inspect Port ED rn * Hy RUSH t MeunldaU Mat etl at dpurererv eee ee taal eerlaerrorieees rhe Hens DWghieee Cetera he AAG Mea, Lak ghialitin( Aa! iar aa OS ew ch? @ 8a vs \ CORNELL UNIVERSITY LIBRARY WILLARD FISKE ENDOWMENT DATE DUE LOA rT ii | QL | ow | Caf | | \ \ | CATALOGUE OF THE PRESH-WATER FISHES OF AFRICA IN THE BRITISH MUSEUM (NATURAL HISTORY). VOLUME Il. BY GEORGE ALBERT BOULENGER, F.R.S. LONDON: PRINTED BY ORDER OF THE TRUSTEES. SOLD BY LONGMANS & Oo., 89 PATERNOSTER ROW, E.C.; B. QUARITOH, 11 GRAFTON STREET, NEW BOND STREET, W.; DULAU & Oo., 87 SOHO SQUARE, W.; AND AT THE BRITISH MUSEUM (NATURAL HISTORY), CROMWELL ROAD, SW. 1909, [ All rights reserved.| ALERE { FLAMMAM, PRINTED BY TAYLOR AND FRANCIS, RED LION COURT, FLEET STREET, LONDON. PREFACE. Tux scope of the present work, which will probably consist of three volumes, is sufficiently indicated by the Author’s Introduction. The first volume contains an account of the Selachii, Crossopterygii, and Dipneusti, which together comprise but a small fraction of the total number of species; and of the Teleostean Suborder Malacopterygii and part of the Ostariophysi. ‘The first of these Suborders includes the Mormyride as its most important constituent so far as the African fresh-water Fauna is concerned, while the second contains the large Families Characinide, Cyprinide, and Siluride. The greater part of the Cyprinidae, as well as the whole of the Siluride, will be dealt with in Volume II. The explorations of the Congo by the Government of the late Congo Free State and the Nile by the Egyptian Government, the ichthyological results of which have been worked out by the author of this Catalogue, have resulted in a large addition to the number of fresh-water fishes known to occur in the respective districts. ‘To the action taken by the two Governments, and to the labours of the individuals who have been concerned in the collection and preservation of the specimens referred to, the possibility of preparing this Catalogue has been mainly due. SIDNEY F. HARMER, Keeper of Zoology. Britisa Museum (Narvrat Hisrory), Lonpon. March, 1909. INTRODUCTION. Tne past decade has been productive of an enormous increase in our knowledge of the Fresh-water Fishes of Africa. ‘The explorations of the Congo and the Nile, undertaken at considerable expense by the Governments of the late Congo Free State and of Egypt, and of the great lakes of Central Africa, initiated in this country, have resulted in the discovery and description of an unexpectedly large number of generic and specific forms, types of most of which are deposited in the Natural History Museum. At the same time, concurrently with the study of the many thousands of specimens collected on the above-mentioned surveys, every effort was made to increase our acquaintance with the fishes of other parts of Africa, with results which are sufficiently apparent from the present Catalogue, intended to form three volumes, which, it is hoped, will afford a sound basis for the discussion of problems of distribution. Much remains to be done, however, in many parts of Africa; and no better incentive could be given to further progress on the part of collectors, as well as of students at home, than the compilation of such a simple handbook, containing a complete list of the specimens preserved in this great ichthyological collection, together with concise descriptions of and keys to the genera and species and text-figures of most of the latter. The greater number of these figures are reproduced, by permission, from the ‘ Annales du Musée du Congo,’ the ‘ Fishes of the Nile,’ the ‘Transactions’ and ‘ Proceedings’ of the Zoological Society of London, and the ‘ Annals and Magazine of Natural History,’ in which works the new species were described by Dr. Gunther and by myself. Other figures are reproduced from the publications of Drs. Peters, Steindachner, vl INTRODUCTION. In most cases when no Gray, Ayres, Day, Sauvage, and Pellegrin. ua original figures had been given, or when the figures were unsatisfactory, drawings have been prepared by Mr. J. Green and Mr. A. H Searle. Whenever a figure is reproduced from a previous publication, the source has been indicated. ‘The initials A. M. C. refer to the quarto plates of the ‘Annales du Musée du Congo,’ F. N. to those of the ‘Fishes of the Nile.’ In making use of the descriptions and keys as drawn up in this work, it should be borne in mind that the fins are not included in the total length or greatest depth, whilst in the indication of the total length of the largest specimen examined, which follows the description of every species, the fish is measured from the end of the snout to the extremity of the middle rays of the caudal fin; by depth of the caudal peduncle, 7.é. the free part of the caudal part of the body between the bases of the dorsal or anal and caudal fins, is meant the least depth, whilst the length is that of the muscular part only, the root of the caudal rays being usually covered with scales. By length of the dorsal and anal fins is meant the basal length. The abbreviations used in the fin and scale formule are too well known to all who work at fishes to need explanation here ; I will only observe that when both Roman and Arabic numerals are used to denote the number of dorsal or anal rays, tlie former refer to the simple or to the spinous rays, and the latter to the soft, branched rays, the last of which is usually cleft to the base, and that the interpolation of a comma between the numerals indicates discontinuity in the fins. G. A. BOULENGER. ZOOLOGICAL DEPARTMENT, 25th February, 1909. SYSTEMATIC INDEX. Subclass SELACHII. Ord. Il. DIPNEUSTI. Ord. I, PLAGIOSTOMI. Fam. 1. Leprposrrpnip.z. Page Subord. I. PLuvrRoTREMI, 1. Protopterus, Owen............ 19 Far. 1. Cipouigema, 1. annectens, Owen ........ 20 Page 2. sethiopicus, Heck. ........ 21 1. Carcharias, Cuv. 2... ....000. 2 3. dollol, Blogs ssa sieved ee wes 22 1. zambesensis, Peters ...... 2 Ord. III. TELEOSTEIL, Subord. II. HyPovRemti. Subord. Il. Matacorprrryatt, Fam. 1. Pristipz, Fam. 1. Enovipm. 1. Pristisy Lathe snascssaas tasks 3 1. perroteti, M. & Five Box. ue BS 3 1. Elops, Woe wittetn Rindden Caradon t 25 Ts: SAUPUS Lis octet ala wee ete 25 Subclass TELEOSTOMI. 2. Nacertay, Cee Vogccact sie 26 2. Megalops, Lacep. ............ 27 Ord. I. CROSSOPTERYGIL. 1. cyprinoides, Brouss. ...... 28 Subord. I, CLADISTIA. Fam. 2. Mormyrips. Hatin ty POLS Beier ya 1. Mormyrops, J. Mall, ........ 30 1. Polypterus, Geoff. .......... 5 1. deliciosus, Leach.......... 82 1. bichir, Geoffr..... 00.00... 6 2. anguilloides, Z. .......... 34 2. lapradii, Stdr...........0. ¢ 3. longiceps, Gthr........... 35 3. congicus, Blgr. ........44 9 4, breviceps, Stdr........... 36 4, endlicheri, Heck. ........ 10 5. engystoma, Blgr. . ...... 36 5. weeksii, Blyr............. 11 6. parvus, Blgr. so... eee. 37 6. ornatipinnis, Blgr......... 12 7. masuianus, Blgr. ........ 38 7. delhezi, Blgr............. 13 8. sirenoides, Blgr. ........ 39 8. senegalus, Cuv. .......... 14 9. zanclirostris, Gir. ...... 40 9. palmas, Ayres.........005 16 10. boulengeri, Pellegr. ...... 40 10. retropinnis, Vaill......... 17 11. curtus, Blgr. .........0.. 4] 2. Calamichthys, J..A. Smith .... 17 12. lineolatus, Blgr........... 42 1. calabaricus, J. A. Smith .. 18 13. nigricans, Blyr........... 48 Vill 1. Mormyrops (con.). 14, microstoma, Blyr. 15. mariv, Sehilth, oc uccwnnae 16. attenuatus, Bly. ise 17. furcidens, Pellegr. .......- 2. Petrocephalus, Marcus. . 1. bane, Lacep. ko . sauvagil, Blyr. . ansorgil, Blgr. .. 1... +54 . ballayi, Sauv. . simus, Sauv. bovei, C. & V. . keatingii, Blgr. ......-+-- . stuhlmanni, Blyr. ......-- . catostoma, Gthr. . gliroides, Vinevg. 11. degeni, Blgr. .........44. 3. Isichthys, Geld 1. henryi, Gall 4. Marcusenius, Gull 1. nigripinnis, Blgr. Dob ww oon Ww > . pulverulentus, Blyr. ...... . marchii, Sawv. . kingsleyee, Gthr. . sphecodes, Sauv. . brachistius, Gil . longianalis, Blgr. ........ . lhuysii, Stdr. . adspersus, thr. Oot oe &W H > . weeksii, Blgr. e - . bates, Blgr. 6 0cdvaewrns KH bo ry . ansorgii, Blgr. ro oo . pauciradiatus, Sidr. ...... _ ix . nigricans, Blgr. .. 6.6... ee . isidori, C. & V. . castor, Pappenh. jue. Or H a ron ~~] . harringtoni, Blgr. ...... . tumifrons, Blgr. . plagiostomus, Bly. w CO te £ bo oS . discorhynchus, Peters .... . petherici, Blgr. .......... . budgetti, Blgr. . macrops, Blgr. ro bo Do ke bo ww bo ns . psittacus, Blgr. 25. wilverthi, Blgr. 5. Stomatorhinus, Blyr. 1. puncticulatus, Blgr. ...... 49 56 58 71 Te. 73 74 75 75 77 78 79 80 81 82 SYSTEMATIC INDEX. 5, Stomatorhinus (co7.). Oe WwW to 6. 6. Myomyrus, Blyr. 1. 7. Guathonemus, Gill 1. wd ww wow DW SHH HPP Pe Ree PP Sankt ANKF OMAN ANKHFHWNH COCO MAIO OF OD bo 0 H ~y 29, 30. 31. 8. Genyomyrus, Blyr. Ls 9. Mormyrus, L. 1; 2, 3. 4, _ walkeri, Gihr. . humilior, Blgr. . corneti, Blgr. . polylepis, Blgr. . bruyerii, Pellegr. . angolensis, Blyr. . cyprinoides, L. . ussheri, Gthr. . greshoffi, Schilth. +. abadii, Blgr. . . tamandua, Gthr. . mirus, Blgr. . elephas, Blyr. microps, Blgr. ose. eevee macrodon, Blgr. ........ moorii, Gthr. . lambouri, Pellegr. ........ . schilthuisie, Blgr. . petersii, Gthr. . longibarbis, Hily. . niger, Gthr. . leopoldianus, Bly. . friteli, Pellegr. . bentleyi, Bigr. . livingstonii, Blgr. ........ . monteiri, Gihr. .. oo... . mento, Blgr. ............ . stanleyanus, Blgr. . senegalensis, Stdr. . macrolepidotus, Peters .... . pictus, Marcus. .......... BUM) BIGr. s.ogicey iaveeaies - kutuensis, Blgr. Se ee ee ee we . rhynchophorus, Blgr. ibis, Blgr. curvirostris, Blgr. .... 0... numenius, Blyr, donnyi, Blgr............, hasselquistii, C.&@ V..... anchietw, Guim macrophthalmus, Gthr. .... | ovis, Blyr. Page SYSTEMATIC INDEX. 9. Mormyrus (con.). Page 5. caballus, Blgr. «2.2.20... 132 6. tapirus, Puppenh. ........ 133 7. tenuirostris, Peters ...... 134 8. kannume, Forsk. .......,. 134 9. Gaschive, Ly ceisewsecpes 136 10. niloticus, Bl.-Schn. ...... 137 11. bozasi, Pellegr, .. 0.2.4... 138 12. longirostris, Peters........ 139 13. rume, C.&@ Ve. oo... eee 140 14. proboscirostris, Blyr....... 141 10. Hyperopisus, Gill .... 6.6... 142 1. bebe, Lacep. .... 0.0... 142 11. Gymnarchus, Cuv. .. 1.2.0... 144 1. niloticus, Cuv. .......... 144 Fam. 3. Noroprerip&. 1. Notopterus, Lacep. .......... 146 Later, Gihts. es ence ve ve ed 146 2. Xenomystus, Gthr. .......... 147 Ie pigris Gtlite. aes assess 147 Fam. 4. OsrmoGLossipx. 1. Heterotis, Hhrend.........5... 149 1. niloticus, Hhrenb. ........ 149 Fam. 5. PANToDONTID&. 1. Pantodon, Peters ............ 151 1. buchholzi, Peters ........ 151 Fam, 6, CLUPEIDE, 1, Clupéa; De vaeaeieee eee aeseers 153 1. finta, Cuv. ...... . 154 2. Pellonula, Gthr, .........0.. 155 1 NOnax, GUnhs. caves nln ons 156 9. miodon, Blgriwcccccs. uses 157 3. obtusirostris, Blgr. ...... 158 4, acutirostris, Blgr. ........ 159 3, Odaxothrissa, Blgr. .......... 160 1. losera, Blgr. .........04. 169 4, Microthrissa, Blgr. . 161 1, royauxi, Blgr. .......04. 161 5, Tlisha, Gray .. 0... ce eee eee 162 1. indica, Swains. .......... 163 6, Chanos, Lacep. .. 1... ee eee e ee 164 1. salmoneus, Forst. ........ 164 VOL. I. 1. Salmo, Art. bo H bo 3 Fam. 7, SaLMoNnID A, Ce a} 1. trutta, Z. Fam. 8. PHRACTOLEMIDE. . Phractolemus, Blgr........... 1. ansorgii, Blgr. Fam. 9. Kyerimp2. . Kmeria, Stdr. 2... eee 1. angolensis, Stdr. 2. spekil, Gar. ca vecaawseas 3. cameronensis, Blgr. ...... . Xenopomatichthys, Pellegr. .... 1. auriculatus, Pellegr. ...... Fam. 10. CRoMERIID A. . Cromeria, Blgr. oo. 6.5... ee 1. nilotica, Blgr. Subord. I]. OstarR1oPHyYSI. Fam. 1. CHARACINIDA. . Sarcodaces, Gthr. ............ 1. odoé, Bi. . Hydrocyon, Cuv. 1. forskalii, Cuv. 2. lineatus, Blkr. 3. vittiger, Blyr. 4, goliath, Blgr.....-....... 5. brevis, Gthr. wo... 0.00.05. . Bryconexthiops, Gthr. 1. microstoma, Gthr......... 2. -yseuxl, Blots a eaves desc . Alestes, M. & T. bs hd VPentexs Fa ciga teak ees . baremose, Joann. bt . macrophthalmus, Githr. .... . liebrechtsii, Blgr. ........ . stuhlmannii, Pfefe ........ . sadleri, Blgr . tholloni, Pellegr... 0.0... . Intermedius, Blgr......... CONann ey . longipinnis, Stdr, ........ ~ So . chaperi, Saw, ix Page 168 168 171 172 172 17 177 179 180 182 184 184 186 187 188 189 . 190 193 195 197 198 199 200 201 201 202 SYSTEMATIC INDEX. x +. Alestes (con.). Page 11. lateralis, Blgr. .....+.--- 204 12. nurse, Riipp. .. 6... - eee 205 13. affinis, Gihr. casaeses es + 208 14. imberi, Peters .......00555 209 15. humilis, Blgr. ...-...++> 211 16, kingsleya, Gthr, ......-- 212 17. bimaculatus, Blygr. .....- 213 18. teniurus, Gthr. .......... 214 19. opisthotenia, Blyr. ...... 215 20. popte, Pellegr. .......--- 216 21. macrolepidotus, C. § Ve .. 217 22. rhodopleura, Blgr......... 218 23. grandisquamis, Blgr. . 220 24, batesii, Blyr. ......-..4.. 221 25, brevis, Blgr. ..... 0.20 eee 222 5. Mieralestes, Blgr. ............ 223 1. aeutidens, Peters ........ 224 2. stormstp Blgr. sods seu 225 3. humilis, Blgr, 22.2.0... 226 4. holargyreus, Gthr......... 227 5. urotenia, Blgr. .......... 228 6. interruptus, Blgr. ........ 229 fe Bltus, BUGKS rsasina ea is 230 6. Petersius, Hilg............... 231 1. modestus, Blgr........... 232 2. hilgendorfi, Blgr. ........ 232 3. conserialis, Hilg, ........ 233 4. tangensis, Lonnb. ........ 234 5. leopoldianus, Blgr. ...... 235 6. brumpti, Pellegr. ........ 235 7. caudalis, Blgr. .......... 236 8. pulcher, Blgrs gcaeyiaeas 237 9. major, Bigr. csaveecceecs 237 10. spilopterus, Blgr. ........ 239 11. woosnami, Blgr, ........ 239 12. occidentalis, Gthr........ 240 7. Eugnatbichthys, Blgr. ........ 241 1. eetveldii, Blgr. .......... 241 2. macroterolepis, Blyr. . 243 4%. Paraphago, Blgr, ............ 244 1. rostratus, Blgr. 0 acces 244 9. Mesoborus, Pellegr. .......... 245 1. crocodilus, Pellegr. ...... 245 1, Bhaso; Gare cas cas wa ayey .. 246 1. loricatus, Githr. ........., 247 10. Phago (con.). 11, 12. 13. 14. 15, 16. 2. intermedius, Blgr......... 3. boulengeri, Schilth. Neoborus, Blqr. 1: 2 1. besse, Joann. ........-005 ornatus, Blgr. quadrilineatus, Pellegr. .... Ichthyoborus, Gthr. .......... Hemistichodus, Pellegr. Te Nannethiops, Gthr. .......... a Neolebias, Std. 1. 2. Distichodus, If. & 7. 1. . altus, Blgr. SOND or W LO Bee ee Ye Wk WD Oo DID A ff WwW lO . notospilus, Gthr. . faseiolatus, Blyr. . niloticus, Z. . niloticus, Joann. . elongatus, Blgr. . ocellicauda, Blgr. . tenia, Blgr. go; 18. Xenocharax, Gthr. 1; 2. vaillanti, Pellegr. uniteniatus, Gthr..... unifasciatus, Stdr..... trilineatus, Blyr. .... affinis, Gihr. ........ . hypostomatus, Pellegr. . maculatus, Blgr. . petersii, Pfeff. . antonil, Schilth. ...... . atroventralis, Blyr. - mossambicus, Peters .. . engycephalus, Gthr. .. lusosso, Schilth. brevis, Blgr. dimidiatus, Pellegr. spilurus, Gthr. crassus, Pellegr. » noboli, Blgr. as ce ecnewees . brevipinnis, Gthr. 1... .... . rostratus, Gihr. ... 06. .4.. . sexfasciatus, Blgr......... ite 17. Nannocharax, Gthr. Le . parvus, Pellegr. ...... . fasciatus, Gthr. 2.0... . intermedius, Blgr..... 2... SYSTEMATIC INDEX. Page 1. Labeo (con.). 19, Citharidium, Blyr. .......... 289 21. cyclorhynchus, Blgr. 1. ansorgii, Blgr. .......... 289 22. falcipinnis, Blgr. ...... 20. Citharinus, Cuv. ............ 290 23. kirkii, Blyr. 0.66000. 1. citharus, Geoffr, «11.4... 291 24. forskalii, Riipp. ........ 4. 2. congicus, Blyr, .......-. 293 25. cylindricus, Peters ...... 3. macrolepis, Blgr. ........ 294 26. nasus, Blgr. ......-.0- 4. latus, 11 & Ge esses 295 27. greenii, Blgr........... 5. gibbosus, Bir. . 297 28. lukule, Blgr. .. 0.06.20. 29. annectens, Blgr. ...... ee ae 30. chariensis, Pellegr. .... ol, parvus, BigP. os. ecea se: 1. Dabeo; Cuts aaxs ca css epee va 300 32. obscurus, Pellegr. ....... 1. niloticus, Forsk. .. 2.6.0... 304 33. umbratus, A. Smith .... 2, horie, Heck. ...... ... 3806 34. capensis, A. Smith .... 3. senegalensis, C. & V. . 808 35. ansorgii, Blgr, ........ 4, altivelis, Peters .......... 309 36. barbatus, Blgr. ........ 5. weeksii, Blgr, .......... 310 2. Discognathus, Heck. 1... 0.2... 6. lineatus, Blgr. .......... 311 1. dembeensis, Riipp. .... Te TOSB, States oye eyn tae ae 312 2. johnstonii, Blgr. ...... 8. mesops, Gthr. ........5- 313 3. vinciguerre, Blgr. .... 9. rudd, Blots seehiweetaas 314 4, makiensis, Blgr, ...... 10. velifer, Blgr. .....- 20-00. 315 5. blanfordii, Blgr. ...... 11. longipinnis, Blgr. ........ 316 6. hindi, Blgr, .......... 12. coubie, Rupp. «1... 317 7. quadrimaculatus, Ripp. 13. congoro, Peters .......... 319 3. Varicorhinus, Riipp........... 14. neumanni, Blgr. ........ 320 1. ansorgii, Blgr. ........ 15. gregorii, Gthr. ........-. 321 2, rien, Blgrs ee nese 16. darlingi, Blgr. ........., 321 3. tornieri, Stdr........... 17. victorianus, Blgr. ........ 322 4, beso, Riipp. .......... 18. fuelleborni, Hilg. & Pappenh. 328 5. maroccanus, Gthr......... 19. brachypoma, Gthr. ...... 324 6. tanganice, Blgr, ........ 20. macrostoma, Blgr......... 325 (To be continued in second volume.) CATALOGUE OF AFRICAN FRESH-WATER FISHES, COOee Subclass SELACHII. Skeleton cartilaginous, no membrane bones; jaws present. Exo- skeleton of dermal denticles, structurally identical with teeth. Males with paired intromittent organs (mixopterygia) attached to the ventral fins. Order I. PLAGIOSTOMI. Several (5-7) gill-clefts opening directly to the exterior. Pterygo- quadrate distinct from the cranium. Suborder I. PLEUROTREMI. Gill-openings lateral, the last in front of or above the base of the pectoral fin. Fam. 1, CARCHARIID. Mouth inferior. Eye with a nictitating membrane; spiracles small or absent; 6 gill-openings, one or two above the base of the pectoral fin. Two dorsal fins and one anal; caudal fin usually of the well-marked heterocercal type. VOL, Te B 2 CARCHARID®. 1, CARCHARIAS. Cuvier, Régne Anim. ii. p. 125 (1817); Mill. & Henle, Plagiost. p. 27 (1841) ; A. Dum. Hist. Poiss. i. p. 341 (1865); Giinth. Cat. Fish. viti. p. 357 (1870). Mouth crescentic; teeth with a single sharp cusp. No spiracles. First dorsal fin opposite to the space between pectorals and ventrals; a pit at the root of the tail. ‘Temperate and tropical seas. One species from the Zam besi. 1. CARCHARIAS ZAMBESENSIS. Carcharias (Prionodon) zambezensis, Peters, Mon. Berl. Ac. 1852, p. 276, and Reise Mossamb. iv. p. 7, pl. i. fig. 2 (1868). Snout very short, with rounded horizontal outline. Anterior border of mouth ona line with anterior border of eye. Teeth ZL finely serrated, upper triangular, with the outer border a little concave, lower much narrower; a small, non-serrated median tooth. Two gill-openings above the pectoral. Anterior dorsal just behind base of pectoral; second Fig. 1. Carcharias zambesensis, Type, after Peters (Reise Mossamb.). 4. dorsal a little anterior to, and of the same size as, anal, which is more distant from vent than from caudal. Ventral a little nearer caudal than pectoral. Caudal as long as distance from end of snout to dorsal. Brownish grey, belly yellowish white, Total length 760 millim. Zambesi, 120 miles from the coast.—Type in Berlin Museum PRISTIS, 3 Suborder Il. HYPOTREMI. Gill-openings ventral, below the base of the pectoral fin. Fam. 1. PRISTID. Mouth inferior. Snout produced into a long flat lamina armed with a series of strong teeth along each edge. No nictitating membrane ; spiracles present, large; 5 gill-openings. Body moderately depressed, elongate. ‘I'wo dorsal fins, no anal; caudal of the heterocercal type. 1. PRISTIS. Latham, Tr. Linn. Soe. ii. 1794, p. 276 ; Mill. & Henle, Plagiost. p. 105 (1841) ; A. Dum. Hist. Poiss. i. p. 471 (1865) ; Giinth. Cat. Fish. viii. p. 436 (1870). Mouth transverse, nearly straight; teeth minute, obtuse, pavement- like. First dorsal fin opposite or close to the base of the ventrals. Seas of the temperate and tropical regions, some species ascending rivers, 1. PRISTIS PERROTETI. Miill. & Henle, t.c. p. 108; A. Dum. t.c. p. 474; Peters, Reise Mossamb. iv. p. 9 (1868) ; Giinth. t. c. p. 486; Day, Fish. Ind. p. 729, pl. exci. fig. 1 (1878). 17-21 pairs of rostral teeth, not trenchant behind, and distant from one another. First dorsal originating in advance of ventrals, second not Pristis perroteta. Indian Ocean, after Day (Fish. Ind.). — $. much smaller. Root of pectoral in advance of first gill-opening. A small lower caudal lobe. Uniform greyish. Reaches a length of 3 metres. BQ 4 POLYPTERID 4. Tropical seas, entering rivers. Ascends the Zambesi and the Shiré and probably other African rivers. —Type in Paris Museum. 1, 2. Saws of half-grown specimens. Zambesi. Sir J. Kirk (P.). 3. Her. Gambia. Sir A. Moloney (P.). Subclass TELEOSTOMI. Skeleton more or less ossified; membrane bones present; jaws present. Exoskeleton of scales or bony plates. No mixopterygia. Order I. CROSSOPTERYGII. Paired fins, at least the pectorals, lobate, with an endoskeletal axis fringed with dermal rays. Mandibular arch suspended from the upper segment of the hyoid or hyomandibular arch (hyostylic skull); no supraoccipital bone; splenial present. Bulb of aorta muscular, with several rows of valves. Suborder I. CLADISTIA. Pectoral fin tribasal. Nostrils on upper surface of snout. Fam, 1. POLYPTERID. Body covered with rhombic bony scales with enamel-like coating. Dorsal fin formed of a series of spines with one or several articulated rays supporting a membrane, forming detached finlets. Tead covered with bony plates; maxillary bone bordering the eye; a spiracle, covered by a bony valve, on each side of the top of the head. Branchiostegal rays replaced by a pair of large bony plates between the mandibular rami. Young with an external opercular gill, Vent opening far back. Vertebral column ossified, diphycercal. Air-bladder double, with pneumatic duct communicating with the ventral side of the esophagus. Two genera, both African :— Ventral fins present. . . . . . . . . 1. Polypterus, Geoffr., p. 5. Novventralhus: ga 4G) @ ew Ga we DE Calamichthys, J. A. Smith, p.17. POLYPTERUS. 5 1. POLYPTERUS. Geoffr. Ann. Mus. Paris, i. 1802, p. 57; A. Dum. Hist. Poiss. ii. p. 391 (1870) ; Giinth. Cat. Fish. vii. p. 326 (1870); Bouleng. Poiss. Bass. Congo, p. 20 (1901), and Fish. Nile, p. 2 (1907). Body more or less elongate, not serpentiform; spines of the dorsal fin bearing two or more articulated rays; ventral fins present ; suboperculum well developed ; teeth small, conical. Nile and Tropical Africa. Synopsts of the Species. I. Mandible projecting slightly beyond the snout; suboperculum much larger than the eye; pectoral extending as far as first dorsal spine, or beyond ; 11-18 dorsal spines ; 11-15 scales between occiput and first dorsal spine. A, 60-70 scales along the body, 46-54 round the middle. , D. XIV-XVIUI; interorbital region slightly convex, 12 (young) to 2} diameters of eye ; 63-70 scales in longitudinal series. . . . 1. P. bichir, Geoffr., p. 6. D. XILI-XYV ; interorbital region quite flat, 2 (young) to 33 diameters of eye ; 60-68 scales in longitudinal series . 2 1 1 ww ww oP. lapradi, Stdr., p. 7. B. 50-59 scales along the body. D. XII-XLV ; interorbital region flat, not more than 3 diameters of eye; usually a small azygous shield between the nasals, in the adult ; 46-52 scales round middle of body . 3. P. congicus, Blgr., p. 9. D. XI-XIV; interorbital region flat or slightly concave, 2 (young) to 32 diameters of eye ; no azygous shield on the snout; 42-46 scales round middle of body . . 2. 1. «4 P. endlicheri, Heck., p. 10. II. Both jaws equal in front, or snout projecting beyond mandible; sub- operculum not much larger, or even smaller than the eye; 5-11 dorsal spines. A. 60-65 scales along the body, 42-46 round the middle ; pectoral widely separated from first dorsal spine ; interorbital region flat or slightly convex, 3 diameters of cye in adult ; 22-26 scales between occiput and first dorsal spine. D. IX-X ; body strongly depressed ; 44-46 scales round middle of body . : D. X-XI ; body feebly depressed, desiaeal: 42 scales round middle of boy . 5. P. weeksit, Blgr., p. 11. 6. P. ornatipinnis, Blgr., p. 12. 6 POLY PTERID. B. 52-61 scales along the body, 32- 40 round the middle ; interorbital region convex, not more than 2 diameters of eye. 1. Pectoral reaching or nearly reaching first dorsal spine. D. X-XI; 56 scales in longitudinal series, 38 round middle of body, 14-17 between occiput and first dorsal spine 2. 0. ee ee 7 P. delhezi, Blgr., p. 13. 2. Pectoral widely separated from first dorsal spine. D. VITI-XI; 53-61 scales in longitudinal series, 34-40 round middle of body, 15-20 between occiput and first dorsal spine. . 6.) 8. Pe senegalus, Cuv., p. 14. D. V-IX ; 52-56 scales in longitudinal series, 36-10 round middle of body, 23-26 between occiput and first dorsal spine . . . 9, P. palmas, Ayres, p. 16. D. VI-VII; 58-59 scales in ean series, 32-36 round middle of body, 35-37 between occiput and first dorsal spine . . .) .) . 10. P. retropinnis, Vaill., p. 17. 1, POLYPTERUS BICHIR. Geoffr, Ann. du Mus. i. 1802, p. 57, pl. v., and Deser. Egypte, Poiss. p. 4, ili, (1809) ; Gnichen. Mag. Zool. 1839, Poiss. p. 10; Giinth. Petherick’s Tray. ii. p. 267 (1869); A. Dum. Hist. Poiss. ii. p. 391, pl. xxiii. fig. 1 (1870) ; Schweinf. Herz. Afr. i. p. 253, fig. (1874) ; Steind. Denkschr. Ak. Wien, xliv. 1882, p. 52, pls. viii. & ix. fig..1 ; Bouleng. Ann. & Mag. N. H. (7) ii. 1898, p. 417; 8. Flower, Proc. Zool. Soc. 1900, p. 968 ; Werner, Zool. Jahrb., Syst. xxi. 1904, p. 271, and Sitzb. Ak. Wien, exv.i. 1906, p. 1118; Bouleng. Fish. Nile, p. 5, pl. i. (1907). Polypterus bichir, part., Giinth, Cat. Fish. viii. p. 326 (1870). Head twice or nearly twice as long as broad, rather strongly flattened, its length 44 to 52 times in total length, with supero-lateral eyes and slightly convex interorbital region; lower jaw projecting slightly beyond upper; length of snout 5 to 7 times in length of the head; eye 83 (young) to 13 times in length of the head, 12 (young) to 24 times in inter- ocular width; latter less than distance between eye and spiracle ; nasal bones in contact on the median line in the adult; a series of 4 to 7 shields between postorbital and spiracular shields ; suboperculum much larger than eye. Dorsal fin with XIV-XVIII Spines; spines over- lapping when folded down. Anal 11-15. Pectoral reaching beyond vertical of first dorsal spine. 63-70 scales in a longitudinal series, 11-15 between occiput and first dorsal spine, 46-54 round middle of body. Greyish or greenish olive above, the young with 10-13 cross-bars on the back and 2 or 3 stripes along the sides of a darker shade, these POLYPTERUS. > ( markings becoming more indistinct or disappearing in the adult; a few scattered black spots sometimes present; paired fins with more or less distinct transverse streaks or series of spots, the other fins without spots ; belly yellow. Total length 720 millim. Nile, Lake Rudolf, Chad Basin.—Types in Paris Museum. Fig. 3. 1 4-! tT Oo 3 Vg . Ad. v 6. Ad., stffd. 7. Her. 8. Ad. 9-12. Hor. & yg. Iie Ye. 15-24. Ad. & hgr. 25. Her., skel. 26. Ad. 27. Ad. 28, Ad. 29, 30. Ad., stffd. Polypterus bichir. Nile Delta (F. N.). 4. Near Damietta. Mansurah, Damietta Nile. Cairo. Near Cairo. Kawa, White Nile. Goz abu Gumah, White Nile. Gharb-el-Aish, 5 Fashoda, - Near Kerro, re Mouth of Lake No, ,, Bahr-el-Gebel. Nile. 2, POLYPTERUS LAPRADII. Dr. J. C. Mitchell (P.). N. R. Harrington, Esq., and Dr. Hunt (E.). Dr. Riippell (C.). L. Loat, Esq. (C.). ” ” ” Capt. S. Flower (P.). Steind. Sitzb. Ak. Wien, lix. i. 1869, p. 103, pl. i. figs. 1 & 2, and pl. ii. figs. 1 & 2; Bouleng. Ann. & Mag. N. Hy (7) it. 1898, p. 418; Budgett, Proc. Cambr. Philos. Soc. x. 1900, p. 236 ; Bouleng. Proc. Zool. Soc. 1902, i. p. 123, pl. x. figs. 1 & 2. Head rather strongly depressed, 1 to 25 times as long as broad, its 8 POLYPTERID. length 43 to 5} times in total length; lower jaw projecting a little beyond snout; eye supero-lateral, its diameter 8 (young) to 12 times in length of head, 2 to 34 times in width of interorbital region, which is quite flat and not less than the distance from eye to spiracle ; all head- shields paired; a series of 4 to 6 shields between postorbital and Fig. 4. Polypterus lapradit. Head of type, after Steindachner (Sitzb. Ak. Wien, 1869). 3. spiracular shields; suboperculum much larger than eye. Dorsal with XIII-XYV spines, anterior more or less overlapping when folded. ‘Anal 11-15. beyond. Pectoral reaching vertical of first dorsal spine, or a little 60-68 scales in a longitudinal series, 11-13 between occipital and first dorsal spine, 46-52 round middle of body. Yellowish or pale olive, with dark stripes and more or less distinct cross-bars ; fins with dark spots or streaks. Total length 740 millim. Senegal, Gambia, Niger.—Types in Vienna Museum. 1. Ad., stffd. 2. Ad. 3-10. Ad. & hgr. 11. Hegr., stffd. 12. Her. 13. Ad. 14. Yg. 15-17. Yg. 18, 19. Her. 20-26. Ad., her., & yg. 27. Ad., skel. 28. Her. Senegal. St. Louis, Senegal. Kaédi, Senegal. Gambia. Abo, Lower Niger. Assay, Niger. 2”? 29 2? Benue River, N. of Ibi. M. P. Delhez (C.). P. Whitfield, Esq. (P.). J.S. Budgett, Esq. (P.). Capt. Vipan (P.). Dr. W. J. Ansorge (C.). Mr. J. T. Dalton (C.). W. A. Forbes, Esq. (P.). H. 1. Norton Traill, Esq. (U.). POLYPTERUS. . 3. POLYPTERUS CONGICUS. Bouleng. Ann. & Mag. N. H. (7) ii. 1898, p. 418, Ann. Mus. Congo, Zool. i. pl. xxx. fig. 1 (1899), Poiss. Bass. Congo, p. 22 (1901), Proc. Zool. Soc. 1902, i. », 123, pl. xi. fig. 1, and Tr. Zool: Soc. xvi. 1901, p. 142, and xvii, 1906, p. 544. Head rather depressed, 14 to 1$ times as long as broad, its length 4 to 5 times in total length; lower jaw projecting a little beyond snout ; eye supero-lateral, its diameter 8 (young) to 12 times in length of head, 12 to 3 times in width of interorbital region, which is flat; nasal shields usually separated, in the adult, by a small azygous shield; a series of 4 or 5 shields between postorbital and spiracular shields; suboperculum much larger than eye. Dorsal with XII-XIV Polypterus congicus. Type. 1. spines, anterior more or less overlapping when folded. Anal 12-15. Pectoral reaching vertical of first dorsal spine, or beyond. 65-59 scales in a longitudinal series, 11-16 between occiput and first dorsal spine, 46-52 round middle of body. Olive above, yellowish beneath ; 6 to 8 more or less regular blackish narrow transverse bands on the body; fins spotted or mottled with blackish. Total length 720 millim. Congo, Lake Tanganyika. 1. Ad., type. Stanley Falls. Rey. W. H. Bentley (C.). 2. Ye. New Antwerp, Upper Congo. Capt. Wilverth (C.). ou XS Bolobo, Upper Congo. Rev. G. Grenfell (P.). 4. Yg. Bangala Country, Upper Congo. M. De Meuse (C.). 5. Ad. Upper Congo. Brussels University. 6-9. Ad. Kalambo, L. Tanganyika. Prof. J. E.8. Moore (C.). 10. Ad. Kituta, 45 Dr.W.A. Cunnington(C.). 10 POLYPTERIDA, 4. POLYPTERUS ENDLICHERI. Heckel, Russegger’s Reise Egypt. ili. p. 310, pl. xxii. fig. 1 (1849) ; A. Dum, Hist. Poiss. ii. p. 393, pl. xxiii. fig. 3 (1870) ; Steind. Denkschr. Ak. Wien, xliv. 1882, p. 52, pls. viii. & ix. fig. 2; Bouleng. Ann. & Mag. N. H. (7) ii. 1898, p. 418, and Proc. Zool. Soc. 1902, i. p. 124, pl. si. fig. 2; Werner, Zool. Jahrb., Syst. xxi. 1904, p. 271, and Sitzb. Ak. Wien, cxv. i. 1906, p-. 1118; Bouleng. Fish. Nile, p. 9, pl. 11. (1907). Polypterus senegalensis, part., Giinth. Petherick’s Trav. ii. p. 268 (1869). Polypterus bichir, part., Giinth. Cat. Fish. viii. p. 326 (1870). Bedy more or less distinctly flattened, at least in front. Head 14 to 14 times as long as broad, 42 to 52 times in total length, much flattened, with supero-lateral eyes and flat or slightly concave interorbital region ; Polypterus endlicheri. White Nile (FL N.). 2. lower jaw projecting slightly beyond upper; length of snout 5 to 7 times in length of head; eye 9 (young) to 15 times in length of head, 2§ (young) to 33 times in the interocular width; latter more than distance between eye and spiracle; nasal bones in contact on the median line in the adult; a series of 4 to 6 shields between postorbital and spiracular shields ; suboperculum much larger than eye. Dorsal with XI-XIV spines, anterior not or but feebly overlapping when folded. Anal 15-18. Pectoral reaching to or beyond vertical of first dorsal spine. 60-88 scales in a longitudinal series, 11-15 between occiput and first dorsal spine, 42-46 round middle of body. Grey or greyish olive above; 4 to 6 more or less regular broad blackish bands across the back: on the POLYPTERUS. 11 sides these bands are directed backwards, or break up into branches or detached spots; lips and fins with blackish spots, these usually forming transverse lines on the pectorals and ventrals, and oblique wavy lines or vermiculations on the dorsal; belly white. Total length 630 millim. White Nile, Bahr-el-Gebel, Lake Chad, Niger—Type in Vienna Museum. 1. Ad., stffd. Khartum. J. Petherick, Esq. (C.). 2. Her. *) 3-4. Ad. & yg. Shederah, near Omdurman. Li. Loat, Esq. (C.). 5d. Yg. Goz Shebesha, White Nile. . 6. Ad. Near Kawa, 53 » 7. Ad., skel, és o ij 8. Ad. Goz abu Gumah, __,, 3 9. Ad. Fashoda, as ” Ic. Ad. Mouth of Lake No, ,, a ll. Ad. Gondokoro, Bahr-el-Gebel. a 12. Her. Lake Chad. Capt. G. B. Gosling (P.). 13. Yg. Agberi, Lower Niger. Dr. W. J. Ansorge (C.). 14, 15,16. Ad. & lgr., Niger. Mr. J. T. Dalton (C.). stftd. 5. POLYPTERUS WEEKSII. Bouleng. Ann. & Mag. N. H. (7) ii. 1898, p. 419, Poiss. Bass. Congo, p. 24 (1901), and Proc. Zool. Soc. 1902, ii. p. 124, pl. x. fig. 4. Polypterus weeksit. Young, type (P. Z.S. 1902). 2. Body depressed. Head 14 to 1§ times as long as broad, 43 to 5} times in total length, much flattened, with supero-lateral eyes and flat interorbital region; snout projecting slightly beyond the lower jaw ; eye 7 (young) to 93 times in length of head, 2} to 3 times in interorbital width; a small azygous shield may be present between the nasals in the adult; a series of 3 or 4 shields between postorbital and spiracular 12 POLYPTERIDE, shields; suboperculum a little smaller or a little larger than eye. Dorsal with IX—X spines, not overlapping when folded. Anal 10-11. Pectoral widely separated from first dorsal spine. 60-65 scales in a longitudinal series, 22-26 between occiput and first dorsal spine, 44-46 round middle of body. Olive above, yellowish beneath ; 7 or 8 blackish bars on the back, bifurcating on the lower part of the sides; fins spotted, marbled, or irregularly barred with blackish; young with a black spot on the membrane behind each dorsal spine. Total length 380 millim. Upper Congo, Katanga. 1. Yg., Type. Monsembe. Rey. J. H. Weeks (P.). 2, Ad. ” ” 6. POLYPTERUS ORNATIPINNIS. Bouleng. Ann. Mus. Congo, Zool. ii. p. 23, pl. vil. fig. 1 (1902), and Ann. & Mag. Ni EL, (7) svi. 1905, ps 641. Head and body less depressed than in the preceding. Head 14 times as long as broad, 6 times in total length; snout projecting very slightly Fig. 8. Polypterus ornatipinnis. Type (A.M.C.). 4, beyond lower jaw; eye lateral, 83 times in length of head, 3 times in width of interorbital region, which is slightly convex; all head-shields paired ; a series of 4 shields between postorbital and spiracular shields ; suboperculum slightly larger than eye. Dorsal with X-XI spines, nik POLYPTERUS. 13 overlapping when folded. Anal 15. Pectoral widely separated from first dorsal spine. 62-63 scales in a longitudinal series, 24-25 between occiput and first dorsal spine, 42 round middle of body. Olive above, with ill-defined light spots, whitish beneath; lips and gill-membrane spotted with black; dorsal fin spotted with black; pectorals, ventrals, anal, and caudal barred with black. Total length 370 millim. Upper Congo (Monsembe and Kassai River). 1. Type. Monsembe. Rev. J. H. Weeks (P.). 7. POLYPTERUS DELHEZI. Bouleng. Ann. Mus. Congo, Zool. i. p. 61, pl. xxx. fig. 2 (1899), and Poiss. Bass. Congo, p. 23 (1901) ; Pellegr. Bull. Mus. Paris, 1904, p. 310. Body feebly compressed, subcylindrical. Head 12 to 12 times as long as broad, 54 to 6 times in total length, with lateral eyes and feebly convex interorbital region ; jaws equal in front; eye 74 times in length of head, 14 times in interorbital width ; head-shields all paired; a series of 3 shields between postorbital and spiracular shields ; suboperculum as large as eye, or a little larger. Dorsal with X-—XI spines, not Fig. 9. Polypterus delhezt. Type (A.M.C.). 2. overlapping when folded. Anal 11. Pectoral reaching or not quite reaching vertical of first dorsal spine. 56 scales in a longitudinal series, 14-17 between occiput and first dorsal spine, 38 round middle of body. Brownish above, whitish beneath; 7 black transverse bands on the back ; fins mottled with dark brown. Total length 340 millim. 14 POLYPTERID-F. Upper Congo (Lake Leopold II. and New Antwerp) ; Shari.—Type in Congo Museum, Tervueren. 1. One of the types. New Antwerp. Capt. Wilverth (C.). 8. POLYPTERUS SENEGALUS. Cuvier, Régne Anim. 2nd ed. p. 330 (1829), and Régne Anim. Illustr., Poiss. pl. ev. fig. 2 (1836) ; Guichen. Mag. de Zool. 1839, Poiss. p. 11, pl. i.; Steind. Sitzb. Ak. Wien, lix. i. 1869, p. 106, pl. i. figs. 3-5 ; A. Dum. Hist. Poiss. ii. p. 394 (1870); Steind. Denk