it a pouch of peritoneum, descends along the inguinal canal in front of the cord, and emerges at the external opening. The anatomical parts concerned are the inguinal canal, with its internal and external abdominal rings, the transversalis fascia {vide Fascia), the peritoneum (vide Peritoneum), and Poupart's ligament. The inguinal or spermatic canal is about one and a half inches in length, extending from the internal abdominal ring to the external abdominal ring. It serves for the passage of the 380 HU.MAN ANATOMY. spermatic cord in the male, and the round ligament with its vessels in the female. Its boundaries are: — In front, skin, superficial fascia, the aponeurosis of the external oblique muscle, the lower border of the internal oblique, and a small portion of the cremaster muscle; Fig. IGS. Dissection ot the inguinal canal: A, external oblique; B B, in- ternal oblique; C, transversalis; D, conjoined tendon; B, rectus ab- dominis witli sheath opened; F, fascia transversalis; H, cremaster; I, infundibular fascia. Behind, the fascia transvei'salis, tlic conjoined tendon of the transversalis and internal oblique muscles, the transversalis fascia, and subperitoneal fat and peritoneum ; Above, by the arched Ijorder of the internal oblique and transversalis muscles ; Below, l)y the connection between Poupart's ligament and the transversalis fascia. SURGICAL ANATOMY. 381 The internal abdominal ring is an oval opening in the transversalis fascia about half an inch above Poupart's ligament, midway between the symphysis pubis and the anterior superior spinous process of the ilium. It is bounded — Above and externally by the arched fibres of the transversalis muscle; below and internally by the deep epigastric vessels. It transmits a funnel-shaped fascia from its margins, the infundibuliform fascia. The external abdominal ring is a triangular opening between the two pillars in the aponeurosis of the external oblique muscle, to the outer side and just above the spine of the pubes. It is about half an inch wide and one inch long, and is bounded — Above, by the intercolumnar fascia ; below, by the spine and crest of the os pubis, and on either side by the pillars or columns of the ring formed by the free margins of the aponeurosis of the external oblique. From the margins of the external abdominal ring arises the intercolumnar fascia. Poupart's ligament, or the femoral arcli, is the rounded lower fibrous margin of the external oblique muscle, extending between the anterior superior spine of the ilium and the spine of the pubes. The portion of the aponeurosis which is inserted into the pectineal line. has received the name of Gimbernat's ligament. The triangular ligament or ligament of Colles is the reflected inner portions of Gimbernat's and Poupart's ligaments attached to the sheath of the rectus. The epigastric artery holds a very important anatomical relation to the inguinal canal and internal abdominal ring. It ascends between the peritoneum and transversalis fascia to reach the sheath of the rectus muscle along the inner and lower margin of the internal ring and beneath the spermatic cord. The coverings of oblique inguinal hernia are : — • (1) Skin; (2) superficial fascia ; (3) intercolumnar fascia ; (4) cremaster muscle, and fascia; (5) fascia transversalis, or infundibuliform fascia; (6) subserous connective tissue; (7) peritoneal sac. 2. Direct inguinal hernia escapes through the abdominal wall, within Hesselbach's triangle, internal to the epigastric artery, and through the external abdominal ring, pushing before it a pouch of peritoneum. Hesselbach's triangle is a space situated at the lower ante- rior part of the abdominal wall on either side. It is bounded as follows : — Externally, epigastric artery ; 382 HTJMAN ANATOMY. Internally, the outer margin of the rectus abdominis muscle ; Below, Pouparfs ligament, forming its base. The struc- tures forming the abdominal wall at this cavity are, from with- out inward : — 1. Skin; 2. Superficial fascia; 3. Intercolumnar fascia ; 4. Conjoined tendon of the transversalis and internal ob- lique ; 5. Fascia transversalis; 6. Subserous tissue; 7. Peritoneum. The anatomical parts concerned in direct inguinal hernia are the same as those in the indirect, already given. The cover- ings of direct inguinal hernia are those just given. 3. Pbmoeal hernia escapes through the femoral ring, carrying before it a pouch of peritoneum, descends through the femoral canal and emerges through the saphenous opening in the fascia lata. The anatomical parts concerned in this hernia are: — The femoral ring, the femoral or crural canal, the septum crurale, crural sheath and deep crural arch, and the saphenous opening. The femoral or crural ring is an oval opening, about an inch and a half in diameter, larger in the female and situated below Pouparfs ligament, between the inner side of the femoral vein and the margin of Gimbernat's ligament. It is the abdom- inal opening of the femoral canal and it is closed in the recent state by the septum crurale and a small lymphatic gland. It is bounded — In front, by the deep femoral arch and Pouparfs ligament ; Behind, by the pubes, covered by the pubic portion of the fascia lata and the pectineus muscle; Externally, by the femoral vein, from which it is separated by a fibrous septum ; Internally, by the deep femoral arch, Gimbernat's ligament, the transversalis fascia, and the conjoined tendon. The femoral canal is a space from a quarter to half an inch in length, extending from Gimbernat's ligament to the upper margin of the saphenous opening. Bounded above by the femoral ring and below by the saphenous opening. It is the innermost compartment of the femoral sheath. Anterior wall — transversalis fascia, separated liy the falci- form process of the fascia lata ; SURGICAL AXAlOiiy. 383 Fig^. I'lG.— Deep Femoral Region — the Femoral "Vessels, etc.. Cut Across as they Emerge Under Poupart's Ligament, AC, anterior crural nerve; CT, edge of the conjoiDed tendon; OE, crural ring; E, dotted line indicates the course of the deep epigastric artery; FS, femoral sheath; G, Gimbcrnat's ligament; IP; ilio-pectineal ligament; P, Poupart's ligament; PS, pectineus muscle. This muscle rests upon the pubic bone and is covered by its fascia,— the pectineal fascia,— which is somewhat thickened immediately beneath Poupart's ligament, where it is known as the pubic ligament of Cooper. It will be noticed that the femoral sheath is divided into three compartments; the outer for the femoral artery; the middle for the femoral vein; the Inner (CR) is the crural ring, the mouth of the crural canal. (McG-ratfi.) 384 HUMAN ANATOMY. Posterior wall — iliac fascia and pubic portion of the fascia lata; External wall — ^fibrous septum of the femoral vein; Internal wall — Gimbernat^s ligament, deep crural arch, and the junction of the iliac and transversalis fascia. The septum crurale is a layer of dense areolar tissue, sup- porting small lymphatic glands and closing in the femoral ring. It is perforated by numerous lymphatic vessels, and forms a bar- rier to the escape of hernia at this point. The crural or femoral sheath is a prolongation downward of the fascia lining the abdomen (transversalis fascia in front, iliac fascia behind the vessels), closely adhering to the femoral vessels to about an inch below the saphenous opening. Its upper part is funnel-shaped and its lower part continuous with the sheath of the vessel. It is divided by septa into three compartments; ia the outer is lodged the femoral artery; the middle is occu- pied by the femoral vein ; and the innermost, the femoral canal, is empty, or occupied by a lymphatic gland. Its outer border is pierced by the genito-crural nerve, its inner by the internal saphenous vein. The deef femoral arch is a fibrous thickening of the trans- versalis fascia which forms the anterior wall of the femoral sheath. It is also known as the deep crural arch. The saphenous opening is an oval aperture of half an inch in width, an inch and a half in length in the upper and inner part of the fascia lata, between its two divisions, the iliac and pubic portions. It is bounded externally by the falsiform process of Burns, internally by the pubic portion of the fascia lata, which curves upward behind the saphenous vein. The open- ing is covered externally by the cribriform fascia and the skin. The coverings of femoral hernia are : — 1. Skin; 3. Superficial fascia; 3. Cribriform fascia; 4. Femoral sheath, crural sheath, or fascia propria; 5. Septum crurale ; 6. Subserous connective tissue; 7. Peritoneal sac. ISCHIO-RECTAL REGION AND PERINEUM. The outlet of the pelvis is a lozenge-shaped space divided by an imaginary line drawn in front of the anus, transversally between the anterior margin of the tuber ischii, into two parts, the ischio-rectal region behind and the perineum in front. SURGICAL ANATOMY. 385 The ischio-rectal region contains the external and internal sphincters, the cornigator cutis ani and the ischio-rectal fossa. The latter is a pyramidal cavity, its apex reaching to the junction of the obturator and anal fascia, and its base formed by the skin. It is bounded — Behind, by the edge of the great sacro-sciatic ligament and gluteus maximus muscle ; internally, by the levator ani, covered by the anal fascia, sphincter ani, and coccygeus muscle; exter- nally, by the tuber ischii and obturator fascia. Fig. 170. Male perineum, superficial dissection. (.Lydston.) It contains: — The internal pudic artery, nerve and veins enclosed in the fascial canal or canal of Alcock ; The inferior hsemorrhoidal vessels and nerves ; The perineal and fourth sacral nerves ; and A mass of areolar tissue. PEEINEUM. The perineum is the triangular space between the anus and scrotum in the male, and between the anus and vulva in the female. The skin is dark, thin, and supplied with sebaceous and sudoriparous glands, and is marked by middle line or raphe. 386 HUMAN ANATOMY. FascicB of the Perineum. The pelvic fascia at the brim of the pelvis is continuous with transverse and iliac fascia; covers sacrum and pyriformis muscle, external iliac artery, and sacral nerves. It forms the anterior and lateral ligament of bladder (true) . It forms arcus tendinse, or white line, corresponding to division of pelvic fascia into two layers, the obturator and recto-vesical, extending from pubis to spine of ischium, from vrhich originates the levator ani muscle, -with recto-vesical and ischio-rectal or anal fascise above and below the muscle respectively. Jlftf^ of CarpuM CatftHutrnm ^nttpnai P